Paraulax Kieffer, 1904
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publication ID |
https://doi.org/ 10.5281/zenodo.189597 |
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publication LSID |
lsid:zoobank.org:pub:AC41ACF9-2D19-45A2-96DE-16470E7D9C7F |
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DOI |
https://doi.org/10.5281/zenodo.5681147 |
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persistent identifier |
https://treatment.plazi.org/id/03ADE415-FFCA-FF9B-FF58-9064FA75B9D3 |
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treatment provided by |
Plazi |
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scientific name |
Paraulax Kieffer, 1904 |
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Type genus Paraulax Kieffer, 1904 .
Diagnosis. Differs from all other cynipids by the presence of the following autapomorphies: modified flagellomere of male antenna always F2, F3 or both (never F1 as in other Cynipidae ) ( Figs. 2 View FIGURE 2 E, 4G, 4I & 5A); presence of a structure of 4–5 rows of sharp, closely spaced and deep costulae on swelling of basal one-third of profemur ( Figs. 3 View FIGURE 3 B & 3C); presence of 5-9 vertical carinae in the ventral region of the gena ( Fig. 4 View FIGURE 4 B). The tribe can also be distinguished by a combination of the following characters: female antenna with 10 flagellomeres, F10 clavate ( Figs. 2 View FIGURE 2 C & 2D); ventral part of clypeus not or only slightly projecting over mandibles; dorsolateral margin of pronotal plate projecting laterally ( Figs. 2 View FIGURE 2 H & 2I); genal occipital carina present; scutellar foveae almost absent ( Figs. 6 View FIGURE 6 D & 9A) or, if present, always shallow or indistinct ( Figs. 2 View FIGURE 2 I & 4C), while the area posterior to the transscutal fissure always is concave; round, distinctly margined posteromedian scutellar impression absent; mesopleural impression present ( Figs. 2 View FIGURE 2 G, 4B & 5C).
Description. Predominantly black, sometimes brown to red-brown, with weakly shining or dull sculpture. Relatively small (1.7–2.9 mm). Female antenna with 10 flagellomeres, male with 13. Face usually with a distinct median vertical carina and facial strigae radiating from clypeus laterally ( Figs. 6 View FIGURE 6 A & 8A). Pronotal plate conspicuous with dorsolateral margins projected laterally. Notauli complete. Scutellar foveae faint or indistinct. Mesopleural impression present, more or less extended. Claws bifid, with a relatively long basal lobe or tooth ( Figs. 3 View FIGURE 3 D & 4D), or, if simple, sometimes with an acute tiny lobe ( Fig. 9 View FIGURE 9 D). Forewing with radial cell closed along anterior margin; R1 tubular albeit slightly depigmented along radial cell ( Figs. 3 View FIGURE 3 E & 4H); vein Rs+M and M, directed towards lower half of median vein ( Figs. 3 View FIGURE 3 E, 4K, 7F & 8H). Female metasoma laterally compressed. Abdominal petiole smooth dorsally, ventrally with deep longitudinal grooves. T2 smooth and shining, covering about 2/3 of metasoma. Projecting part of hypopygial spine relatively short, 3–4 times longer than wide ( Fig. 3 View FIGURE 3 G).
Diversity and distribution. Includes two genera occurring in the South of the Neotropical Region, in temperate Nothofagus forests of Argentina and Chile, one of them associated with galls of Aditrochus (Pteromalidae) on Nothofagus .
Remarks. A recent morphological phylogenetic study found Paraulax to be closely related to the Pediaspidini (Liljeblad et al. 2008). We justify the erection of a new tribe as follows: with several members, including a new genus, the new tribe itself is strongly supported by the molecular data of the present analysis (100% bootstrap) whereas the grouping with Pediaspis is only recovered with a 75% of bootstrap support; the new group is morphologically distinct and supported by several synapomorphies; the erection of a new tribe for the cynipids associated with Nothofagaceae is congruent with the traditional tribal classification of the Cynipidae , which to a large degree is based on biological associations such as higher level systematics of the host plants.
The tribe Paraulacini , as here defined, share with the Pediaspidini , among other character states, a relatively long pronotum medially, the pronotal plate being extended dorsally, a mostly smooth mesopleuron, usually with a visible mesopleural impression, and the scutellar foveae faint or absent. The two tribes can, however, be readily distinguished as follows:
1. Female antenna with 12 or more flagellomeres ( Fig. 1 View FIGURE 1 A); last flagellomere not wider than the penultimate; male antenna with modified F1 ( Fig. 1 View FIGURE 1 B). Ventral area of gena without vertical carinae. Ventral part of clypeus broadly projecting over mandibles ( Fig. 1 View FIGURE 1 C); genal carina absent. Dorsolateral margin of pronotal plate not projecting laterally ( Fig. 1 View FIGURE 1 D). Scutellar foveae absent ( Fig. 1 View FIGURE 1 E); area behind transscutal fissure flat or convex. A round, distinctly margined posteromedian scutellar impression present ( Fig. 1 View FIGURE 1 E). Mesopleural impression absent or faint ( Fig. 1 View FIGURE 1 F). Profemur not modified. Includes one Palaearctic genus inducing galls on Acer View in CoL and another genus from the Himalayan region with biology unknown ............................................................................................................ Pediaspidini
- Female antenna with 10 flagellomeres ( Fig. 2 View FIGURE 2 C); last flagellomere wider than the penultimate; male antenna with either F2, F3 or both modified ( Figs. 2 View FIGURE 2 F & 4G). Ventral area of gena with 5–9 vertical carinae ( Fig. 4 View FIGURE 4 B); genal carina present. Ventral part of clypeus at most slightly projecting over mandibles ( Fig. 2 View FIGURE 2 A). Dorsolateral margin of pronotal plate strongly projecting laterally ( Figs. 2 View FIGURE 2 H & 2I). Scutellar foveae absent or present, always shallow or indistinct, while the area posterior to the transscutal fissure always is concave ( Figs. 2 View FIGURE 2 G & 2I). Round, distinctly margined posteromedian scutellar impression absent ( Fig. 2 View FIGURE 2 I). Mesopleural impression present ( Figs. 2 View FIGURE 2 G & 7A). Profemur ventrally with a swelling composed of 4–5 rows of sharp, closely spaced, deep costulae ( Figs. 3 View FIGURE 3 B & 3C). Includes two neotropical genera associated with galls of Aditrochus (Pteromalidae) View in CoL on Nothofagus View in CoL ........................... Paraulacini
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