Tinearia Schellenberg, 1803

Ježek, Jan, Pont, François Le, Martinez, Eddy, Mollinedo, Sergio, Insad & Ibba, n Edificio, 2011, Three new species of non-biting moth flies (Diptera: Psychodidae: Psychodinae) from Bolivia, with notes on higher taxa of the subfamily, Acta Entomologica Musei Nationalis Pragae 51 (1), pp. 183-210 : 191

Tinearia Schellenberg, 1803

Tinearia Schellenberg, 1803 : pl. 40. Type species: Psychoda alternata Say, 1825 , designation by COQUILLET (1910: 615).

Ulomyia (partim) ( Tinearia as synonym of genus Ulomyia Haliday, 1838 in CURTIS (1838), not Ulomyia Walker, 1856 ): ENDERLEIN (1937: 84) (invalid designation of Trichoptera fuliginosa Meigen, 1804 as type species of Tinearia ); SARÀ (1952: 2).

Psychoda (partim) ( Tinearia as synonym of genus Psychoda Latreille, 1796 ): RAPP (1945: 28); WITHERS (1988: 69); IBÁÑEZ- BERNAL (1992: 97); DUCKHOUSE & LEWIS (2007: unpaginated); IBÁÑEZ- BERNAL (2008: 97).

Tinearia: JEŽEK (1977: 232) (reinstated as separate genus); JEŽEK (1983: 258); JEŽEK (1984a: 141); SASAKAWA (1986: 2, 3); WAGNER (1984: 4); WAGNER (1988: 12); WAGNER (1990: 45); WAGNER & BÁEZ (1993: 2); WAGNER (1994: 91); WAGNER & JOOST (1994: 85); JEŽEK & VAN HARTEN (1996: 70); WAGNER (1997a: 209, 213, 220, 222); WAGNER (1997b: 138, 140, 143–144); JEŽEK (2000: 27); WAGNER (2000: 244); WAGNER (2001: 89); BERNOTIENĖ (2002: 6); WAGNER (2003: 119); WAGNER & HRIBAR (2004: 510); JEŽEK & VAN HARTEN (2005: 217); WAGNER (2007: unpaginated); WAGNER & ANDERSEN (2007: 295); JEŽEK & VAN HARTEN (2009: 706).

Psychoda (Tinearia) (as subgenus of Psychoda Latreille, 1796 ): BRAVO et al. (2006: 3).

Only important references including synonymies are presented above.

Differential diagnosis. Antennomere 13 without a fold ( Figs. 40, 41 ), sensory filaments with small branches ( Fig. 50 ), shorter than flagellar nodes, terminal lobe of labium with five digitiform projections ( Fig. 42 ), dark patches of hairs near apices of wing veins developed ( Figs. 38, 39 ). Thus it differs from Ypsydocha Ježek, 1984 , which has antennomere 13 with conspicuous circular distal fold, sensory filaments with large branches, longer than flagellar nodes, terminal lobe of labium with three digitiform projections, and apices of wing veins without dark patches of hairs.

Discussion. The taxonomy of the tribe Psychodini is not fixed so far due to rather incompatible views of different psychodid specialists. JEŽEK (1984a) and JEŽEK & VAN HARTEN (1996) proposed a new classification for the species commonly ascribed to Psychoda Latreille, 1796 (pteropleurite trapezoidal = Psychodini in their sense) and the reinstatement of the genus Tinearia Schellenberg, 1803 ( JEŽEK 1977) was the first historical step in this reclassification. FAIRCHILD (1951) examined SCHELLENBERG’ s (1803) original publication (plates and the names appearing on them); the plate XL bears the supposed generic name Tinearia , but no specific name. No mention of the name Tinearia appears in the text by anonymous authors. COQUILLET’ s (1910) designation of Psychoda alternata Say, 1825 as the type species thus validates Tinearia . The generic classification of JEŽEK (1984) was partially incorporated into the Catalog of the Diptera of the Australasian and Oceanian Regions (DUCKHOUSE & LEWIS 2007) in contrast to the paper on a supraspecific classification of the genus Psychoda ( BRAVO et al. 2006) , where Tinearia is considered only one of 12 subgenera of the genus Psychoda sensu lato.

Currently there are 16 species of the genus Tinearia Schellenberg, 1803 distributed throughout the World including Tinearia boliviensis sp. nov. described in this paper. In addition, IPE et al. (1986) described three problematic species from India – Psychoda kishorei Ipe, Ipe & Kishore, 1986 , Psychoda panajiensis Ipe, Ipe & Kishore, 1986 , and Psychoda sanctijohani Ipe, Ipe & Kishore, 1986 – all of which are most likely synonyms of Tinearia alternata ( Say, 1825) , see the opinion of JEŽEK & VAN HARTEN (2009).

Figures

Figs. 40–48. Tinearia boliviensis sp. nov.,J and♀. 40 – apical antennomeres (♀); 41 – same (J); 42 – terminal lobe of labium and epipharynx (♀); 43 – maxilla and palpus maxillaris (♀); 44 – haltere (J); 45 – lateral view of gonostyle; 46 – lateral view of epandrium and surstylus; 47 – dorsal view of aedeagal complex and gonopods (a little deflected); 48 – dorsal view of epandrium and surstyli. Scale bars: 0.1 mm, except 0.05 in Figs. 41–42 and 45.

Figs. 49–58. Tinearia boliviensis sp. nov., J and ♀. 49 – basal antennomeres (♀); 50 – ascoidal filament (♀); 51 – lateral view of thoracic sclerites (J); 52 – last tarsomeres and claw of P 1 (♀); 53 – lateral view of gonopod; 54 – caudal view of aedeagal complex and gonocoxites; 55 – lateral view of terminal part of aedeagus; 56 – same, diagonal view; 57 – ventral view of ovipositor (from chloralphenol); 58 – female subgenital plate. Scale bars: 0.05 mm, except 0.1 in Figs. 51, 53–54 and 57.

Figs.34–39. Tinearia boliviensis sp. nov.,J and♀. 34–35 – frontoclypeus and facets (34 –J; 35 –♀); 36–37 – cibarium and epipharynx (36 – ♀; 37 – J); 38–39 – wing (38 – J; 39 – ♀). Scale bars: 0.1 mm, except 0.5 in Figs. 38–39.