Tonsilla jinyunensis, Zhang & Irfan & Wang & Zhang, 2022

Zhang, Meng, Irfan, Muhammad, Wang, Lu-Yu & Zhang, Zhi-Sheng, 2022, Six new species of Tonsilla Wang & Yin, 1992 (Araneae: Agelenidae) from southern China, with the first description of the male of T. yanlingensis (Zhang, Yin & Kim, 2000), Zootaxa 5091 (2), pp. 357-372 : 358-360

Tonsilla jinyunensis sp. nov.

Figures 1 , 2A–E , 9A–B , 10

Type material. Holotype ♂: CHINA, Chongqing City, Jinyun Mountain, Guankou , 29.839486°N, 106.388354°E, 650 m, 24.X.2011, L.Y. Wang, D. Wang & M.X. Liu leg. (SWUC-T-AG-49-01) ; Paratypes (4 ♂ and 8 ♀): 2 ♀, same data as holotype (SWUC-T-AG-49-02~03) ; 1 ♀, Gele Mountain Forest Park, 29.573881°N, 106.434339°E, 728 m, 14.IV.2011, L.Y. Wang & Z.X. Li leg. (SWUC-T-AG-49-04) ; 4 ♂ and 5 ♀, Beibei District , Jindaoxia Valley, 30.028285°N, 106.624741°E, 880 m, 23.XI.2008, Z.S. Zhang leg (SWUC-T-AG-49-05~13) .

Etymology. The specific epithet refers to the type locality; adjective.

Diagnosis. Males of this new species resemble those of T. rostrum ( Jiang et al. 2018) in having similar tegular sclerites, conductor’s dorsal apophysis and tibial apophysis, but can be distinguished by the following characters: the apex of the conductor slightly curved, narrowing towards tip with a blunt end in ventral view ( Fig. 2B, C ), vs. apex bifurcated with pointed end ( Jiang et al. 2018, fig. 23A); ventral margin of median apophysis wider than long in retrolateral view ( Fig. 2B, C ), vs. longer than wide ( Jiang et al. 2018, fig. 23B); the anterior margin of conductor lamella round in ventral view ( Fig. 2B ), vs. protruding laterally with a blunt tip ( Jiang et al. 2018, fig. 23A); the patellar apophysis extends to the ventral side of the tibia in retrolateral view ( Fig. 2C ), vs. extending above the tibia ( Jiang et al. 2018, fig. 23B). Female of the new species can be distinguished from all other congeners by the following characters: anterior and lateral margins of atria somewhat rectangular, narrowing posteriorly, epigynal teeth absent; spermathecae human brain-shaped and the copulatory ducts long and indistinct ( Fig. 2D, E ).

Description. Male (holotype, Fig. 9A ). Total length 15.65. Carapace 8.27 long, 4.92 wide; opisthosoma 7.01 long, 4.07 wide. Eye sizes and interdistances: AME 0.23, ALE 0.31, PME 0.26, PLE 0.33; AME–AME 0.23, AME– ALE 0.15, PME–PME 0.18, PME–PLE 0.40, ALE–PLE 0.14. MOA 0.81 long, front width 0.69, back width 0.75. Clypeus height 0.35. Leg measurements: I 21.43 (5.67, 6.69, 5.39, 3.68); II 18.80 (5.04, 5.99, 4.62, 3.15); III 16.89 (4.63, 5.04, 4.51, 2.71); IV 22.38 (5.79, 7.05, 6.39, 3.15). Spination of legs: femur I-II 220, III 240, IV 230; patella I-II 000, III-IV 020; tibia I-II 026, III-IV 046; metatarsus I 026, II 066, III-IV 088.

Palp ( Fig. 2A–C ). Patella short, patellar apophysis longer than patella, slightly curved with pointed end, extending forward. Tibia with retrolateral and lateral apophyses with blunt tips. Cymbium approximately 3 times longer than wide; cymbial furrow extensively sclerotized, 1/3 of cymbium length. Median apophysis spoon-like. Conductor sclerotized, longer than wide, slightly curved in ventral view, distal end beak-shaped in ventral view. Conductor lamella conspicuous with round margin, conductor’s dorsal apophysis long, rod-shaped with pointed end. Embolus filiform, originating at the 7 o’clock position.

Female (one of the paratypes from Jinyun Mt., Fig. 9B ). Total length 16.05. Carapace 8.25 long, 4.86 wide; opisthosoma 8.28 long, 5.66 wide. Eye sizes and interdistances: AME 0.24, ALE 0.32, PME 0.32, PLE 0.30; AME– AME 0.21, AME–ALE 0.14, PME–PME 0.20, PME–PLE 0.44, ALE–PLE 0.15. MOA 0.80 long, front width 0.65, back width 0.75. Clypeus height 0.30. Leg measurements: I 17.97 (4.85, 6.29, 4.07, 2.76); II 14.14 (4.01, 4.58, 3.34, 2.21); III 13.87 (3.95, 4.44, 3.64, 1.84); IV 19.48 (5.31, 6.40, 5.24, 2.53). Spination of legs: femur I-II 220, III 140, IV 210; patella I-II 000, III-IV 020; tibia I 016, II 026, III-IV 046; metatarsus I 006, II 046, III 096, IV 086.

Epigyne ( Fig. 2D, E ). Epigynal plate somewhat rectangular, teeth absent. Atrial margins distinct, narrowing posteriorly. Epigynal hoods shallow, indistinct, located ventro-mesally. Copulatory opening located in anterior margin of epigynal plate. Copulatory ducts long, indistinct. Spermathecae human brain-shaped, separated by less than 1/3 of their width.

Distribution. China (Chongqing) ( Fig. 10 ).

Figures

FIGURE 1. Live female specimen of Tonsilla jinyunensis sp. nov. from Jinyun Mountain.

FIGURE 2. Tonsilla jinyunensis sp. nov.A.Male holotype, left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Epigyne, ventral view; E. Same, dorsal view. Abbreviations: At, atrium; CD, copulatory duct; CDA, Conductor’s dorsal apophysis; CF, cymbial furrow; CL, conductor lamella; Co, conductor; CO, copulatory opening; EH, epigynal hood; Em, embolus; FD, fertilization ducts; LTA, lateral tibial apophysis; MA, median apophysis; PA, patellar apophysis; RTA, retrolateral tibial apophysis; SH, spermathecal heads; Sp, spermathecae.

FIGURE 9. Habitus of Tonsilla spp. A–B, male holotype and female paratype of T. jinyunensis sp. nov.; C–D, male holotype and female paratype of T. jiugongensis sp. nov.; E–F, male holotype and female paratype of T. rutunda sp. nov.; G–H, male holotype and female paratype of T. subrostrum sp. nov.; I–J, male holotype and female paratype of T. subtruculenta sp. nov.; K–L, male and female of T. yanlingensis (Zhang, Yin & Kim, 2000); M, male holotype of T. yueliangensis sp. nov.

FIGURE 10. Distribution of Tonsilla species treated here.