Landouria ciliocincta ( Möllendorff, 1897 )

Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, European Journal of Taxonomy 526, pp. 1-73 : 61-62

publication ID	https://doi.org/ 10.5852/ejt.2019.526
publication LSID	lsid:zoobank.org:pub:3FFC527D-E9CA-4D9F-BF59-076FA60171AC
persistent identifier	https://treatment.plazi.org/id/03B1DB4C-8B11-4E24-4C28-FC983835FC36
treatment provided by	Plazi
scientific name	Landouria ciliocincta ( Möllendorff, 1897 )
status

Treatment

Landouria ciliocincta ( Möllendorff, 1897) View in CoL

Figs 45 View Figs 29–47 , 54 View Figs 52–55 , 69 View Figs 63–71 , 83–84 View Figs 81–84 ; Tables 3–4 View Table 3 View Table 4

Plectotropis ciliocincta Möllendorff, 1897: 67 (“ Java ”; more exactly ( Zilch 1966): “O-Java: Zuider- Gebirge”).

Landouria ciliocincta – van Benthem Jutting 1950: 466 View in CoL (in part). — Zilch 1966: 293, pl. 7, fig. 1 (in part). — Dharma 2015: 8, figs 2, 4 (in part). — Nurinsiyah et al. 2016: 559.

Diagnosis

Landouria ciliocincta is characterized by a whitish, almost disc-like, keeled shell with an only slightly elevated spire, a reddish-brown band at the angular periphery and a wide umbilicus, a long, cylindrical penis with a small bulge at its proximal end, and a short, very broad flagellum, which tapers towards its proximal end.

Material examined

Lectotype INDONESIA • lectotype (designated by Zilch 1966); East Java, Zuider (Southern) Mountains ; SMF 9176 View Materials a.

Paralectotype

INDONESIA • 1 spec.; same data as for lectotype; SMF 9716 View Materials b .

Other material

INDONESIA – East Java • 1 spec. (voucher for Dharma 2015); Malang, Ngliyep; 8°23′ S, 112°25′ E; near sea level; BDJ GoogleMaps • 1 spec. (det. anat.) (voucher for Nurinsiyah et al. 2016); Malang, Kondangmerak, nature/protected forest; 8°23′22″ S, 112°31′02″ E; 51 m a.s.l.; MZB 19211 • 1 spec. (voucher for Nurinsiyah et al. 2016); Malang, Kondangmerak, nature/protected forest; 8°23′21″ S, 112°31′05″ E; 100 m a.s.l.; MZB 17645 • 2 spec.; Tengger Mountains ; ZMB 45408 .

Description

SHELL (Figs 83–84; Tables 3–4 View Table 3 View Table 4 ). Almost disc-like with slightly elevated spire, with 5–6 moderately convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles, indistinct incised spiral lines and sparse scaly processes, mainly at periphery; tuberculate around umbilicus; whitishcorneous with a brownish band at periphery; body whorl keeled; aperture rounded rhombic; upper insertion of peristome descending; peristome expanded, reflexed and thickened; umbilicus slightly eccentric, wide, comprising 22–30% of shell diameter, hardly obscured by columellar edge.

GENITALIA ( Figs 45 View Figs 29–47 , 69 View Figs 63–71 ; Table 4 View Table 4 ). Atrium short; penis long, cylindrical, with a small bulge at its proximal end, without penis coecum; membranaceous sheath around distal part of penis connected with flagellum by tissue; penial retractor runs from diaphragm to middle part of cylindrical epiphallus; flagellum short, very broad, without node, tapering towards its proximal end; vas deferens narrow and long; vagina long; oviduct short, broad, proximally tapering; peduncle of bursa copulatrix subdivided into a broad, proximally tapering distal part and a narrower, longer proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina.

Remarks

Landouria ciliocincta shares with L. pakidulan sp. nov. and L. menorehensis sp. nov. the whitish shell with a brown band at the periphery. For differences, see remarks under L. pakidulan sp. nov.

Distribution

Landouria ciliocincta View in CoL is known from Malang Regency in East Java (Fig. 54). Records from other areas of Java ( Oostingh 1923; van Benthem Jutting 1941, 1950; Zilch 1966; Dharma 2015) and from Sumatra ( van Benthem Jutting 1959; Heryanto 2013) probably refer to other species.

References

Figures

Figs 29–47. Flagella of species of Landouria from Java. 29. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 30. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 31. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 32. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 33. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 34. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 35. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 36. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 37. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 38. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). 39. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 40. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 41. L. pacitanensis sp. nov., East Java, Pacitan, near Srau Beach (MZB 19231). 42. L. intumescens (Martens, 1867), East Java, Guwoterus (MZB 19207). 43. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). 44. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 45. L. ciliocincta (Möllendorff, 1897), East Java, Malang, Kondangmerak (MZB 19211). 46. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 47. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bar: 1 mm.