Tonsilla subrostrum, Zhang & Irfan & Wang & Zhang, 2022

Zhang, Meng, Irfan, Muhammad, Wang, Lu-Yu & Zhang, Zhi-Sheng, 2022, Six new species of Tonsilla Wang & Yin, 1992 (Araneae: Agelenidae) from southern China, with the first description of the male of T. yanlingensis (Zhang, Yin & Kim, 2000), Zootaxa 5091 (2), pp. 357-372 : 363-365

Tonsilla subrostrum sp. nov.

Figures 5A–E , 9G–H , 10

Type material. Holotype ♂: CHINA, Hunan Province, Zhangjiajie City, Wulingyuan District, Wulingyuan Scenic Spot , Jinbianxi , 29.338675°N, 110.451479°E, 876 m, 07.XI.2020, Y.N. Mu leg. (SWUC-T-AG-52-01) ; Paratypes: 1 ♂ and 4 ♀, same data as holotype (SWUC-T-AG-52-02~06) .

Etymology. The specific epithet refers to its similarity to Tonsilla rostrum Jiang, Chen & Zhang, 2018 ; noun.

Diagnosis. The male of this new species resembles Tonsilla rostrum ( Jiang et al. 2018) in having the similar tegular sclerite, conductor’s dorsal apophysis, patellar and tibial apophysis of male palp. The females of both species have epigynes with long epigynal teeth. Both species can be distinguished by the following characters: apical end of conductor horn-shaped, relatively broad with blunt end and conductor’s dorsal apophysis short, slightly extending above the inner margin of cymbium in ventral view in T. subrostrum sp. nov. ( Fig. 5B, C ), vs. spine-shaped with sharp pointed end and conductor’s dorsal apophysis long, almost touching the outer margin of cymbium in T. rostrum ( Jiang et al. 2018, figs 23A, B, 24B, C). Proximal margin of median apophysis round in retrolateral view in T. subrostrum sp. nov. ( Fig. 5B, C ), vs. longer than wide in T. rostrum ( Jiang et al. 2018, figs 23A, B, 24B, C). The female of T. subrostrum sp. nov. resembles T. rostrum in having similar copulatory ducts situated dorso-mesally but can be distinguished by the atrial septum longer than wide in T. subrostrum sp. nov. ( Fig. 5D ), vs. almost triangular in T. rostrum ( Jiang et al. 2018, figs 23C, 24D). The spermathecae are oval in T. subrostrum sp. nov. ( Fig. 5E ) but bean-shaped in T. rostrum ( Jiang et al. 2018, figs 23C, 24D).

Description. Male (holotype, Fig. 9G ). Total length 7.98. Carapace 4.45 long, 2.86 wide; opisthosoma 3.54 long, 2.36 wide. Eye sizes and interdistances: AME 0.16, ALE 0.22, PME 0.24, PLE 0.23; AME–AME 0.06, AME– ALE 0.06, PME–PME 0.10, PME–PLE 0.14, ALE–PLE 0.07. MOA 0.48 long, front width 0.35, back width 0.52. Clypeus height 0.18. Leg measurements: I 11.66 (3.12, 3.91, 2.73, 1.90); II 9.80 (2.78, 2.93, 2.44, 1.65); III 9.21 (2.58, 2.70, 2.59, 1.34); IV 12.73 (3.42, 3.95, 3.61, 1.75). Spination of legs: femur I-III 220, IV 230; patella I-II 000, III 020, IV 110; tibia I 006, II 026, III 046, IV 146; metatarsus I 006, II 046, III 056, IV 086.

Palp ( Fig. 5A–C ). Patella as long as the patellar apophysis; patellar apophysis long, finger-shaped with blunt end, extending forward in retrolateral view. Retrolateral and lateral tibial apophysis distinct with blunt end. Cymbium approximately 3 times longer than wide; cymbial furrow extensively sclerotized, 1/3 of the cymbial length. Median apophysis spoon-like, proximal margin round. Conductor apex horn-shaped, relatively broad, slightly curved with blunt end, lower arm spine-like with pointed end. Conductor’s dorsal apophysis short and with a pointed end. Conductor lamella distinct. Embolus filiform, originating at the 7 o’clock position.

Female (one of the paratypes, Fig. 9H ). Total length 9.08. Carapace 4.65 long, 2.75 wide; opisthosoma 4.71 long, 3.13 wide. Eye sizes and interdistances: AME 0.15, ALE 0.22, PME 0.23, PLE 0.24; AME–AME 0.11, AME– ALE 0.08, PME–PME 0.12, PME–PLE 0.19, ALE–PLE 0.08. MOA 0.55 long, front width 0.40, back width 0.60. Clypeus height 0.13. Leg measurements: I 10.61 (2.87, 3.57, 2.54, 1.63); II 9.04 (2.60, 2.84, 2.18, 1.42); III 8.44 (2.22, 2.69, 2.32, 1.21); IV 11.86 (3.22, 3.92, 3.12, 1.60). Spination of legs: femur I 210, II 220, III 140, IV 220; patella I 000, II 022, III 120, IV 110; tibia I 006, II 025, III-IV 046; metatarsus I 006, II 056, III 0106, IV 086.

Epigyne ( Fig. 5D, E ). Epigynal plate almost round, teeth situated anteriorly. Atrial ridges smooth. Atrium with distinct septum. Epigynal hoods distinct, located ventro-mesally. Copulatory opening situated posteriorly in the atrium. Copulatory ducts situated in between spermathecae. Spermathecae oval, separated by a distance half of their length.

Distribution. China (Hunan) ( Fig. 10 ).

Figures

FIGURE 5. Tonsilla subrostrum sp. nov. A. Male holotype, left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Epigyne, ventral view; E. Vulva, dorsal view. Abbreviations: CD, copulatory duct; CDA, Conductor’s dorsal apophysis; CF, cymbial furrow; CL, conductor lamella; Co, conductor; CO, copulatory opening; EH, epigynal hood; Em, embolus; ET, epigynal teeth; FD, fertilization ducts; LTA, lateral tibial apophysis; MA, median apophysis; PA, patellar apophysis; RTA, retrolateral tibial apophysis; Sp, spermathecae.

FIGURE 9. Habitus of Tonsilla spp. A–B, male holotype and female paratype of T. jinyunensis sp. nov.; C–D, male holotype and female paratype of T. jiugongensis sp. nov.; E–F, male holotype and female paratype of T. rutunda sp. nov.; G–H, male holotype and female paratype of T. subrostrum sp. nov.; I–J, male holotype and female paratype of T. subtruculenta sp. nov.; K–L, male and female of T. yanlingensis (Zhang, Yin & Kim, 2000); M, male holotype of T. yueliangensis sp. nov.

FIGURE 10. Distribution of Tonsilla species treated here.