Coptoprepes campanensis, RAMÍREZ, 2003

RAMÍREZ, MARTÍN J., 2003, The Spider Subfamily Amaurobioidinae (Araneae, Anyphaenidae): A Phylogenetic Revision At The Generic Level, Bulletin of the American Museum of Natural History 2003 (277), pp. 1-262 : 87-90

Coptoprepes campanensis , new species Figures 39–41

TYPES: Male holotype from Chile, Región V, Quillota province , Palmas de Ocoa, La Campana Natl. Park, unburned site, 23.VIII.1985, pitfall 1, R. Calderón ; female paratype from the same locality, trap 5, 22.VI.1984, R. Calderón, deposited in AMNH .

ETYMOLOGY: The specific name refers to the type locality, where this species seems to be very common.

DIAGNOSIS: Distinguished from other Coptoprepes by having a huge embolus and primary conductor, and long, contorted female copulatory ducts.

FEMALE (paratype): Total length 3.17. Carapace length 1.40, width 1.02, wider between legs II and III. Length of tibia/metatarsus: I, 0.72/0.61; II, 0.66/0.59; III, 0.57/0.63; IV, 0.98/1.07. Palpal tarsus length 0.41. Chelicerae with three teeth on promargin, four on retromargin. Sternum length 0.81, width 0.61. Spines: leg I, femur d 1–1–1, p d1ap; tibia v 2–2–0; metatarsus v 2bas. II = I. III, femur d 1–1–1, p and r d1ap; patella r 1; tibia v p1­ p1–2, p 1­ d1­ 1­ 0, r d1–1, d r1bas; metatarsus v 2–0–1 (and apical preening comb, fig. 40), p and r d1–1–1, d 0­ p1–2. IV, femur = III; tibia v p1–2–2, p and r 1­ d1­ 1­ 0, d r1bas; metatarsus v 2–2–1, p and r d1–1–1, d 0­ p1–2. Abdomen length 1.87, width 1.07, spiracle–epigastrium 1.00, spiracle–spinnerets 0.16. Color: grayish, spotted in dark brown. Sternum and mouthparts dark brown, leg coxae paler. Abdomen with dark dorsum, darker on cardiac area, with three pairs of paler spots covered by whitish hairs, spots in two posterior pairs closer to each other. Venter grayish, slightly paler than dorsum. Epigyne (fig. 41C, D) partially displaced posteriorly, lateral lobes widely separate, copulatory openings near epigastric fold. Copulatory ducts long, convoluted, fused with those of opposite side though in part of their length. Spermathecae contiguous, lumen small, fertilization ducts separate from posterior border.

MALE (holotype): Total length 3.00. Carapace length 1.43, width 0.97. Length of tibia/metatarsus: I, 1.11/1.01; II, 0.96/0.93; III, 0.73/0.90; IV, 1.11/1.37. Chelicerae slightly narrower than those of female. Sternum length 0.83, width 0.67. Spines as in female, except: leg I, tibia v 2–2–2. II, tibia v r1­ r1– 2. IV, patella r 1. Abdomen length 1.60, width 0.91, spiracle–epigastrium 0.80, spiracle–spinnerets 0.18. Color as in female, but darker carapace, with paler central strip. Palp (figs. 39, 41A, B): femur short; tibia short, as wide as long, RTA absent. Cymbium relatively large, flattened, with deep apical retrolateral notch where median apophysis and secondary conductor fit; cymbial conductor wide. Tegulum basal. Sperm duct with pronounced loop at base of secondary conductor. Embolus with basal process ample, flattened, rounded. Median apophysis apical, large, heavily sclerotized, bifid, apical projection forming right angle (fig. 39B–D). Primary conductor huge, canal deep (fig. 39A– C), tip simple. Secondary conductor triangular (fig. 39B). Additional projection of tegulum near tip of primary conductor. Paramedian apophysis absent.

NATURAL HISTORY: Unknown. Most specimens were collected in pitfall traps.

DISTRIBUTION: Relict forests in central Chile.

OTHER MATERIAL EXAMINED: CHILE: Región IV (Coquimbo): Elqui: 34 km SE La Serena , 29°58̍S, 70°57̍W, 300 m, riparian litter, 23.X.1994, R. Leschen, C. Carlton no. 002, 13 ( AMNH). Limarı´: Many vials from type locality and Fray Jorge Natl. Park, pitfall traps, R. Calderón ( AMNH). Región V (Valparaíso): Petorca: Cuesta El Melón, 520 m, chaparral, 10.I.1985, N. Platnick and O. Francke, 3♀ ( AMNH) ; Los Molles, Rt. 5, km 188, elev. 10 m, 9.XI.1993, 32°14̍S, 71°30̍W, N. Platnick, K. Catley, M. Ramírez, T. Allen , 2♀ ( AMNH). Valparaíso: Quintero, 9.III.19??, 1♀ ( MACN­Ar). Región Metropolitana (Santiago): Santiago: Batuco , 1964, 3♀ 1 immature ( MHNS) ; Quebrada La Plata, fundo La Rinconada de Maipu´, 8.X.1958 – 10.V.1960, W. Noodt, many specimens ( MHNS) ; Valle del Río Mapocho between El Arrayán and Farellones (Barber traps), 15.X.1958 – 8.VI.1960, W. Noodt, many specimens ( MHNS). Cordillera: Río Clarillo Natl. Res. , 940 m, 26.XI.1993, 33°44̍S, 70°28̍W, N. Platnick, K. Catley, M .

Ramírez, T. Allen, 1♀ ( AMNH) ; Río Clarillo , flight intersection trap, 3.XI–7.XII.1989, S.A. Marshall, 1♀ ( AMNH) .

Figures

Fig. 39. Coptoprepes campanensis, n. sp., male copulatory bulb (Quillota, Palmas de Ocoa). A. Ventral view. B. Ventral­apical view. C, D. Apical­prolateral view. (C1 = primary conductor; C2 = secondary conductor; E = embolus; MA = median apophysis; PBE = process on base of embolus.)

Fig. 40. Coptoprepes campanensis, n. sp., female left metatarsus III, ventral view, showing apical unpaired spine and irregular preening comb (Quillota, Palmas de Ocoa).

Fig. 41. Coptoprepes campanensis, n. sp. A. Male palp, ventral view (holotype). B. Same, retrolateral view. C. Epigyne, ventral view (paratype). D. Same, cleared, dorsal view. Scale bars = 0.2 mm.