Ancyropilus hornensis, Bartlett & Lambkin, 2022

Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, Zootaxa 5220 (1), pp. 1-81 : 25

Ancyropilus hornensis sp. nov.

ZooBank registration: urn:lsid:zoobank.org:act:91850EDB-374C-48AE-BB1E-28B7B0E321E2

( Figs 35 , 74 , 110 , 141 ; Map 1)

HOLOTYPE ♁: Northern Territory: Horn Islet Pellew Group, N.T. 15-21 Feb. 1968 B. Cantrell ( QM, type reg. T258549) . PARATYPES (2): Northern Territory: Horn Islet Pellew Group, N.T. 15-21 Feb. 1968 B. Cantrell (1 ♁, QM); Horn Islet Sir Edward Pellew Group, N.T. 15-21.ii.1968 B. Cantrell (1 ♀, QM) .

Diagnosis. Pronotum rotund (sub-globose); elytra dark with orange humeral fasciate and apical maculations (humeral maculae less well-defined), 8 th stria beginning near anterior margin of fascia or within fascia, striae reaching apical maculae, interstrial setae not dense as in other Ancyropilus species; femora yellow basally, brown apically, tarsi with three ventral tarsal pads.

Description. Habitus: Fig. 141 . Total length: 7.3–9.8 mm (holotype, 8 mm). Head: Vertex and frons black to dark brown, clypeus reddish black or brown, anteclypeus semi-transparent orange, labrum, antennae and palpi orange to orange-brown; eyes separated by about 0.54–0.65 eye widths (holotype, 0.54); vertex and upper part of frons punctate-rugulose, lower part of frons (above epistomal suture) transversely rugulose with occasional punctation near eye margins; clypeus relatively heavily punctate; genae and submentum wrinkled; exterior margins of terminal palpomeres about 1.5 times (maxillary) and 2 times (labial) the length of inside edges; antennae not reaching base of pronotum; eyes and cranium with long erect pale setae, frons and vertex with denser shorter medially-directed setae. Prothorax: Dark brown, pronotal arch and basal collar generally paler than disc, prosternum dark to pale brown; pronotum robust (sub-globose), about 1.14–1.25 times longer than wide (holotype, 1.14), sides rounded to subtly angulate, middle wider than pronotal arch; subapical depression v-shaped, central impression obsolete; disc moderately punctate though with a smooth reflective general appearance, punctures more dense laterally, a medial glabrous stripe in basal half, base with paired paramedical glabrous tumescences, anterior paramedical tumescent areas partly glabrous; disc with fine short multi-directional setae and sparser long erect setae. Pterothorax: Ventrites dark orange-brown, with fine pale posteriorly- or medially-directed setae; elytra dark brown with yellow-orange markings (each elytron with a weakly defined humeral macula running obliquely towards but not reaching the suture, a broad median fascia that crosses the suture and is broadest laterally—anteriorly meeting the humal macula in the female paratype, plus a trigonal apical macula), length to width ratio 2.62–2.72:1 (holotype, 2.72:1); 8 th stria beginning near anterior margin of median fascia, or within fascia (after about 7–10 punctures from base of the 7 th stria), all ten striae reaching apical macula, or 6 th and/or 10 th striae terminating before apical macula; punctation circular, without nodules, diameter of apical punctation similar or slightly smaller than diameter of basal punctures, striae separated by approximately one puncture diameter; epipleurae extending into apical maculae; interstrial setae generally arranged single-file along striae (two setae across strial width observed in striae posterior of median fascia); intrafoveal setae shorter than puncture diameter in the two observed males and as long as puncture diameter in the only known female specimen. Legs: Approximately basal half of profemora and basal three-fifths of mesoand metafemora yellow, remaining parts of femora brown; tibiae and tarsi brown, tibial carinae darker; ventral tarsal pads yellowish; meso- and metafemora slender, profemora weakly swollen. Abdomen: Ventrites brown. Male genitalia: Tegmen ( Fig. 35 ), widest near middle, slightly narrowed between middle and parameroid lobes (the apices of which are tapered to a slender, pronounced, slightly inwards-directed digitiform process), dorsal sinus just under one-third tegmen length, ventral sinus about as long as dorsal sinus, apodeme very short, about one-fifthteenth tegmen length; median lobe as in Fig. 74 ; pygidium as in Fig. 110 .

Etymology. This species is named after the type locality of Horn Islet, of the Sir Edward Pellow group of islands located in the south-west corner of the Gulf of Carpentaria, Northern Territory.

Biology. All three adult specimens were collected in mid-February.

Distribution (Map 1). Ancyropilus hornensis sp. nov. is so far known only from Horn Islet, a small rocky landmass about 2 km long, near the larger Centre Island.

Figures

FIGURES 27–47. Tegmina, dorsal: (27) Ancyropilus brigalowae, paratype; (28) Ancyropilus emmotti, holotype; (29) Ancyropilus exossuarius, paratype; (30) Ancyropilus monteithi, holotype; (31) Ancyropilus noonbahensis, holotype; (32) Ancyropilus packsaddlensis, holotype; (33) Ancyropilus simplex, paratype; (34) Ancyropilus tricolor, paratype; (35) Ancyropilus hornensis, paratype; (36) Culcipternotus mareebensis, paratype; (37) Infectostriatus absentis, holotype; (38) Infectostriatus differens, holotype; (39) Monilonotum bunyense, paratype; (40) Monilonotum doddi; (41) Monilonotum pascoei; (42) Monilonotum sundholmi, paratype; (43–47) Notopilo, beswickensis species group—(43) Notopilo beswickensis, holotype; (44) Notopilo calicis, paratype; (45) Notopilo tompricensis, holotype; (46) Notopilo xanthoimprocerus, holotype; (47) Notopilo xanthoprolatus, holotype (scale bars: 0.5 mm).

FIGURES 66–101. Median lobes: (66) Ancyropilus brigalowae, paratype; (67) Ancyropilus emmotti, holotype; (68) Ancyropilus exossuarius, paratype; (69) Ancyropilus monteithi, holotype; (70) Ancyropilus noonbahensis, holotype; (71) Ancyropilus packsaddlensis, holotype; (72) Ancyropilus simplex, paratype; (73) Ancyropilus tricolor, holotype; (74) Ancyropilus hornensis, paratype; (75) Culcipternotus mareebensis, paratype; (76) Infectostriatus absentis, holotype; (77) Infectostriatus differens, holotype; (78) Monilonotum bunyense, paratype; (79) Monilonotum pascoei; (80) Monilonotum sundholmi, paratype; (81–85) Notopilo, beswickensis species group—(81) Notopilo beswickensis, holotype; (82) Notopilo calicis, paratype; (83) Notopilo tompricensis, holotype; (84) Notopilo xanthoimprocerus, paratype; (85) Notopilo xanthoprolatus, holotype; (86–88) Notopilo, cambageicola species group—(86) Notopilo cambageicola, paratype; (87) Notopilo interfabulatus, paratype; (88) Notopilo tanybasilaris, paratype; (89) Notopilo congruus (congruus species group); (90) Notopilo reduncus (reduncus species group); (91–93) Notopilo, variipes species group—(91) Notopilo brevistriatus, paratype; (92) Notopilo gerstmeieri, holotype; (93) Notopilo variipes; (94–96) Notopilo, unplaced to species group—(94) Notopilo elstoni, holotype; (95) Notopilo katherinensis, holotype; (96) Notopilo obesus, paratype; (97) Platynotum bulli, paratype; (98) Platynotum culgoense, holotype; (99) Platynotum femorale; (100) Platynotum foveosetosa, holotype; (101) Platynotum gracile, holotype (scale bars: 0.5 mm).

FIGURES 102–137. Male pygidiae: (102) Ancyropilus brigalowae, paratype; (103) Ancyropilus emmotti, holotype; (104) Ancyropilus exossuarius, paratype; (105) Ancyropilus monteithi, holotype; (106) Ancyropilus noonbahensis, holotype; (107) Ancyropilus packsaddlensis, holotype; (108) Ancyropilus simplex, paratype; (109) Ancyropilus tricolor, paratype; (110) Ancyropilus hornensis, paratype; (111) Culcipternotus mareebensis, paratype; (112) Infectostriatus absentis, holotype; (113) Monilonotum doddi; (114) Monilonotum pascoei; (115) Monilonotum sundholmi, paratype; (116–120) Notopilo, beswickensis species group—(116) Notopilo beswickensis, holotype; (117) Notopilo calicis, paratype; (118) Notopilo tompricensis, holotype; (119) Notopilo xanthoimprocerus, paratype; (120) Notopilo xanthoprolatus, holotype; (121–124) Notopilo, cambageicola species group—(121) Notopilo cambageicola, paratype; (122) Notopilo interfabulatus, paratype; (123) Notopilo lawnhillensis, paratype; (124) Notopilo tanybasilaris, holotype; (125) Notopilo congruus (congruus species group); (126–128) Notopilo, variipes species group—(126) Notopilo brevistriatus, paratype; (127) Notopilo gerstmeieri, holotype; (128) Notopilo variipes; (129–132) Notopilo, unplaced to species group—(129) Notopilo confusus, holotype; (130) Notopilo elstoni, holotype; (131) Notopilo katherinensis, holotype; (132) Notopilo obesus, paratype; (133) Platynotum bulli, paratype; (134) Platynotum culgoense, holotype; (135) Platynotum femorale; (136) Platynotum foveosetosa, holotype; (137) Platynotum gracile, holotype (scale bars: 0.5 mm).

FIGURES 138–149. Habitus: (138) Ancyropilus brigalowae, paratype; (139) Ancyropilus emmotti, holotype; (140) Ancyropilus exossuarius, paratype; (141) Ancyropilus hornensis, paratype; (142) Ancyropilus labris, holotype; (143) Ancyropilus monteithi, paratype; (144) Ancyropilus noonbahensis, holotype; (145) Ancyropilus packsaddlensis, holotype; (146) Ancyropilus simplex, paratype; (147) Ancyropilus tricolor, paratype; (148) Culcipternotus mareebensis, holotype; (149) Infectostriatus absentis, holotype (scale bars: 1.0 mm).