Gayennoides losvilos, RAMÍREZ, 2003

RAMÍREZ, MARTÍN J., 2003, The Spider Subfamily Amaurobioidinae (Araneae, Anyphaenidae): A Phylogenetic Revision At The Generic Level, Bulletin of the American Museum of Natural History 2003 (277), pp. 1-262 : 128-130

Gayennoides losvilos , new species Figures 64A , 66

TYPES: Female holotype from Chile, Región IV, Choapa province, 5 km N Los Vilos, 3 m, under succulent rock cover along cove, ca. 31°55̍S, 71°31̍W, 5.I.1985, N. Platnick, O. Francke , deposited in AMNH.

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Similar to G. molles , but distinguished by having three teeth on the cheliceral retromargin, the male palpal cymbium with a marked retrolateral notch, and a small, procurved epigynal median field.

FEMALE (holotype): Total length 13.30. Carapace length 5.72, width 4.00, wider on legs II–III. Length of tibia/metatarsus: I, 3.72/3.13; II, 3.46/3.00; III, 2.77/2.90; IV, 3.86/4.40. Palpal tarsus length 1.67. Chelicerae unmodified, with three teeth on retromargin. Sternum length 2.93, width 2.07. Spines: leg I, femur d 1–1–1, p 2ap; tibia v p1–2–2; metatarsus v 2bas. II, femur d 1–1– 1, p 0­ d1­ d1, r d1; tibia v r1–2–2, p 0–1; metatarsus v 2bas. III, femur d 1–1–1, p and r 0­ d1­ d1; patella r 1; tibia v p1–2–2, p 1­ d1­ 1­ 0, r d1–1, d r1bas; metatarsus v 2–2–2, p and r d1–1–1, d 0­ p1–2. IV, femur d 1–1– 1, p 0­ d1­ d1, r d1ap; patella r 1; tibia v p1– 2–2, p and r 1­ d1­ 1­ 0, d r1bas; metatarsus = III. Abdomen length 7.70, width 4.40, spiracle–epigastrium 3.70, spiracle–spinnerets 1.45. Color (fig. 64A): carapace brown, reddish to cephalic area, with lateral diffuse dark bands. Chelicerae reddish brown. Legs pale grayish with dark dots at dorsal and lateral spine bases, some spots on patellae and tibiae. Sternum grayish, slightly darker on margins. Abdomen pale gray with dark spots, anterior third of cardiac area gray, posterior third pale, rest of dorsum with diffuse pattern of dark spots. Venter pale with small spots, more concentrated on median stripe. Epigyne (fig. 66F, G): median field elevated, small, short, with ample anterior pouch of hemispheric lumen; two diverging ridges anterior of and at sides of anterior pouch. Copulatory ducts short.

MALE (Carrizal Bajo, not type): Total length 11.15. Carapace length 4.50, width 3.50. Length of tibia/metatarsus: I, 4.52/4.00; II, 3.86/3.30; III, 2.83/3.00; IV, regenerated. Chelicerae longer than those of female, median promarginal tooth quite larger, closer to apical tooth. Sternum length 2.87, width 1.83. Spines as in female, except: leg II, femur p 0­ p1–2; tibia v 0–2–2. III, tibia v p1­ p1–2 or p1–2–2. IV, right absent, left regenerated, with abnormal spines. Abdomen length 6.12, width 3.33, spiracle–epigastrium 2.90, spiracle–spinnerets 1.07. Color as in female. Palp (fig. 66A–E): tibia long, width/ length 0.48, cymbium with conspicuous notch at retrolateral basal margin, slightly advanced compared to that of G. molles . Copulatory bulb: embolus with basal process conspicuous (fig. 66C). Paramedian apophysis with retrolateral cusp squared, prolateral cusp with ventral peak. Primary conductor low, triangular.

NATURAL HISTORY: Collected near seashore, under the Crassulaceae lining stones.

DISTRIBUTION: Known only from seashore of Chile, in Huasco, Choapa, and Elqui provinces.

OTHER MATERIAL EXAMINED: CHILE: Región III (Atacama): Huasco : N of Carrizal Bajo, 22.X.1980, L. Peña, 13 ( AMNH). Región IV (Coquimbo): Elqui: 6 km S Cruz Grande, 20 m, beach, 5 m, 11.XI.1993, 29°29̍S, 71°19̍W, N. Platnick, K. Catley, M. Ramírez, T. Allen , 1♀ ( AMNH, photos MJR 1329–1330) ; 5 km N Los Vilos , 3 m, under succulent rock cover along cove, 5.I.1985, N. Platnick, O. Francke , 3♀ 1 immature ( AMNH) ; 20 km N La Serena (Rt. 5 km 491), 120 m, 7.X.1992, N. Platnick, P. Goloboff, K. Catley , 2♀ ( AMNH) .

ARACHOSIA O.P.­ CAMBRIDGE

Table 21

Arachosia O.P.­ Cambridge, 1882: 425 (type species by monotypy Arachosia anyphaenoides O.P.­ Cambridge, 1882). Keyserling, 1891: 83, 126. Simon, 1897a: 92, 94, 96, 98, 100. F.O.P.­ Cambridge, 1900: 93. Mello­Leitão, 1922: 22. Ramírez, 1995a: 381, 1997: 178.

Samuza Keyserling, 1891: 83 , 134 (type species Samuza praesignis Keyserling, 1891 , designat­ ed by Petrunkevitch, 1928: 174). NEW SYNONY­ MY.

Abuzaida Keyserling, 1891: 83 , 132 (type species Abuzaida striata Keyserling, 1891 , designated by Simon, 1897a: 104). NEW SYNONYMY.

Eusamuza Mello­Leitão, 1915: 144 (type species by original designation Samuza praesignis Keyserling, 1891 ).

TABLE 21 Synapomorphies of Arachosia and Internal Clades

Figures

Fig. 64. Gayennoides spp. A. G. losvilos, sp. n., female (Elqui, Cruz Grande, photo MJR 1330). B. G. molles, sp. n., male (Petorca, Los Molles, photo MJR 1331).

Fig. 66. Gayennoides losvilos, n. sp., male from Carrizal Bajo, Huasco, female holotype. A. Male copulatory bulb, ventral view. B. Same, retrolateral view. C. Detail of embolus, ventral­apical view. D. Male palp, retrolateral view. E. Median and paramedian apophyses, retrolateral view. F. Epigyne, ventral view. G. Same, cleared. Scale bars = A, B, F, G, 0.25 mm; C, E, 0.2 mm; D, 0.5 mm. (MA = median apophysis; PBE = process on base of embolus; PMA = paramedian apophysis; T = tegulum.)