Phthiracarus (Trachyhoplophora) Berlese, 1923

Niedbała, Wojciech & Liu, Dong, 2023, Systematic, synonymic and biogeographical list of ptyctimous mites (Acari, Oribatida) in the world (1799 – 2022), Zootaxa 5265 (1), pp. 1-442 : 214-223

Probably Oriental species introduced to western Pacific Islands solomonensis Notophthiracarus Niedbała, 1998

Notophthiracarus solomonensis: Niedbała 2006 d, Niedbała & Liu 2018 .

Notophthiracarus (Notophthiracarus) solomonensis: Subías 2004 .

Hoplophthiracarus (Notophthiracarus) solomonensis: Subías 2022 .

Australasia: Solomon Islands ( Balogh & Balogh 2002, Niedbała 1998b, 2006d, 2012a, Subías 2022). Australasian, Solomon Islands, endemic somalicus Phthiracarus (Trachyhoplophora) Berlese, 1923 incertae sedis

Phthiracarus echidninus Berl. var. somalicus Berlese, 1923 (in part): Niedbała 1994b.

Phthiracarus (Trachyhoplophora) somalicus: Castagnoli & Pegazzano 1985 , Niedbała 1986b, 1992. Steganacarus somalicus: Hammen 1959 .

Hoplophthiracarus somalicus: Mahunka 1991 a, Mahunka & Mahunka-Papp 1995 .

Notophthiracarus (?) somalicus: Niedbała 1994b, 2001 a , Niedbała & Liu 2018.

Notophthiracarus (Notophthiracarus) somalicus: Subías 2004 .

Hoplophorella (Rhacaplacarus) somalica: Subías 2022 .

Afrotropic: Somalia ( Berlese 1923, Castagnoli & Pegazzano 1985, Hammen 1959, Mahunka 1991 a, Mahunka & Mahunka-Papp 1995, Niedbała 1986b, 1992, 1994b, 2001a, Subías 2022).

Afrotropical, Somalia, endemic

Notes. Species represented in the collection of Berlese is poorly preserved and most important characters could not be examined.

Subías (2022) considered this species as congeneric with Plonaphacarus persimilis Niedbała, 1994 . See note of P. persimilis .

soniae Hauseroplophora Mahunka, 1977

Hauseroplophora soniae: Niedbała 2004b , Niedbała & Liu 2018, Subías 2022.

Afrotropic: Kenya ( Balogh & Balogh 2002, Mahunka 1977b, Niedbała 2004b, 2008c, Subías 2022). Afrotropical, Kenya, endemic sordidus Notophthiracarus Niedbała et Colloff, 1997

Notophthiracarus sordidus: Niedbała 2000 , Liu & Zhang 2014 d, Niedbała & Liu 2018.

Notophthiracarus (Notophthiracarus) sordidus: Subías 2004 .

Hoplophthiracarus (Notophthiracarus) sordidus: Subías 2022 .

Australasia: Australia (Tasmania) ( Balogh & Balogh 2002, Niedbała 2000, 2012a, Niedbała & Colloff 1997, Subías 2022).

Australasian, Australia, endemic sororius Arphthicarus (Niedbała, 1982)

Hoplophthiracarus sororius Niedbała, 1982

Austrophthiracarus sororius: Niedbała 1986b, 1992 .

Arphthiracarus sororius: Niedbała 1994a, 2003a , 2004a, 2008e, Niedbała & Liu 2018.

Notophthiracarus (Notophthiracarus) sororius: Subías 2004 .

Hoplophorella (Hoplophorella) sororia: Subías 2022 .

Neotropic: Mexico, Guatemala, Costa Rica, Panama ( Balogh & Balogh 1987, 1988, 2002, Niedbała 1982d, 1986b, 1992, 2003a, 2004a, 2008e, Subías 2022).

Neotropical, Mesoamerica spadix Phthiracarus Niedbała, 1983

Phthiracarus spadix: Niedbała 1986b, 1992 , 2008f, 2011b, 2012b, Olszanowski et al. 1996, Niedbała & Olszanowski 1997, 2004, 2008, Niedbała & Liu 2018.

Phthiracarus (Phthiracarus) spadix: Subías 2022 .

Palaearctic: Switzerland, Austria, Germany, Poland, Czech Republic, Slovakia, Romania, Ukraine (western part) ( Beck et al. 2014, Mahunka & Mahunka-Papp 2003, Niedbała 1983c, 1986b, 1992, 2008f, 2011b, 2012b, 2015b, Olszanowski et al. 1996, Subías 2022, Weigmann et al. 2015).

Western Palaearctic, distributed in Central Europe and Balkan region. Carpathian species sparsa Mesoplophora (Mesoplophora) Niedbała, 2004

Mesoplophora (Mesoplophora) sparsa: Liu & Chen 2010c , Liu 2016 a, Niedbała & Liu 2018, Subías 2022.

Neotropic: Peru ( Niedbała 2004 a, Oliveira et al. 2017, Subías 2022).

Neotropical, Peru, endemic spathulatus Notophthiracarus Niedbała, 2019

Hoplophthiracarus (Steganacarellus) spathulatus: Subías 2022 .

South Africa: Western Cape (Niedbała et al. 2019).

South African, endemic spatulata Hoplophorella Parry, 1980 synonym of Atropacarus (Hoplophorella) hamatus (Ewing, 1909)

sphaerula Hoplophora Banks, 1895 synonym of Phthiracarus globosus (C.L. Koch, 1841)

spiculifera Acrotritia (Mahunka, 1991)

Rhysotritia clavata spiculifera Mahunka, 1991

Rhysotritia spiculifera: Niedbała 1998a , b, 2000, 2001a, 2004d, Niedbała & Corpuz-Raros 1998, Mahunka 1999b, Niedbała & Penttinen 2007.

Acrotritia spiculifera: Niedbała 2006d, 2012 a , Liu & Zhang 2014a, Liu 2015 d, Niedbała & Liu 2018. Afrotropic: Ivory Coast, Ghana, Cameroon, Tanzania, Seychelles, Comoro Islands, Madagascar. Orient: Sri Lanka, Indonesia (West Sumatra), Malaysia, Philippines. Australasia: Papua New Guinea, Australia (northern and southern parts), New Caledonia, Fiji, Tahiti ( Balogh & Balogh 2002, Mahunka 1991c, 1999, 2002, Niedbała 1998a, b, 2000, 2001a, 2004d, 2006d, 2008b, c, d, 2012a, 2014b, 2017a, b, c, Niedbała & Corpuz-Raros 1998, Niedbała & Osiejuk 2018, Niedbała & Penttinen 2007, Subías 2022).

Pantropical

Note. It is unlikely that this species is congeneric with A. sinensis Jacot, 1923 as suggested by Subías (2022). Both species differ in the shape of trichobothria.

spinatus Steganacarus (Rhacaplacarus) Niedbała, 2001

Steganacarus (Rhacaplacarus) spinatus: Niedbała & Liu 2018 .

Rhacaplacarus (Rhacaplacarus) spinatus: Subías 2004 .

Hoplophorella (Rhacaplacarus) spinata: Subías 2022 .

Afrotropic: Democratic Republic Congo ( Niedbała 2001a, Subías 2022).

Afrotropical, Democratic Republic Congo, endemic spindleformis Indotritia (Indotritia) Pan, Xu et Liu, 2018

Indotritia spindleformis Pan, Xu et Liu, 2018

Indotritia spindleformis: Liu 2021b .

Indotritia (Indotritia) spindleformis: Subías 2022 .

Orient: China (Guangxi) (Liu 2021b, Pan et al. 2018).

Oriental, China, endemic spiniferus Steganacarus (Rhacaplacarus) (Mahunka, 1993)

Rhacaplacarus spiniferus Mahunka, 1993

Steganacarus (Rhacaplacarus) spiniferus: Niedbała 2001 a, Niedbała & Liu 2018 .

Rhacaplacarus (Rhacaplacarus) spiniferus: Subías 2004 .

Hoplophorella (Rhacaplacarus) spinifera: Subías 2022 .

Afrotropic: Tanzania ( Balogh & Balogh 2002, Mahunka 1993d, Niedbała 2001a, 2017c, Subías 2022). Afrotropical, Tanzania, endemic spiniformis Arphthicarus Liu, Wu et Chen, 2014

Arphthicarus spiniformis: Niedbała & Liu 2018 , Liu 2021b.

Hoplophthiracarus (Notophthiracarus) spiniformis: Subías 2022 .

Orient: China (Guangxi) (Liu 2021b, Liu et al. 2014, Subías 2022).

Oriental, China, indigenous spiniformis Notophthiracarus Niedbała, 2004

Notophthiracarus spiniformis: Niedbała & Liu 2018 .

Hoplophthiracarus (Steganacarellus) spiniformis: Subías 2022 .

Neotropic: Venezuela ( Niedbała 2004a, Subías 2006).

Neotropical, Venezuela, endemic spiniformis Plonaphacarus Liu et Wu, 2019

Plonaphacarus spiniformis: Liu 2021b .

Hoplophthiracarus (Hoplophthiracarus) spiniformis: Subías 2022 .

Orient: China (Chongqing) (Liu 2021 b, Liu & Wu 2019).

Oriental, China, endemic spiniger Steganacarus (Rhacaplacarus) (Aoki, 1980)

Hoplophorella spiniger Aoki, 1980

Steganacarus (Rhacaplacarus) spiniger: Niedbała 1986b, 1992 , 2000, 2011b, 2012b, Niedbała & Liu 2018.

Rhacaplacarus (Rhacaplacarus) spiniger: Subías 2004 .

Hoplophorella (Kakophthiracarus) spiniger: Subías 2022 .

Palaearctic: Japan ( Aoki 1980a, Niedbała 1986b, 1992, 2000, 2011b, 2012b, Subías 2022).

Eastern Palaearctic, Japan, endemic spinosa Apoplophora Mahunka, 1987

Apoplophora spinosa: Mahunka 1988d, 1991b , Niedbała 2000, 2001c, 2004c, Liu et al. 2009a, Liu 2016 a, Niedbała & Liu 2018, Subías 2022.

Orient: Malaysia (Sabah, Sarawak), Indonesia (Sumatra) ( Balogh & Balogh 2002, Mahunka 1987f, 1988d, 1991b, Niedbała 2000, 2001c, 2004c, 2008b, Subías 2022).

Oriental, Borneo and Sumatra spinosa Mesotritia Aoki, 1980

Mesotritia spinosa: Aoki 1980a , Niedbała 2011b, 2012b, Niedbała & Liu 2018.

Mesotritia markeli (lapsus linguae): Niedbała 2000.

Palaearctic: Japan ( Aoki 1980a, Niedbała 2011b, 2012b, Subías 2022).

Eastern Palaearctic, Japan, endemic

Note. This species was mistakenly synonymized with M. maerkeli ( Liu & Chen 2010 a, Niedbała & Liu 2018).

spinosa Oribotritia (Oribotritia) (Mahunka, 1988)

Philotritia spinosa Mahunka, 1988

Oribotritia spinosa: Niedbała 1998a, 2004d , Mahunka 1999b, Subías 2004, Liu 2015 a, Niedbała & Liu 2018.

Oribotritia (Oribotritia) spinosa: Subías 2022 .

Afrotropic: Madagascar, Mauritius, Reunion ( Balogh & Balogh 2002, Mahunka 1988b, 1999b, 2002, Niedbała 1998a, 2001, 2004d, 2008c, 2017a, Subías 2022).

Afrotropical, eastern islands spinosus Steganacarus (Steganacarus) ( Sellnick, 1920)

Phthiracarus spinosus Sellnick, 1920 : Castagnoli & Pegazzano 1985.

Steganacarus spinosus: Hammen 1952 , Krivolutsky 1975, Balogh & Mahunka 1980 a, 1983, Olszanowski et al. 1996, Niedbała & Olszanowski 1997, 2004, 2008.

Steganacarus (Steganacarus) spinosus: Niedbała 1986b, 1992 , 1994a, 2008b, 2011b, 2012b, Avanzati & Bernini 1989, Olszanowski & Niedbała 2000, Mahunka & Mahunka-Papp 2004, Subías 2004, Weigmann 2006, Mortazavi et al. 2010, Niedbała & Liu 2018, Niedbała et al. 2021, Subías 2022.

Steganacarus barborae: Niedbała 1992 .

Steganacarus barborae Mahunka et Miko, 1989 syn. by Niedbała 1994a.

Steganacarus (Steganacarus) punctulatus Sergienko, 1985 syn. by Niedbała 1986b, 1992.

Steganacarus (Steganacarus) punctulatus Sergienko : Subías 2022

Palaearctic: Norway, Finland, Belgium, the Netherlands, Germany, Austria, Poland, Czech Republik, Slovakia, Hungary, Serbia, Macedonia, Romania, Bulgaria, Ukraine (western part), Azerbaijan, Russia (South European Territory), Turkey, Iran, India (Meghalaya) ( Avanzati & Bernini 1989, Balogh & Mahunka 1980 a, 1983, Castagnoli & Pegazzano 1985, Hammen 1952, Mahunka et al. 2013, Mahunka & MahunkaPapp 2004, Mahunka & Miko 1989, Mortazavi et al. 2010, Niedbała 1986b, 1994a, 1992, 2011b, 2012b, 2015a, b, Niedbała et al. 2020, 2021, Olszanowski et al. 1996, Schatz 1983, Sellnick 1920, Seniczak et al. 2021, Sergienko 1985, Subías 2022, Weigmann et al. 2015).

South-Western Palaearctic, Panearopean, except Mediterranean subregion and also reported from Anatolian, Pontic and Iraniano-Kashmiran areas

Note. Subías (2022) treated Steganacarus (S.) personatus Niedbała, 1983 as subspecies of this species. However, we find no reason to establish the status of the subspecies.

spinulosa Synichotritia Walker, 1965

Synichotritia spinulosa: Lions & Norton 1999 , Niedbała 2002a, 2011b, 2012b, Niedbała & Liu 2018, Subías 2022.

Nearctic: Canada (British Columbia), USA (Oregon, Mississippi, California) .

Palaearctic: Russia (Far East) ( Lions & Norton 1999, Mahunka 1990, Marshall et al. 1987, Niedbała 2002a, 2011b, 2012b, 2015b, Niedbała & Starý 2015e, Subías 2022, Walker 1965).

Nearctic, western part with south-eastern disjunction (Mississippi), introduced to eastern Palaearctic spinus Euphthiracarus (Euphthiracarus) Niedbała, 2002

Euphthiracarus spinus Niedbała, 2002

Euphthiracarus spinus: Niedbała & Liu 2018 .

Euphthiracarus (Euphthiracarus) spinus: Subías 2022 .

Nearctic: USA (Washington, Nevada) ( Niedbała 2002a, Subías 2022).

Nearctic, western part spinus Hoplophthiracarus Niedbała et Starý, 2014

Hoplophthiracarus spinus: Niedbała & Liu 2018 .

Hoplophorella (Rhacaplacarus) spina: Subías 2022 .

Afrotropic: Cameroon ( Niedbała & Osiejuk 2018, Niedbała & Starý 2014 d, Subías 2022).

Afrotropical, Cameroon, endemic spinus Notophthiracarus Niedbała, 2006

Notophthiracarus spinus: Niedbała & Liu 2018 .

Hoplophthiracarus (Notophthiracarus) spinus: Subías 2022 .

South Africa: Western Cape ( Niedbała 2006c, Subías 2022).

South African, endemic spinus Steganacarus (Rhacaplacarus) Niedbała, 2014

Steganacarus (Rhacaplacarus) spinus: Niedbała & Ermilov 2014b , Niedbała & Liu 2018.

Hoplophorella (Kakophthiracarus) spinus: Subías 2022 .

Orient: Southern Vietnam ( Niedbała & Ermilov 2014b, Subías 2022).

Oriental, Southern Vietnam, endemic spurcus Notophthiracarus Niedbała et Colloff, 1997

Notophthiracarus spurcus: Niedbała 2000, 2006 d , Liu & Zhang 2014 d, Niedbała & Liu 2018. Notophthiracarus (Notophthiracarus) spurcus: Subías 2004 .

Hoplophthiracarus (Notophthiracarus) spurcus: Subías 2022 .

Australasia: Australia (Tasmania) ( Balogh & Balogh 2002, Niedbała 2000, 2006d, 2012a, Niedbała & Colloff 1997, Subías 2022).

Australasian, Australia, endemic stenodes Steganacarus (Rhacaplacarus) Niedbała, 2004

Steganacarus (Rhacaplacarus) stenodes: Niedbała & Liu 2018 .

Hoplophorella (Mantigueracarus) stenodes: Subías 2022 .

Neotropic: Peru, Brazil ( Niedbała 2004 a, Oliveira 2004, Oliveira et al. 2017, Subías 2022).

Neotropical, Guyano-Brasilian and Patagono-Andean stenos Atropacarus (Hoplophorella) Niedbała et Starý, 2014

Atropacarus (Hoplophorella) stenos: Niedbała & Liu 2018 .

Hoplophorella (Kakophthiracarus) stenos: Subías 2022 .

Afrotropic: Tanzania (Niedbała 2017c, Niedbała & Starý 2014a, Subías 2022).

Afrotropical, Tanzania, endemic stenotus Notophthiracarus Niedbała et Corpuz-Raros, 1998

Notophthiracarus stenotus: Niedbała 2000 , Niedbała et al. 2006, Niedbała & Liu 2018.

Notophthiracarus (Notophthiracarus) stenotus: Subías 2004 .

Hoplophthiracarus (Steganacarellus) stenotus: Subías 2022 .

Orient: Philippines, Indonesia (Luzon, Samar). Australasia: Papua New Guinea ( Niedbała 2000, Niedbała

& Corpuz-Raros 1998, Niedbała & Starý 2015e, Niedbała et al. 2006, Subías 2022).

Oriental, Philippinian subregion stenotus Phthiracarus Niedbała, 2002

Phthiracarus stenotus: Niedbała 2004 a, Niedbała & Liu 2018 .

Phthiracarus (Phthiracarus) stenotus: Subías 2022 .

Nearctic: Bermuda. Neotropic: Antilles ( Niedbała 2002b, 2004a, Subías 2022).

Nearctic and Neotropical, Mesoamerica sterigma Acrotritia (Niedbała, 1998)

Rhysotritia sterigma Niedbała, 1998

Acrotritia sterigma: Subías 2004 , Niedbała 2007a, 2008b, Liu & Zhang 2014a, Liu 2015 d, Niedbała & Liu 2018, Niedbała & Ermilov 2020.

Orient: Malaysia ( Sarawak, Sabah), Indonesia (Bali). Australasia: Society Islands, Australia (Queensland, New South Wales) ( Balogh & Balogh 2002, Niedbała 1998b, 2007a, 2008b, 2012a, 2014b, 2017b, Niedbała & Ermilov 2020, Subías 2022).

Pantropical, eastern part

Note. Subías (2022) considered this species as congeneric with A. brasiliana (Mahunka, 1983) . However, these species differ from others in the shape of trichobothria (rodlike in A. brasiliana versus with distinct head in A. sterigma ) as well as in their geographical distance.

stigmosus Hoplophthiracarus Niedbała, 2000

Hoplophthiracarus stigmosus: Subías 2004 , Niedbała & Liu 2018.

Hoplophthiracarus (Hoplophthiracarus) stigmosus: Subías 2022 .

Orient: Northern Vietnam, Indonesia ( Flores) ( Niedbała 2000, Subías 2022).

Oriental, Indo-Chinese-Malaysian and Sundaian subregions, disjunctive)

stilifer Atropacarus (Hoplophorella) ( Hammer, 1961)

Steganacarus stilifer Hammer, 1961

Hoplophorella stilifer: Niedbała 1984b .

Atropacarus (Hoplophorella) stilifer: Niedbała 1986b, 1992 , 1998b, 2000, 2001a, 2003a, 2004a, 2008e, Niedbała & Schatz 1996, Niedbała & Penttinen 2007, Niedbała & Liu 2018.

Hoplophorella (Hoplophorella) stilifera: Subías 2022 .

Hoplophorella benoiti Mahunka, 1984 syn. by Niedbała 2001a.

Hoplophorella benoiti: Mahunka 1992 , Niedbała 1992.

Neotropic: Guatemala, Belize, Costa Rica, Panama, Antilles ( Cuba), Galapagos Islands, Peru, Brazil. Afrotropic: Cameroon, Angola, Zambia, Seychelles. Orient: India (southern), Southern and Northern Vietnam, Philippines. Australasia: Tuamotu Islands, Marquesas Islands, Henderson Island ( Balogh & Balogh 1987, 1988, 2002, Ermilov & Niedbała 2013, Hammer 1961, Mahunka 1984b, 1992, Niedbała 1984b, 1986b, 1992, 1998b, 2000, 2001a, 2003a, 2004a, 2008c, e, 2012a, 2017 a, Niedbała & Ermilov 2014b, 2015, 2017a, Niedbała & Penttinen 2007, Niedbała & Schatz 1996, Oliveira 2004, Oliveira et al. 2017, Schatz 1998, Subías 2022).

Pantropical storkani Oribotritia (Oribotritia) Feider et Suciu, 1957 species inquirenda

Oribotritia storkani Feider et Suciu, 1957

Oribotritia storkani: Krivolutzky 1975 , Balogh & Mahunka 1980 a, 1983, Subías 2004, Niedbała 2011b, Liu 2015 a, Niedbała & Liu 2018.

Oribotritia (Oribotritia) storkani: Subías 2022 .

Palaearctic: Romania ( Balogh & Mahunka 1980 a, 1983, Feider & Suciu 1957, Mahunka et al. 2013, Subías 2022).

Note. Oribotritia storkani sensu Grandjean, 1967 is probably synonymous with Oribotritia serrata Feider et Suciu, 1958 (presence of three aggenital setae, two anal and three adanal setae). Many morphological features have not been mentioned in the original description, e.g. the presence of rostral and c 3 setae, number of genital setae, number of anal setae.

straminea Hoplophora C.L. Koch, 1841 species inquirenda

Hoplophora straminea: Oudemans 1937 , Niedbała 1986b, 1992, 2011b, Niedbała & Liu 2018.

Phthiracarus stramineus: Jacot 1936 , Sellnick 1960, Berg et al. 1990, Niedbała 2002 a, Mahunka & Mahunka-Papp 2004.

Archiphthiracarus stramineus: Balogh & Mahunka 1980a .

Phthiracarus (Archiphthiracarus) stramineus: Subías 2004 .

Nearctic: USA (Connecticut, New York). Palaearctic: Germany ( Balogh & Mahunka 1980 a, 1983, Jacot 1936, C.L. Koch 1841, Niedbała 1986b, 1992, 2002a, 2015b, Subías 2022).

Note. Type specimens do not exist. On the basis of the redescription by Jacot (1936), this species cannot be unambiguously classified. Subías (2022) recognized H. stramineae as probably congeneric with P. bryobius Jacot, 1930 . However, with regard to Jacot (1936) description of P. stramineus based on topotypic material, P. bryobius differs from P. stramineus by the longer setae of prodorsum and notogaster and the trichobothria rounded distally (versus pointed distally in P. stramineus ).

stria Microtritia Liu et Zhang, 2014

Microtritia stria: Liu & Chen 2014 a, Niedbała & Liu 2018 .

Microtritia striata (lapsus linguae): Subías 2022.

Australasia: New Zealand ( Liu & Zhang 2014a, Subías 2022).

Australasian, New Zealand, endemic striata Mesoplophora Mahunka, 1985 synonym of Apoplophora pantotrema ( Berlese, 1913)

striata Oribotritia (Oribotritia) Mahunka, 2009

Oribotritia striata Mahunka, 2009

Oribotritia striata: Liu 2015 a, Niedbała & Liu 2018 .

Oribotritia (Oribotritia) striata: Subías 2022 .

Afrotropic: Madagascar (Mahunka 2009a, Niedbała 2017a, Subías 2010).

Afrotropical, Madagascar, endemic striata Sabahtritia Liu et Chen, 2009

Sabahtritia striata: Niedbała & Liu 2018 , Liu 2021b, Subías 2022.

Orient: China (Hainan Island) (Liu 2021b, Liu & Chen 2009, Subías 2022).

Oriental, China, endemic striatissima Microtritia Mahunka, 1999

Microtritia striatissima: Niedbała 2001a, 2004 d , Liu & Chen 2014 a, Niedbała & Liu 2018.

Afrotropic: Madagascar ( Balogh & Balogh 2002, Mahunka 1999b, 2002, Niedbała 2001a, 2004d, 2017a, Subías 2022).

Afrotropical, Madagascar, endemic strictus Arphthicarus Niedbała, 2017

Arphthicarus strictus: Niedbała & Ermilov 2017b , Niedbała & Liu 2018.

Hoplophorella (Hoplophorella) stricta: Subías 2022 .

Neotropic: Antilles ( Trinidad) ( Niedbała & Ermilov 2017b, Subías 2022).

Neotropical, Trinidad, endemic striculus Atropacarus (Atropacarus) (C.L. Koch, 1836)

Hoplophora stricula C.L. Koch, 1836

Hoplophora stricula: Ewing 1917b , Oudemans 1937, Grandjean 1953.

Phthiracarus stricula: Castagnoli & Pegazzano 1985 .

Steganacarus striculus: Grandjean 1933a , d, Hammen 1952, Feider & Suciu 1957.

Atropacarus striculum: Grandjean 1934a , c.

Atropacarus striculus: Aoki 1958b, 1980 a , Kamill & Baker 1980, Mahunka 1990b, Olszanowski et al. 1996, Niedbała & Olszanowski 1997, 2004, 2008, Wang et al. 2002, Mahunka & Mahunka-Papp 2004, Chen et al. 2010, Liu 2021b.

Atropacarus striculus striculus: Subías 2022 .

Atropacarus (Atropacarus) striculus: Doumandji et al. 1985 , Niedbała 1985b, 1986b, 1992, 1994b, c, 2000, 2002a, 2004a, 2006d, 2007c, 2008e, f, 2011b, 2012b, Niedbała & Ermilov 2018a, 2022, Olszanowski & Niedbała 2000, Niedbała & Szywilewska-Szczykutowicz 2017, Niedbała & Liu 2018, Niedbała et al. 2021.

Steganacarus striculus: Hammen 1952 , Feider & Suciu 1957, Aoki 1958b, Krivolutsky 1975, Balogh &

Mahunka 1980 a, 1983, Niedbała et al. 1981, 1982, 1990, Subías 2022.

Steganacarus (Atropacarus) striculus: Sergienko 1985 , Weigmann 2006.

Phthiracarus echidninus (nomen nudum): Hammen 1959, Castagnoli & Pegazzano 1985, Niedbała 1994b, Mahunka & Mahunka-Papp 1995.

Steganacarus diaphanum Jacot, 1930 syn. by Kamill & Baker 1980.

Steganacarus diaphanum: Niedbała 1986b .

Steganacarus senex Aoki, 1958 syn. by Fujikawa 1972.

Steganacarus senex: Aoki 1958b, 1980 a , Balogh & Mahunka 1983.

Steganacarus striculus insularis Weigmann, 1976 : Kamill and Baker (1980) (wrongly synonymized with Atropacarus striculus (C.L. Koch) .

Steganacarus trichosus Mahunka, 1982 syn. by Niedbała 1986b.

Nearctic: Canada (Yukon, British Columbia, Alberta, Manitoba, Ontario, Quebec, New Brunswick, Nova Scotia ), USA (Washington, Oregon, California, Wyoming, Utah, Colorado, Arizona, New Mexico, South Dakota, Minnesota, Wisconsin, Iowa, Illinois, Michigan, Indiana, Ohio, Maine, Vermont, New Hampshire, Massachusetts, Pennsylvania, New York, New Jersey, West Virginia, Virginia, Tennessee, North Carolina, South Carolina, Georgia). Palaearctic:Azores, England, Spain, Portugal, France, Denmark, the Netherlands, Belgium, Norway, Sweden, Finland, Germany, Luxembourg, Switzerland, Austria, Italy, Poland, Czech Republik, Slovakia, Hungary, Slovenia, Croatia, Greece, Serbia, Romania, Bulgaria, Estonia, Latvia, Lithuania, Belarus (Brest, Minsk, Palm houses), Ukraine (western and eastern parts, Crimea), Georgia, Russia (North, Central and South European Territory, Western and Eastern Siberia, Far East), Algeria, Iran, Kazakhstan, Tajikistan, India (Himachal Pradesh), Nepal, Mongolia, China (Beijing, Heilongjiang, Jilin), North and South Korea, Japan. Neotropic: Mexico, Guatemala. Afrotropic: Cameroon, Tanzania ( Zanzibar), Madagascar. Orient: Philippines, Northern Vietnam, China ( Taiwan, Zhejiang). Australasia: Australia (New South Wales, Victoria, Tasmania), Hawaii Islands ( Aoki 1958b, 1970, 1980, Balogh & Mahunka 1980 a, 1983, Bayartogtokh & Yondon 2019, Castagnoli & Pegazzano 1985, Chen et al. 2010, Doumandji et al. 1985, Ewing 1917b, Feider & Suciu 1957, Grandjean 1933a, d, 1934c, 1953, Hammen 1952, Hammer 1952, Jacot 1930, 1938, Kaczmarek & Niedbała 2002, Kamill & Baker 1980, C.L. Koch 1836, Liu 2021b, Mahunka 1982c, 1990 b, Mahunka & Mahunka-Papp 1999, 2003, 2004, Mahunka et al. 2013, Marshall et al. 1987, Niedbała 1985b, 1986b, 1992, 1994b, c, 1998c, 2000, 2002a, 2004a, 2006d, 2007c, 2008e, f, 2012a, 2015b, 2017a, b, 2018, Niedbała & Buczkowska 2020, Niedbała & Ermilov 2014a, 2017a, 2018a, 2019b, 2020, 2021, 2022, Niedbała et al. 2020, 2021, Niedbała & Osiejuk 2018, Niedbała & Starý 2015e, Niedbała & Szywilewska-Szczykutowicz 2017, Norton & MacNamara 1976, Olszanowski et al. 1996, Oudemans 1937, Ryabinin et al. 2018, Schatz 1983, Seniczak et al. 2021, Sergienko 1985, Subías 2022, Wang et al. 2002, Weigmann et al. 2015).

Semicosmopolitan

Note. Subías (2022) treated Atropacarus (A.) insularis ( Weigmann, 1976) as subspecies of this species. See note of A. (A.) insularis .

striculus insularis Steganacarus Weigmann, 1976 = Atropacarus (Atropacarus) insularis ( Weigmann, 1976)

striculus var. clavatus Atropacarus Aoki, 1980 = Atropacarus (Atropacarus) clavatus Aoki, 1980

strigosus Austrophthiracarus (Niedbała, 1984)

Neophthiracarus strigosus Niedbała, 1984

Calyptophthiracarus strigosus: Niedbała 1986b, 1992 .

Austrophthiracarus strigosus: Niedbała 1994a, 2004a , Liu & Zhang 2016 a, Niedbała & Liu 2018.

Notophthiracarus (Protophthiracarus) strigosus: Subías 2004 .

Hoplophthiracarus (Protophthiracarus) strigosus: Subías 2022 .

Neotropic: Argentina ( Balogh & Balogh 1987, 1988, 2002, Niedbała 1984m, 1986b, 1992, 2004a, Subías 2022).

Neotropical, Argentina, endemic striolatus Arphthicarus Niedbała, 2004

Arphthicarus striolatus: Niedbała & Liu 2018 .

Hoplophthiracarus (Notophthiracarus) striolatus: Subías 2022 .

Neotropic: Colombia, Ecuador ( Niedbała 2004 a, Niedbała & Roszkowska 2017, Subías 2022).

Neotropical, Guyano-Brasilian styphelos Plonaphacarus Niedbała, 2001

Plonaphacarus styphelos: Liu et al. 2011b , Niedbała & Liu 2018.

Rhacaplacarus (Rhacaplacarus) styphelos: Subías 2004 .

Hoplophorella (Rhacaplacarus) styphelos: Subías 2022 .

Afrotropic: Ethiopia, Tanzania ( Niedbała 2001a, 2017c, Niedbała & Ermilov 2022c, Subías 2022). Afrotropical, eastern part subcedrus Euphthiracarus (Euphthiracarus) Niedbała, 2006

Euphthiracarus subcedrus Niedbała, 2006

Euphthiracarus subcedrus: Niedbała 2011b, 2012b , Niedbała & Liu 2018.

Euphthiracarus (Euphthiracarus) subcedrus: Subías 2022 .

Palaearctic: Russia (Western Siberia, Far East) ( Niedbała 2006a, 2011b, 2012b, Subías 2022). Central-Eastern Palaearctic, in Western Siberia and Amurian areas, disjunctive subciliatus Atropacarus (Hoplophorella) (Mahunka, 1983)

Hoplophorella subciliata Mahunka, 1983

Hoplophorella subciliata: Niedbała 1986b, 1992 .

Atropacarus (Hoplophorella) subciliatus: Niedbała 2001 a, Niedbała & Liu 2018 .

Hoplophorella (Hoplophorella) subciliata: Subías 2022 .

Afrotropic: Tanzania ( Balogh & Balogh 2002, Mahunka 1983b, Niedbała 1986b, 1992, 2001a, 2017c, Subías 2022).

Afrotropical, Tanzania, endemic subdolus Phthiracarus Niedbała, 1983

Phthiracarus subdolus: Niedbała 1986b, 1992 , 2011b, 2012b, Niedbała & Liu 2018.

Phthiracarus (Archiphthiracarus) subdolus: Subías 2004 .

Phthiracarus (Phthiracarus) subdolus: Subías 2022 .

Palaearctic: Greece, Ukraine (Crimea), Azerbaijan, Georgia, Russia (South European Territory) ( Mahunka et al. 2013, Niedbała 1983a, 1986b, 1992, 2011b, 2012b, 2015a, Subías 2022).

Western Palaearctic, distributed in Ponto-Mediterranean and Black Sea-Pontic areas subglobosus Phthiracarus Berlese, 1924 synonym of Phthiracarus globosus (C.L. Koch, 1841)

subiasi Phthiracarus Niedbała, 1986

Phthiracarus subiasi: Niedbała 1992, 2011b , 2012b, Niedbała & Liu 2018.

Phthiracarus (Archiphthiracarus) subiasi: Subías 2022 .

Palaearctic: Canary Islands, Spain ( Niedbała 1986 d, 1992, 2011b, 2012b, Subías 2022).

Western Palaearctic, known from Macaronesian and Iberian areas subitus Notophthiracarus (Niedbała, 1983)

Hoplophorella subita Niedbała, 1983

Atropacarus (Hoplophorella) subitus: Niedbała 1986b, 1992 , 2001 a, Niedbała & Liu 2018. Kakophthiracarus subitus: Mahunka & Mahunka-Papp 2007 .

Hoplophorella (Hoplophorella) subita: Subías 2022 .

Notophthiracarus subitus: Niedbała & Ermilov 2022c .

Afrotropic: Ethiopia, Kenya ( Balogh & Balogh 2002, Niedbała 1983i, 1986b, 1992, 2001a, Niedbała & Ermilov 2022c, Subías 2022).

Afrotropical, eastern part submolesta Oribotritia (Oribotritia) Niedbała, 2000

Oribotritia submolesta Niedbała, 2000

Oribotritia submolesta: Subías 2004 , Niedbała 2011b, 2012b, Liu 2015 a, Niedbała & Liu 2018. Oribotritia (Oribotritia) submolesta: Subías 2022 .

Orient: Southern and Northern Vietnam. Palaearctic: North India ( Niedbała 2000, 2011b, 2012b, Niedbała & Ermilov 2014b, Subías 2022).

Oriental, introduced to northern India subsellatus Protophthiracarus ( Balogh et Mahunka, 1977)

Hoplophorella subsellata Balogh et Mahunka, 1977

Protophthiracarus subsellatus: Niedbała 1986b, 1992 , 2004 a, Niedbała & Liu 2018.

Steganacarus galeatus Balogh et Mahunka, 1978 syn. by Niedbała 1986b.

Steganacarus galeatus: Niedbała 1986b, 1992 .

Atropacarus (Hoplophorella) galeatus: Niedbała 1994a .

Hoplophorella (Hoplophorella) subsellata: Subías 2022 .

Neotropic: Brazil ( Balogh & Balogh 1987, 1988, 2002, Balogh & Mahunka 1977, 1978b, Niedbała 1986b, 1992, 1994a, 2004 a, Oliveira 2004, Oliveira et al. 2017, Subías 2022).

Neotropical, Brazil, endemic substrictus Atropacarus (Atropacarus) (Niedbała, 1983)

Steganacarus (Atropacarus) substrictus Niedbała, 1983

Atropacarus substrictus: Subías 2004 .

Atropacarus (Atropacarus) substrictus: Niedbała 1986b, 1992 , 2011b, 2012b, Niedbała & Liu 2018, Subías 2022.

Palaearctic: Georgia, Russia (South European Territory) ( Niedbała 1983a, 1986b, 1992, 2011b, 2012b, Subías 2022).

Western Palaearctic, known from Pontic subregion subtilis Mesoplophora (Parplophora) Niedbała, 1981

Mesoplophora subtilis Niedbała, 1981

Mesoplophora (Parplophora) subtilis: Niedbała 1985a, 2000 , 2004a, Mahunka 1998b, Liu & Chen 2010c, Liu 2016 a, Niedbała & Liu 2018, Subías 2022.

Neotropic: Antilles, Colombia, Honduras, Panama, Surinam, Venezuela, Peru, Brazil. Orient: Sri Lanka, Indonesia (Sumatra). Australasia: Papua New Guinea ( Balogh & Balogh 1988, 2002, Behan-Pelletier et al. 1993, Liu & Oconnor 2015, Mahunka 1998 b, Niedbała, 1981, 1985a, 2000, 2004a, 2012 a, Oliveira 2004, Oliveira et al. 2017, Subías 2022).

Pantropical, except Afrotropical Region succincta Oribotritia (Oribotritia) Niedbała, 1998

Oribotritia succincta Niedbała, 1998

Oribotritia succincta: Niedbała 2004d , Liu 2015 a, Niedbała & Liu 2018.

Oribotritia succinta (lapsus linguae): Subías 2004

Oribotritia (Oribotritia) succinta: Subías 2022 .

Afrotropic: Madagascar ( Balogh & Balogh 2002, Mahunka 2002, Niedbała 1998a, 2004d, 2017a, Subías 2022).

Afrotropical, Madagascar, endemic succinctus Steganacarus (Rhacaplacarus) Niedbała, 2001

Steganacarus (Rhacaplacarus) succinctus: Niedbała & Liu 2018 .

Rhacaplacarus (Rhacaplacarus) succinctus: Subías 2004 .

Hoplophorella (Rhacaplacarus) succincta: Subías 2022 .

Afrotropic: Democratic Republic Congo ( Niedbała 2001a, Subías 2022).

Afrotropical, Democratic Republic Congo, endemic sudafricanus Phthiracarus (Archiphthiracarus) Subías 2010 synonym of Phthiracarus humilis Niedbała, 2006 sudamericanus Austrophthiracarus ( Pérez-Íñigo et Baggio, 1993)

Archiphthiracarus sudamericanus Pérez-Íñigo et Baggio, 1993

Austrophthiracarus sudamericanus: Niedbała 2004a , Liu & Zhang 2016 a, Niedbała & Liu 2018, Niedbała & Ermilov 2022b.

Phthiracarus (Archiphthiracarus) sudamericanus: Subías 2022 .

Neotropic: Brazil, Argentina ( Balogh & Balogh 2002, Niedbała 2004a, Niedbała & Ermilov 2022b, Oliveira 2004, Oliveira et al. 2017, Pérez-Íñigo & Baggio 1993, Subías 2022).

Neotropical, Guyano-Brasilian and Patagono-Andean sudamericanus Euphthiracarus (Pocsia) Niedbała, 2004

Euphthiracarus (Pocsia) sudamericanus: Niedbała & Liu 2018 , Subías 2022.

Neotropic: Brazil ( Niedbała 2004 a, 2014 a, Oliveira 2004, Oliveira et al. 2017, Subías 2022).