Tenomerga mucida ( Chevrolat, 1844 )

Yoshitomi, Hiroyuki, 2024, Revision of Tenomerga (Coleoptera, Cupedidae) of Japan and adjacent regions, Zootaxa 5555 (2), pp. 201-226 : 203-212

Tenomerga mucida ( Chevrolat, 1844)

[Japanese name: Nagahiratamushi]

Figs 1A, B , 2A , 3A, B , 4A , 5 , 6 , 7 , 13A ; Table 1

Cupes mucida Chevrolat, 1844: 58 .

Cupes mucidus : Atkins 1963: 154.

Tenomerga mucida : Neboiss 1984: 453; Miyatake 1985a: 21; 1985b: 1, pl. 1, 1 [figure and short description]; Lawrence et al. 2021: 464, fig. 6C [hindwing venation]; Park & Cho 2023: 33 [recorded from Korea].

Cupes clathratus Solsky, 1871: 370 . Tamanuki 1928: 250; Yabe 1929: 61; Nakane 1973: 2. (Synonymized with Tenomerga mucida ( Chevrolat, 1844) by Neboiss 1984).

Cupes ocularis Pascoe, 1872: 319 . (synonymized with Cupes clathratus Solsky, 1871 by Harold 1877)

Cupes clathratus var. fuscus Tamanuki, 1928: 251 . (Synonymized with Tenomerga mucida ( Chevrolat, 1844) by Neboiss 1984).

Type specimen examined. Holotype of Cupes clathratus var. fuscus Tamanuki, 1928 ( Fig. 2A ): male ( SEHU), “ Japan Matsumura”, “ Cupes clathratus var. fuscus Tam. Type”, “ Tenomerga mucida (Chevrolat) det. Neboiss 1983”, “0000001300 Sys.Ent Hokkaido Univ. Japan [ SEHU]”.

Specimens examined. [ JAPAN] <Hokkaido> 1 male, 2 females, ( SEHU), Date , no data, M. Matsushita leg. ; 1 female, ( SEHU), Jozankei, Sapporo-shi , 17. IX. 1954, T. Kumata leg. ; 1 male, ( SEHU), Jozankei, Sapporo-shi , 25. VIII. 1954, M. Konishi leg. ; 2 females, ( SEHU), Jozankei, Sapporo-shi , 7. VIII. 1987, Y. Nishijima leg. ; 1 male, ( NSMT), Kitami-Aioi , 8. VIII. 1950, S. Shibanai leg. ; 1 male, ( KUM), Maruyama, Sapporo-shi , 31. VII–4. VIII. 2000, S. Ogata leg. ; 1 female, ( NSMT), Poroshiri-dake, Hidaka , 30. VII. 1971, J. Aoki leg. ; 1 female, ( SEHU), Sakkuru , 7. VII. 1950, T. Nakashima leg. ; 1 female, ( SEHU), Sapporo-shi , 1. VIII. 1918, S. Matsumura leg. ; 1 female, ( SEHU), Sapporo-shi , 18. VIII. 1971, S. Matsumura leg. ; 1 male, ( SEHU), Sapporo-shi , 26. VII. 1984, K. Furukawa leg. ; 2 males, ( SEHU), Sapporo-shi , 3, VIII. 1918, S. Matsumura leg. ; 1 male, ( NSMT), Sapporo-shi , 3. VIII. 1924 ; 1 female, ( SEHU), Sapporo-shi , 31. VII. 1980 (A. Tanaka coll. 2005) ; 2 females, ( SEHU), Sapporo-shi , 9. VIII. 1987 (A. Tanaka coll. 2005) ; 1 female, ( SEHU), Sapporo-shi , VIII. 1907 ; 1 male, ( NSMT), Shumarinai, Sorachi-shicho , 2–3. VIII. 1982, S. Tsuyuki leg. ; 1 male, ( SEHU), Soranuma-dake , 9. VIII. 1962, T. Kumata leg. (Nakane coll. 1999) ; 1 male, ( EUMJ), Teinekanayama, Sapporo-shi , 17. VIII. 2022, Y. Omatsu leg. ; 1 female, ( SEHU), Tomuraushi , 26. VIII. 1954, S. Takagi leg. ; 1 male, ( SEHU), Toyotaki, Sapporo-shi , 18. VIII. 1979, T. Fujisawa leg. ; 1 male, ( SEHU), Yamahana, Sapporo-shi , 7. VIII. 1903 ; 1 female, ( NSMT), Yasukuni, Abashirishicho , 30–31. VII. 1982, S. Tsuyuki leg. < Aomori Pref.> 1 male, ( EUMJ), Hachinohe , 9. VIII. 1938, A. Fukuda leg. ; 1 female, ( NSMT), Takinomata, Hirakawa-shi , 18. VIII. 1952, K. Shimoyama leg. < Yamagata Pref.> 1 male, ( SEHU), Sakata , 2. VII. 1948, T. Nakane leg. (Nakane coll. 1999) ; 1 male, ( SEHU), Sakata , 7. VII. 1948, K. Shirahata leg. (Nakane coll. 1999) ; 2 males, ( SEHU), Sakata , 7. VII. 1948, K. Shirahata leg. (Nakane coll. 1999) ; 1 female, ( NSMT), Yonezawa , 21. VIII. 1941, Y. Kurosawa leg. ; 1 female, ( EUMJ), Nakamaru, Nagasaka-shi , 17. VII. 1984, S. Nomura leg. < Iwate Pref.> 2 males, ( EUMJ), Genpeidaira, nr. Hayachine , 31. VI. 1983, Y. Notsu leg. ; 2 males, ( EUMJ), Genpeidaira, nr. Hayachine , 31. VII. 1982, N. Ohbayashi leg. < Miyagi Pref.> 1 male, ( KUM), Makiyama, Ishinomaki-shi , 30. VII. 1995 ; 1 male, ( EUMJ), Shironuma, Zaou , 19. VIII. 1978, M. Sato leg. < Fukushima Pref.> 1 female, ( EUMJ), Tateiwa-mura , 25–27. VII. 1980, Y. Notsu leg. ; 1 male, ( EUMJ), Tateiwa-mura , 28. VII. 1984, K. Nagata leg. ; 1 female, ( NSMT), Wakamatsu , 10. V. 1950, Y. Kurosawa leg. ; 1 female, ( EUMJ), Yugura, Onuma-gun , 5. VIII. 1974, J. Yokoi leg. < Chiba Pref.> 1 male, ( NSMT), Katori, Katori-shi , 7. VII. 1935, T. Tani leg. ; 1 male, ( EUMJ), Nishizaki, Tateyama-shi , 7. VII. 1979, S. Maru leg. < Saitama Pref.> 1 male, ( NARO), Shimura , 24. VII. 1949, E. Yoshida leg. ; 1 male, ( NARO), Shimura , 28. VII. 1949, Y. Hirano leg. ; 1 male, ( NARO), Yono , 8. VII. 1954, Y. Hirano leg. < Gunma Pref.> 1 male, ( SEHU), Numata , 5. VIII. 1950, S. Ueno leg. (Nakane coll. 1999) . < Tochigi Pref.> 2 females, ( NARO), Yumoto, Nikko-shi , 10. VII. 1949, M. Okamoto leg. < Ibaraki Pref.> 1 female, ( EUMJ), Migimomi, Tsuchiura-shi , 10. VII. 1979, S. Ohmomo leg. ; 1 female, ( NSMT), Mito-shi , 23. VII. 1978, T. Tani leg. < Kanagawa Pref.> 1 male, ( NARO), Fukazawa, Tanzawa , 24. VIII. 1986, Y. Hirano leg. ; 1 female, ( EUMJ), Hiratsuka , 6. VII. 1978, Y. Notsu leg. ; 1 male, ( NARO), Komayama, Ooiso-shi , 21. VI. 1996, S. Tsuyuki leg. ; 1 male, ( NARO), Kurokura, Tanzawa , 22. VIII. 1987, Y. Hirano leg. ; 1 male, ( NARO), Manazuru , 23. VII. 1983, Y. Hirano leg. ; 1 male, ( NARO), Mt. Kohbo, Hakone , 18. VII. 1982, T. Maenami leg. ; 1 female, ( EUMJ), Nakai-machi , 19. VII. 1980, Y. Notsu leg. ; 1 male, ( EUMJ), Ninomiya , 18. VII. 1979, N. Ohbayashi leg. ; 1 male, ( NARO), Odawara , 14. VII. 1956, Y. Hirano leg. ; 1 female, ( NARO), Odawara , 3. VIII. 1982, Y. Hirano leg. ; 1 female, ( NARO), Ooiso , 30. VII. 1973, Y. Hirano leg. < Tokyo Met .> 1 male, ( KMNH), Amanuma, Suginami-ku , 20. VI. 1934, N. Gyotoku leg. ; 1 male, ( KMNH), Amanuma, Suginami-ku , 20. VII. 1932, N. Gyotoku leg. LT; 1 male, ( KMNH), Amanuma, Suginami-ku , 6. VII. 1934, N. Gyotoku leg. ; 2 females, ( KMNH), Fujimidai, Meguro-ku , 14. VII. 1975, T. Shibuya leg. ; 1 male, ( NSMT), Kinuta , 20. VII. 1962, Y. Kurosawa leg. ; 1 female, ( SEHU), Kiyokawa, Taito-ku , 13. VII. 1974, T. Ichikawa leg. (Nakane coll. 1999) ; 1 female, ( KMNH), Nakameguro, Meguro-ku , 6. VIII. 1939, T. Shibuya leg. ; 1 female, ( NSMT), Nippara, Okutama , 17–18. X. 1976, Y. Kurosawa leg. ; 1 male, ( NARO), Nishigahara , 25. VII. 1948, I. Hattori leg. ; 1 male, ( SEHU), no data, 2–10. VIII. 1933, S. Matsumura leg. ; 1 female, ( SEHU), S. Matsumura leg. ; 1 male, ( NARO), no data, VIII. 1918 ; 1 male, ( NSMT), Ooi-machi , 10. VII. 1940, K. M. leg .; 1 male, ( NSMT), Ooi-machi , 24. VII. 1940, K. M. leg .; 1 male, ( NARO), Setagaya , 10. VII. 1950, I. Hattori leg. ; 1 male, ( NARO), Setagaya , 26. VII. 1951, I. Hattori leg. ; 1 male, ( NARO), Shiba , 18. VII. 1967, S. Kinoshita leg. ; 2 males, ( EUMJ), Suginami , 20. VII. 1952, S. Hisamatsu leg. ; 1 male, ( EUMJ), Suginami , 26. VII. 1952, S. Hisamatsu leg. ; 1 female, ( SEHU), Takao-san , 3–4. VII. 1997, M. Maruyama leg. leg.; 1 male, ( NSMT), Ushigome , 5. VII. 1933, T. Wakamori leg. < Izu Isls. : Miyake-jima > 1 male, ( EUMJ), Ako (34°03’N 139°29’E), 7–30. VII. 2008, H. Makihara leg. MT; 1 male, ( EUMJ), Hinoyama-shita , 10– 11. VII. 2008, H. Makihara leg. ; 1 male, ( EUMJ), Tsubota-rindo , 10–11. VII. 2008, H. Makihara leg. < Izu Isls. : Mikura-jima > 1 male, ( EUMJ), 14. VII. 1986, K. Shimizu leg. ; 1 male, ( NSMT), 9–12. VII. 1974, K. Kinugasa leg. ; 2 males, ( SEHU), Kawada , 2/ 3. V. 1974 (larva), 21/ 30. VI. 1974 (Imago), T. Ichikawa leg. (Nakane coll. 1999) ; 1 male, ( EUMJ), Sato , 14. VII. 1973, S. Nagao leg. < Ogasawara Isls .> 1 male, ( NSMT), Chichijima , 11. VI. 1976, Y. Kurosawa leg. ; 1 male, ( EUMJ), Hahajima , 15–17. VI. 1985, H. Makihara leg. ; 1 male, ( SEHU), Hyogidaira, Hahajima , 20. VI. 1984, H. W. leg. (Nakane coll. 1999) . < Niigata Pref.> 1 female, ( KUM), Kurokawa , 10. V. 1972, K. Baba leg. ; 1 female, ( KUM), Kurokawa , 10. VII. 1977, K. Baba leg. ; 1 male, ( EUMJ), Kurokawa , 13. VII. 1975, K. Baba leg. ; 1 female, ( KUM), Kurokawa , 2. VIII. 1954, K. Baba leg. ; 1 male, ( EUMJ), Kurokawa , 20. VII. 1957, K. Baba leg. ; 1 female, ( KUM), Kurokawa , 20. VII. 1961, K. Baba leg. ; 1 female, ( KUM), Kurokawa , 21. VIII. 1954, K. Baba leg. ; 1 female, ( KUM), Kurokawa , 23. VII. 1954, K. Baba leg. ; 1 male, ( EUMJ), Kurokawa , 26. VII. 1956, K. Baba leg. ; 1 male, ( EUMJ), Miomote , 25. VIII. 1973, A. Seino leg. ; 1 female, ( KUM), Momozaki-hama , 10. VII. 1957, K. Baba leg. ; 1 male, ( EUMJ), Murakami , 6. VIII. 1961, M. Sato leg. ; 1 male, ( SEHU), Nagaoka , 27. VII. 1946, T. Nakane leg. (Nakane coll. 1999) . < Shizuoka Pref.> 1 male, ( SEHU), Gotemba , 8. 1934, K. Nakamura leg. (Nakane coll. 1999) ; 1 female, ( KUM), Shimotogari , 10. VII. 1973, M. Chujo leg. < Nagano Pref.> 1 female, ( SEHU), Kamikochi , 14. VII. 1918, Issiki leg. ; 1 male, ( SEHU), Karuizawa , VII. 1979 (A. Tanaka coll. 2005) ; 1 male, ( NSMT), Nyugasa-yama , 3. V. 1960, Y. Yoshida leg. < Aichi Pref.> 1 female, ( EUMJ), Dando , 20. VII. 1969, M. Sato leg. ; 1 female, ( EUMJ), Hagiyama, Mizuho-ku, Nagoya-shi , 20. VII. 1977, Y. Hori leg. ; 1 female, ( EUMJ), Higashi-ku, Nagoya-shi , 18. VII. 1965, Z. Naruse leg. ; 1 female, ( EUMJ), Higashisugi, Nagoya-shi , 12. VII. 1964, H. Yamada leg. ; 1 female, ( EUMJ), Higashiyama, Nagoya-shi , VII. 1971, M. Sato leg. figured by Miyatake, 1985b: 1, pl. 1, 1; 1 male, ( EUMJ), Kamagahora, Midori-ku, Nagoya-shi , 5. VI. 2002, M. Sato leg. ; 1 male, ( EUMJ), Matsukawabashi, Nagoya-shi , 26. VII. 1973, M. Sato leg. ; 1 male, ( EUMJ), Midori-ku, Nagoya-shi , 6. VII. 2004, M. Sato leg. ; 1 female, ( EUMJ), Nagoya-shi , 17. VIII. 1963, N. Ohbayashi leg. ; 1 male, ( EUMJ), Nagoya-shi , 6. VII. 1958, N. Ohbayashi leg. ; 1 male, ( EUMJ), Shimada, Tempaku-ku, Nagoya-shi , VII. 1991, H. Yoshitomi leg. ; 1 male, ( SEHU), Shindeki-machi, Nagoya-shi , 5. VII. 1941, S. Osawa leg. (Nakane coll. 1999) ; 1 male, ( SEHU), Shirakabe, Nagoya-shi , 16. VII. 1942, T. Horiuti leg. (Nakane coll. 1999) ; 1 male, ( EUMJ), Shitara-cho , 8–12. VII. 2002, S. Nagashima leg. < Gifu Pref.> 1 female, ( SEHU), Kinkazan , 5. VIII. 1966, T. Nakane leg. (Nakane coll. 1999) . < Fukui Pref.> 1 male, ( KUM), Aoba-yama , 21–24. VII/ 1974, H. Sasaji leg. ; 1 male, ( SEHU), Ikeizumicho, Chikuzen-shi , 12. VII. 1961, K. Fukuda leg. (Nakane coll. 1999) ; 1 male, ( KUM), Mikuni , 8–10. VII. 1974, H. Sasaji leg. ; 1 male, 1 female, ( KUM), Sanri-yama , 26. VII. 1974, H. Sasaji leg. < Ishikawa Pref.> 1 male, ( KUM), Kanazawa-jo, Kanazawa-shi , 16. VII. 1968 . < Toyama Pref.> 1 female, ( KUM), Senjugahara, Tateyama-machi , 1–2. VIII. 1981, K. Baba leg. < Kyoto Pref.> 1 male, ( EUMJ), Kyoto-shi , 14. VII. 1957, M. Ohtake leg. < Shiga Pref.> 1 male, ( SEHU), Hiei-zan , 22. VIII. 1957, H. Ishida leg. (Nakane coll. 1999) . < Wakayama Pref.> 1 male, ( NARO), Nachi-san , 10. V. 1954, T. Fujimura leg. ; 1 male, ( SEHU), Shingu , 12. VII. 1946, S. Osawa leg. (Nakane coll. 1999) . < Hyogo Pref.> 1 male, ( EUMJ), Akashi-shi , 13. VII. 1973 ; 1 female, ( EUMJ), Akashi-shi , 24. VII. 1970, M. Tomokuni leg. ; 1 female, ( SEHU), Nishinomiya , 13. VII. 1947, K. Sakaguchi leg. (Nakane coll. 1999) ; 1 male, ( SEHU), Rokko , 3. VII. 1948, N. Yano leg. (Nakane, 1963, fig. 1-1) . < Tottori Pref.> 1 female, ( NSMT), Daisen , 22. VII. 1940 ; 1 female, ( EUMJ), Daisen , 31. VII. 1975, Y. Notsu leg. < Kagawa Pref.> 1 female, ( EUMJ), Hanazono-cho, Takamatsu-shi , 22. VI. 1966, M. Takagi leg. ; 1 male, ( EUMJ), Sakamoto , 5. VII. 1948, T. Ko leg. ; 1 male, ( EUMJ), Yashima , 13. VII. 1965, M. Takagi leg. ; 1 female, ( KUM), Yashima , 19. VII. 1973, M. Chujo leg. ; 1 male, ( EUMJ), Zozu-san , 16. VII. 1978, M. Sakai leg. ; 1 male, ( EUMJ), Zozu-san , 16. VII. 1978, S. Hisamatsu leg. < Tokushima Pref.> 1 male, ( EUMJ), Bizan , 4. VII. 1923, M. Yoshida leg. ; 1 male, ( EUMJ), Ishima, Anan-shi , 30. VII. 2002, Y. Kamite leg. ; 1 female, ( EUMJ), Nyodo-cho, Tokushima-shi , 27. VII. 1973, K. Yoshida leg. ; 1 male, ( EUMJ), Otaki-san , 2. VIII. 1972, M. Sakai leg. ; 1 male, ( EUMJ), Otaki-san , 2. VIII. 1972, M. Yoshida leg. < Ehime Pref.> 1 male, ( EUMJ), Dogo, Matsuyama-shi , 7. VII. 1950, T. Yano leg. ; 1 male, ( EUMJ), Edamatsu, Matsuyama-shi , 12. VI. 1994, K. Aita leg. ; 1 male, ( EUMJ), Gosho, Yamanouchi, Toon-shi , 7. VII. 1987, Y. Utsunomiya leg. ; 1 female, ( EUMJ), Ishitegawa dam, Matsuyama-shi , 30. VII. 1976, A. Oda leg. ; 1 male, ( OMM), Ishizuchi-san, Kumakogen-cho , 29. V. 1989, Y. Utsunomiya leg. ; 1 male, ( EUMJ), Komenono, Matsuyama-shi , 24. VI. 1979, I. Amano leg. ; 1 male, ( EUMJ), Kono-mura, nr. Matsuyama-shi , 21. VII. 1954, T. Yano leg. ; 1 female, ( EUMJ), Kuwabara, Matsuyama-shi , 30. VI. 1978, S. Suwa leg. ; 1 female, ( EUMJ), Matsuyama-jo, Matsuyama-shi , 24. VII. 1999, T. Inoue leg. ; 1 male, ( EUMJ), Matsuyama-jo, Matsuyama-shi , VII. 2022, N. Isa leg. ; 2 males, ( EUMJ), Matsuyama-shi , 10. VII. 1974, M. Miyatake leg. ; 1 female, ( EUMJ), Matsuyama-shi , 18. VII. 1966, T. Ishihara leg. ; 1 male, ( EUMJ), Matsuyama-shi , 2. VIII. 1956, S. Hisamatsu leg. ; 1 female, ( EUMJ), Matsuyama-shi , 29. VI. 2022, Li & Shiiba leg. ; 1 female, ( EUMJ), Matsuyama-shi , 3. VIII. 1955, S. Hisamatsu leg. ; 2 females, ( EUMJ), Matsuyama-shi , 8. VII. 1949, T. Ishihara leg. ; 1 male, ( EUMJ), Matsuyama-shi , VII. 1961, T. Ishikawa leg. ; 1 male, ( EUMJ), Momiji-machi, Matsuyama-shi , 3. VII. 1995, K. Okada leg. ; 1 female, ( OMM), Nojiri, Kumakogen-cho , 29. VII. 2012, S. Yano leg. LT; 1 male, ( EUMJ), Oda-cho , 15. VII. 1998, E. Yamamoto leg. ; 1 male, ( EUMJ), Omogokei , 4. VIII. 1952, T. Koyama leg. ; 1 female, ( EUMJ), Shiratsue-Myojingamori, Matsuyama-shi , 22. VII. 1972, S. Kinoshita leg. LT; 1 female, ( EUMJ), Sue-machi, Tobe-cho , 2. VII. 2015, K. Yoshida leg. ; 1 male, ( EUMJ), Sugitate, Matsuyama-shi , 2. VIII. 1976, N. Yashiro leg. ; 1 male, ( EUMJ), Sute-machi, Tobe-cho , 2. VII. 2015, K. Yoshida leg. ; 1 male, ( EUMJ), Takanawa-san, Matsuyama-shi , 11–12. VIII. 2015, K. Kuroda leg. LT; 1 male, ( EUMJ), Takanawa-san, Matsuyama-shi , 16. VIII. 1979, H. Kan leg. ; 1 female, ( EUMJ), Takanawa-san, Matsuyama-shi , 2. VIII. 2015, H. Yoshitomi leg. ; 1 male, ( EUMJ), Takanawa-san, Matsuyama-shi , 20. VII. 1998, H. Kan leg. ; 1 female, ( EUMJ), Tarumi, Matsuyama-shi , 1. VII/ 2008, Y. Sato leg. ; 1 female, ( EUMJ), Tarumi, Matsuyama-shi , 23. VI. 2022, R. Akeo leg. ; 1 female, ( EUMJ), Tarumi, Matsuyama-shi , 3. VII. 1999, S. Hayakawa leg. ; 1 female, ( EUMJ), Tarumi, Matsuyama-shi , 30. VI. 2022, R. Akeo leg. ; 1 female, ( EUMJ), Tarumi, Matsuyama-shi , 4. VII. 2021, H. Kono leg. ; 1 female, ( EUMJ), Toon-shi , 12. VI. 1983, M. Sakai leg. ; 1 female, ( EUMJ), Toon-shi , 17. VII. 1988, M. Sakai leg. ; 1 male, ( EUMJ), Toon-shi , 20. VII. 1987, M. Sakai leg. ; 2 males, ( EUMJ), Top of Sugitate, Matsuyama-shi , 9. VIII. 1983, H. Kan leg. LT; 1 male, ( EUMJ), Yokogawara , 12. VII. 1995, M. Sakai leg. ; 1 female, ( EUMJ), Yokogawara , 20. VIII. 1999, M. Sakai leg. LT; 1 male, ( EUMJ), Yokogawara, Toon-shi , 30. VII. 1990, M. Sakai leg. LT; 1 male, ( EUMJ), Yuwatari, Matsuyama-shi , 24. VI. 1979, Y. Seiyama leg. LT; 1 male, ( EUMJ), Yuyamayanagi, Matsuyama-shi , 2. VII. 2003, T. Inoue leg. ; 1 female, ( OMNH), Nojiri, Kumakogen-cho , 29. VII. 2012, S. Yano leg. < Kochi Pref.> 1 female, ( EUMJ), Asakura, Kochi-shi , 17. VII. 1980, A. Ishida leg. ; 1 female, ( EUMJ), Asakuraotsu , 6. VIII. 1978, R. Shimamoto leg. LT; 1 male, ( EUMJ), Hirohata, Iburi, Tosashimizu-shi , 17–18. VII. 2000, K. Aita leg. LT; 1 female, ( EUMJ), Kuroson , 4–7. VII. 1966, N. Ohbayashi leg. ; 1 female, ( EUMJ), Kuroson , 6–10. VIII. 1966, K. Iwao leg. ; 1 male, ( EUMJ), Okinoshima , 11. VIII. 1951, T. Esaki leg. < Fukuoka Pref.> 1 female, ( KMNH), Adachi-yama , 13. VIII. 1966, K. Adachi leg. ; 1 male, ( KUM), Ainoshima, Shingu-cho , 14. VII. 1991, K. Kido leg. ; 1 male, ( KUM), Bunda, Shinkawa, Ukiha-cho , Ukiha-shi , 24. VII–5. VIII. 2022, K. Kido leg. FIT; 1 male & 1 female, ( KUM), Bunda, Shinkawa, Ukiha-cho , Ukiha-shi , 24–29. VII. 2022, K. Kido leg. LT; 1 male, ( KUM), Bunda, Shinkawa, Ukiha-cho , Ukiha-shi , 27. VI–2. VII. 2022, K. Kido leg. LT; 1 female, ( KUM), Bunda, Shinkawa, Ukiha-cho , Ukiha-shi , 2–8. VII. 2022, K. Kido leg. LT; 1 male, ( KMNH), Chikujo, Chikujo-machi , 18. VI. 1987, Y. Takakura leg. ; 1 male, ( KUM), Ebitsu, Okagaki-machi , 8. VII. 1972, T. Ishikawa leg. ; 1 female, ( KMNH), Fujimatsu, Moji-ku , 15. VII. 1967, M.Amano leg. ; 1 female, ( KMNH), Fukakura, Hiko-san , 11. VIII. 1981, M. Amano leg. ; 3 males, 2 females, ( EUMJ), Fukuchi-yama , 4. VIII. 1954, S. Kimoto leg. ; 1 female, ( KUM), Hakomatsu, Higashi-ku, Fukuoka-shi , 16. VII. 1996, H. Kojima leg. ; 1 female, ( NSMT), Hakozaki , 3. VII. 1946, Y. Kurosawa leg. ; 1 female, ( EUMJ), Hiko-san, Soeta-machi , 1. VIII. 1956, S. Kimoto leg. ; 1 female, ( NSMT), Hiko-san, Soeta-machi , 2. VIII. 1946, Y. Takakura leg. ; 1 female, ( KMNH), Hiko-san, Soeta-machi , 21. VII. 1958, M. Amano leg. ; 1 male, ( KMNH), Hinago, Chikujo-machi , 11. VII. 1987, Y. Takakura leg. ; 1 female, ( KMNH), Ishitsubo-cho, Yahata , 25. VII. 2007, K. Ueda leg. ; 1 female, ( KMNH), Kamifujimatsu, Moji-ku, Kitakyushu-shi , 27. VII. 1999, M.Amano leg. ; 1 male, ( KMNH), Kashiwara, Fukuoka-shi , 6. VII. 1978, N. Gyotoku leg. ; 1 female, ( KMNH), Koga, Asakura-shi , 10. VIII. 1954, N. Gyotoku leg. ; 1 male, ( KUM), Konosu-yama, Minami-ku, Fukuoka-shi , 1. VII. 1971, T. Ishikawa leg. ; 1 male, ( KMNH), Kugumiya , 10. VII. 1942, N. Gyotoku leg. ; 1 male, ( KMNH), Kugumiya, Asakura-shi , 16. VII. 1950, N. Gyotoku leg. ; 1 male, ( KMNH), Kugumiya, Chikuzen , 13. VII. 1944, N. Gyotoku leg. ; 1 female, ( KMNH), Miwa , 17. VII. 1970, K. Miyoshi leg. ; 1 female, ( KMNH), Miwa , 19. VII. 1972, K. Miyoshi leg. ; 1 male, ( KMNH), Shimoshin-nyo, Nogata-shi , 16. VI. 2021, Y. Minoshima leg. LT; 1 male, ( KUM), Shiroyama, Munakata-shi , 4. VIII. 1985, K. Kido leg. ; 1 male, ( SEHU), Shojidake , 21. VII. 1961, S. Nakao leg. (Nakane coll. 1999) ; 1 female, ( KMNH), Tabuse , 9. VII. 1988, K. Miyoshi leg. ; 1 female, ( KMNH), Taga , 26. VII. 1986, K. Miyoshi leg. ; 1 female, ( KMNH), Takasu-yama , 25. VII. 1977, Y. Takakura leg. ; 1 male, ( KMNH), Tanushimaru, Kurume-shi , 28. VII. 1935, I. Tateishi leg. ; 1 male, ( KMNH), Toyama-jinja, Kawasaki-cho , 2. VII. 1986, Y. Takakura leg. ; 1 male, ( KUM), Wakasugi-yama, Sasaguri-machi , 7. VIII. 1967, K. Kido leg. ; 1 female, ( KMNH), Yoshii-machi, Ukiha-shi , 6. VIII. 1958, N. Gyotoku leg. ; 1 male, ( KMNH), Yoshiki, Chikushino-shi , 17. V. 1993, N. Gyotoku leg. LT; 1 female, ( KMNH), Yoshikuni , 2. VII. 1929 . < Nagasaki Pref.> 1 female, ( KUM), Shimabara , 14. VII. 1977, Fukunaga leg. < Tsushima > 1 female, ( EUMJ), Ariake-san , 25–28. VII. 1979, Y. Seiyama leg. < Kumamoto Pref.> 1 female, ( EUMJ), Kadoyama, Amakusa-shi , 31. VII. 1973, M. Miyahara leg. ; 1 female, ( NSMT), Kyoragi, Amakusa-Kamishima , 6. VII. 1980, H. Irie leg. < Oita Pref.> 1 male, ( KUM), Kuenhira-yama, Kokonoe-cho , 12. VIII. 2004, S. Ogata leg. LT; 1 male, ( KUM), Kuenhira-yama, Kokonoe-cho , 15. VIII. 2003, S. Ogata leg. LT; 1 male, ( KUM), Kuenhira-yama, Kokonoe-cho , 20. VIII. 2002, S. Ogata leg. LT; 1 male, ( KUM), Kuenhira-yama, Kokonoe-cho , 22. VIII. 1998, S. Ogata leg. LT; 1 male, ( EUMJ), Sobo-san , 11. VIII. 1962, M. Amano leg. ; 1 female, ( KMNH), Sobo-san , 19. VI. 1939, N. Gyotoku leg. ; 1 male, ( KMNH), Sobo-san , 20. VI. 1938, N. Gyotoku leg. ; 1 female, ( KUM), Sobo-san , 22. VII. 1938, H. Tagawa leg. ; 1 female, ( KMNH), Sobo-san , 23. VII. 1948, M. Amano leg. < Miyazaki Pref.> 1 female, ( KUM), Aburatsu , 17. VI. 1946, T. Mitono leg. LT; 1 female, ( SEHU), Hosomi-cho, Nobeoka-shi , 5. VIII. 1983, S. Masuda leg. (Nakane coll. 1999) ; 1 female, ( EUMJ), Kirishima-onsen , 21. VII. 1968, K. Suga leg. ; 1 female, ( SEHU), Omata , 17. VII. 1954, K. Nohara leg. (Nakane coll. 1999) . < Kagoshima Pref.> 1 male, ( SEHU), Kurino , 30. VII. 1983, T. & T. Nakane leg. (Nakane coll. 1999) ; 1 male, ( SEHU), Togo , 12. VII. 1985, Sk. Yamane leg. ; 1 male, ( SEHU), Toso , 29. VI. 1983, Y. Takai leg. (Nakane coll. 1999) . < Yakushima > 1 male, ( SEHU), Miyanoura , 20. VII. 1971, T. Hatayama leg. (Nakane coll. 1999) ; 1 male, ( EUMJ), Miyanoura , 23. VI. 1975, Y. Kurosawa leg. ; 1 female, ( SEHU), Shiratani , 24. VII. 1981, M. Miura leg. (Nakane coll. 1999) . < Amami-Oshima > 1 male, ( EUMJ), Nazesato , 13. VI. 2020, Fukida leg. < Ryukyu : Okinoerabu-jima > 1 male, ( KUM), Ooyama-shokubutu-koen, 23–26. VI. 2022, S. Imada leg.

[ CHINA] 1 male, ( TARI), Houchai, 11. VI. 2015, Y.-T. Chung leg.

[ TAIWAN] 1 female, ( NSMT), "Akazayazaya", 17. III. 1927, T. Kano leg. ; 1 male, ( TARI), Holisha, 20. V. 1919 .

[data unclear] 1 female, ( SEHU), no data ; 1 male, ( NSMT), "Furjai", 4. VIII. 1921 ; 1 male, ( NARO), no data, 20 VII. 1919, Go leg. ; 1 female, ( NARO), no data ; 1 male, 2 females, ( EUMJ), no data ; 1 female, ( SEHU), no data, T. Otsuka leg. ; 1 female, ( SEHU), Sanbongi, 8. VIII. 1919 .

Diagnosis. Color of body blackish-brown, generally monocolor. Scales on head, pronotum, scutellar shield, and elytra black, but cream-colored on a part of head, anterior part of pronotum, posterior part of intervals V and VII, and apical part of elytra. Elytra ( Fig. 4A ) with 9 longitudinal rows of window punctures; window punctures round to square.

Male ( Fig. 1A ). Eyes ( Fig. 3A ) large, strongly prominent. Antennae long and slender, reaching about proximal 3/4 of elytra; AL/TL 0.64–0.88 (0.81); approximate ratio of each antennomere (n = 10; average) as 3.37: 1.00: 3.50: 4.14: 4.73: 5.17: 5.47: 5.55: 5.49: 5.36: 7.03. Anterior pronotal angles ( Fig. 3A ) acute. HL/HW 0.53–0.62 (0.58); PW/PL 1.25–1.47 (1.35); EL/EW 2.73–2.96 (2.83); EL/PL 5.54–5.94 (5.73); EW/PW 1.43–1.57 (1.51); TL/ EW 3.60–3.92 (3.75); EYL/EYW 1.68–2.33 (1.91); TEL/HL 0.10–0.19 (0.15). Apex of sternite IX ( Fig. 6C ) simply pointed. Bifurcate process ( Fig. 7F ) with short median strut (Y-shaped in Hörnschemeyer, 2009). Parameres ( Fig. 7A ) with posterior projection of dorsal margin (pp) triangular; apical hook ( Fig. 7E ) bi-pointed; ventro-marginal spines (vms) long, reaching apical hook. Apical margin of dorsal plate (dp, Fig. 7B ) deeply notched. Apical margin of mesal lobe (ml, Fig. 7B ) clearly excised.

Female ( Fig. 1B ). Eyes ( Fig. 3B ) small. Antennae short and stout, reaching about proximal 1/2 of elytra; AL/TL 0.59–0.65 (0.63); approximate ratio of each antennomere (n = 10; average) as 3.40: 1.00: 3.47: 3.80: 4.16: 4.38: 4.43: 4.38: 4.41: 4.22: 5.82. Anterior angles of pronotum ( Fig. 3B ) acute. HL/HW 0.61–0.73 (0.67); PW/PL 1.29–1.53 (1.39); EL/EW 2.73–2.94 (2.81); EL/PL 5.71–6.50 (6.06); EW/PW 1.47–1.63 (1.55); TL/EW 3.58–3.88 (3.69); EYL/EYW 1.97–2.29 (2.09); TEL/HL 0.19–0.25 (0.21).

Distribution. Japan: Hokkaido, Honshu, Shikoku, Kyushu, Oki, Tsushima, Yakushima, Izu Isls. (Miyake-jima, Mikura-jima), Ogasawara Isls. (Chichi-jima, Haha-jima), Amami-Oshima, Okinoerabu-jima; Korea, China, Caroline Is., the Philippines, East Siberia, Hawaii.

Remarks. This is the first record of this species from Taiwan, Amami-Oshima, and Okinawa-jima.

Biological notes. The larva and pupa of this species were described by Fukuda (1938, 1939a, b) and Yavorskaya et al. (2015). The larvae were collected from rotten wood of cherry tree ( Fukuda 1938), brown-rottened logs of a willow ( Yavorskaya et al. 2015), and pine tree stump affected by brown-rot fungus ( Fukuda 1941). I collected many larvae of this species from a dead branches of cherry trees affected by white rot fungus (Kaizu-cho, Gifu Pref., 15. XII. 1991).

Adults are frequently attracted to light, and they are also collected from the trunks of withered standing trees at night. Nomura (2017) reported that many adults of this species were collected from inside the warehouse.

Ohno (1974) reported that the adults of this species stridulate by rubbing their hind wings and elytra together, producing a sound like 'tsk, tsk, tsk, tsk....'.

Figures

FIGURE 1. Habitus of Tenomerga spp. A, B) Tenomerga mucida, C, D) T. anguliscutis, E, F) T. japonica, G, H) T. trabecula, I, J) T. kurosawai, K, L) T. yamato. A, C, E, G, I, K) males; B, D, F, H, J, L) females. I) holotype, J) paratype. Scale bars = 5.0 mm.

FIGURE 2. Holotypes and associated labels for Tenomerga spp. A) Cupes clathratus var. fuscus Tamanuki, 1928, B) Cupes formosanus Tamanuki, 1928, C) Cupes japonicus Tamanuki, 1928, D) Tenomerga yamato Miyatake, 1985.

FIGURE 3. Heads and pronota of Tenomerga spp. A, B) Tenomerga mucida, C, D) T. anguliscutis; E, F) T. japonica, G, H) T. trabecula, I, J) T. kurosawai, K, L) T. yamato. A, C, E, G, I, K) males; B, D, F, H, J, L) females. I) holotype, J) paratype. Scale bars = 1.0 mm.

FIGURE 4. Dorso-lateral views of elytra of Tenomerga spp. A) Tenomerga mucida, B) T. anguliscutis, C) T. japonica, D) T. trabecula, E) T. kurosawai, F) T. yamato. Scale bars = 1.0 mm.

FIGURE 5. Mouthparts of Tenomerga mucida. A) labrum, dorsal view, B) labium, dorsal view, C) left maxilla, ventral view, D) labial palpus, E) right mandible, ventral view, F) left mandible, dorsal view. dls—dorso-lateral sensory patch, ga— garea, la—lacinia, pa—palpifer, sp—sensory patch.

FIGURE 6. Male (A–C) and female (D–F) genitalia of Tenomerga mucida. A) tergite VIII, B) sternite VIII, C) stertnite IX (9S) and tergite IX (9T), D) sternite VIII, E) tergite VIII, F) ovipositor. st1—stylus I, st2—stylus II, co—coxite, va—valvifer.

FIGURE 7. Male genitalia of Tenomerga mucida. A) tegmen, dorsal view, B–D) penis, dorsal (B), ventral (C) and lateral (D) views, E) apical hook, lateral view, F) bifurcate process. ah—apical hook, ap—anterior projection of dorsal margin, bo—basal opening, dp—dorsal plate, ml—mesal lobe, pp—posterior projection of dorsal margin, vms—ventro-marginal spine.

FIGURE 13. Distribution maps of Tenomerga spp. A) Tenomerga mucida, B) T. anguliscutis, C) T. japonica (rhombus) and T. trabecula (triangle), D) T. yamato (square) and T. kurosawai (star).