Pinodytes shoshone Peck & Cook, 2011

Peck, Stewart B. & Cook, Joyce, 2011, Systematics, distributions and bionomics of the Catopocerini (eyeless soil fungivore beetles) of North America (Coleoptera: Leiodidae: Catopocerinae) 3077, Zootaxa 3077 (1), pp. 1-118 : 48-49

Pinodytes shoshone Peck & Cook , new species

( Figs. 42 , 359 , 376–383 )

Type material. Holotype: male ( WFBM). UNITED STATES. Idaho: Shoshone Co.: Hobo Cedar Grove, P.F., VIII-23/ X-18-1985, F.W. Merickel . Paratypes (11): Same locality as holotype, 5.VI.1982, F.W. Merickel, cedar leaf litter Ber., 5 ( WFBM); Latah Co.: 10miNE Harvard, 29.X.1985, F.W. Merickel, B.F., 6 ( WFBM) .

Material examined. We have examined 12 specimens.

Distribution. Specimens ( Fig. 359 ) are known only from Latah and Shoshone counties, northern Idaho.

Diagnostic description. Total length 2.18–2.36 mm.; greatest width 0.98–1.08 mm. Reddish brown; elongateoval in shape ( Fig. 42 ). Head. Finely, sparsely punctate, shining, with substriate microsculpture. Antenna ( Fig. 376 ) with antennomere 2 slightly longer than 3; antennomere 5 slightly larger than 4 and 6; antennomere 7 clearly larger than 8; antennomeres 9 and 10 lack visible sensory vesicles. Pronotum. With fine punctures separated by 2– 4 diameters and a few scattered larger punctures, shining, with reticulate microsculpture. Widest near base; sides nearly parallel in basal one-half, weakly converging in apical one-half; apical margin emarginate, basal margin nearly straight; apical angles rounded, basal angles nearly right-angled. Elytra. Punctures larger than on pronotum, in sublinear rows in basal one-half, smaller and more irregular apically; punctures joined transversely by fine strioles. Slightly wider than pronotum, widest at about basal one-third, sides roundly converging to apex. Legs. Protibia ( Fig. 377 ) widened apically; apex spinose; apical one-half of outer margin with a few spines, apical onehalf of inner margin finely spinose. Mesotibia ( Fig. 378 ) narrow, nearly straight; apex, outer margin and apical onehalf of inner margin strongly spinose. Metatibia ( Fig. 379 ) elongate, narrow, straight; apex spinose, apical one-half with scattered small spines. Metafemur ( Fig. 379 ) elongate, narrow. Male protarsomeres ( Fig. 377 ) not expanded; protarsomere 1 about as long as 2 and 3 combined; with elongate setae laterally and a double row of thin, colorless, transverse phanerae ventrally. Mesotarsomeres without phanerae. Venter. Mesoventrite ( Fig. 383 ) carinate; longitudinal carina shallowly emarginate; large excavation behind transverse carina. Male genitalia. Median lobe of aedeagus ( Figs. 380, 381 ) elongate, broad, flattened before apex; strongly dorsoventrally curved near middle; apex triangular, rounded. Everted internal sac ( Fig. 380 ) with broad, flattened spines of various sizes. Parameres ( Figs. 380, 381 ) slender, apices weakly expanded; not reaching apex of median lobe; each bearing two apical setae. Spermatheca. Elongate ( Fig. 382 ), cylindrical, strongly curved before duct.

Etymology. The name shoshone , a noun in apposition, refers to the occurrence of this species in Shoshone Co., Idaho.

Figures

FIGURE 359. Distributions of species in the Pinodytes cryptophagoides species group in northwestern United States. P. isabella, open triangle in northern Idaho. P.orca, closed square in southwestern Oregon. P.setosus, closed triangles in southwestern Oregon and northwestern California. P. shoshone, closed circles in northern Idaho. P. sinuatus, open squares in southwestern Oregon.

FIGURES 37–43. Pinodytes habitus photos. 37. P.garibaldi, total length 2.80 mm. 38. P. haidagwaii, total length 3.04 mm. 39. P. isabella, total length 2.18 mm. 40. P.orca, total length 4.20 mm. 41. P.setosus, total length 1.96 mm. 42. P. shoshone, total length 2.18 mm. 43. P. sinuatus, total length 2.28 mm.

FIGURES 376–383. Pinodytes shoshone. 376. Male right antenna, dorsal. 377. Male right protibia and protarsus. 378. Male right mesotibia. 379. Male right metafemur and metatibia. 380. Aedeagus, lateral. 381. Aedeagus, dorsal. 382. Spermatheca. 383. Mesoventrite, lateral. Scale bar = 0.15 mm (Fig. 382), 0.3 mm (Figs. 381, 383), 0.4 mm (Figs. 377–380), 0.45 mm (Fig. 376).