Scaphinotus (Pseudonomaretus) relictus (G. H. Horn) 1881

Kavanaugh, David H. & Angel, Kathleen W., 2015, A Taxonomic Review of the Northwestern Nearctic Subgenus Pseudonomaretus Roeschke 1907, including Description of the New Species Scaphinotus hoodooensis (Coleoptera: Carabidae: Cychrini) from the Bitterroot Mountains of Montana, Proceedings of the California Academy of Sciences 62 (14), pp. 381-396 : 393

Scaphinotus (Pseudonomaretus) relictus (G. H. Horn) 1881

Figures 2d, 3d , 6 , 12

RECOGNITION.— Adults of this species can be distinguished from those of other species of Pseudonomaretus by the following combination of character states: SBL = 13.0 mm or more (males 13.0–16. 6 mm, females 14.7–20. 3 mm); elytra moderately shiny, microsculpture shallowly impressed, without metallic reflection; head with a distinct transverse impression at level of posterior margin of eyes clearly delineating frontal and occipital regions; labrum ( Fig. 2d) with lateral lobes moderate in length and width; antennomere 4 ( Fig. 3d ) not pubescent, with only a very few scattered setae in addition to apical whorl of setae; pronotum with basolateral pair of setiferous pores present; male protarsomere 1 ( Fig. 6b ) with pad of adhesive setae ventrally over virtually entire ventral surface, protarsomere 4 without a pad; elytra with 13–16 apparent striae, with at least lateral striae disrupted by punctures in and between striae, elytral intervals 4 and 8 each with two to six discal setiferous pores; median lobe of male aedeagus ( Fig. 12) moderate in thickness and of nearly equal thickness throughout, ventral margin arcuate in basal one-fourth, straight in middle half, and slightly deflected ventrally in apical one-fourth in lateral aspect ( Fig. 12b), widest near basal one-fourth of shaft length, straight, gradually tapered to a flattened point apically in dorsal aspect ( Fig. 12a), armature of internal sac as in Fig. 12.

GEOGRAPHICAL DISTRIBUTION.— This species is known from southwestern Alberta and southeastern British Columbia, northwestern Montana, northern Idaho and eastern Washington ( Gidaspow 1973; Bergdahl 2014).

Figures

FIGURE 3. Digital photographs of right antenna, antennomeres 1 to 5, of Scaphinotus (Pseudonomaretus) species, dorsal aspect, with antennomeres identified by number; a, S. hoodooensis Kavanaugh and Angel sp. nov. (near Hoodoo Pass, Mineral County, Montana); b, S. merkelii (G. Horn) (St. Joe National Forest, Idaho); c, S. regularis (LeConte) (Hayden Creek, Kootenai County, Washington); d, S. relictus (G. Horn) (Selway River at O’Hara Campground, Idaho County, Idaho); e, S. mannii Wickham (Steptoe Canyon, Whitman County, Washington). Scale lines = 1.0 mm.

FIGURES 4–7. Digital photographs of male left protarsus of Scaphinotus (Pseudonomaretus) species, with tarsomeres identified by number; a, dorsal aspect; b, ventral aspect; as = pad of adhesive setae. Fig. 4. S. hoodooensis Kavanaugh and Angel sp. nov. (near Hoodoo Pass, Mineral County, Montana). Fig. 5. S. merkelii (G. Horn) (9 km SE of Lowell, Idaho County, Idaho). Fig. 6. S. relictus (G. Horn) (Selway River at O’Hara Campground, Idaho County, Idaho). Fig. 7. S. mannii Wickham (Steptoe Canyon, Whitman County, Washington). Scale lines = 1.0 mm.