Eulichas robusta Hájek, 2009

Hájek, Jiří, 2009, Revision of the genus Eulichas Jacobson, 1913 (Coleoptera: Eulichadidae) II. E. dudgeoni species group, Zootaxa 2192, pp. 1-44 : 29-31

Eulichas robusta Hájek , sp. nov.

( Figs. 14 , 35 , 55 , 75 )

Type locality. Malaysia, Pahang, Bukit Fraser, ca. 03°43’N 101°44’E, 1500 m.

Type material. 130 specimens — Holotype 3 ( NMPC), labelled: “ Malaysia 1500m / Bukit Fraser / 22.– 28. 11. 1998 / R. Kocina lgt. [printed]”. Paratypes: 233 (nos. 1–2) 1Ƥ (no. 3), same label data as holotype ( NMPC); 13 (no. 4) 2ƤƤ (nos. 5–6), “ MALAYSIA 24.11.[19]98 / Bukit fraser hill / H. Habal lgt. [printed]” ( NMPC); 13 (no. 7), “ MALAYSIA / BUKIT FRASER / 1. – 5. 5. 2003 / B. Makovský lgt. [printed]” ( NMPC); 533 (nos. 8–12), “ MALAYSIA / BUKIT FRASER / 25. – 31.3 2004 / M. Machytka lgt. [printed]” ( NMPC); 633 (nos. 13–18) 1Ƥ (no. 19), same label data, but “B. Makovský lgt. [printed]” ( NMPC); 1Ƥ (no. 20), “ MALAYSIA / BUKIT FRASER / 8. – 14. 4. 2008 / B. Makovský lgt. [printed]” ( NMPC); 13 (no. 21), “W. Malaysia: 1140 m / Fraser’s Hill, / Jeriau Road / 5–12.viii.1986 [printed] // G.S.Robinson / BM 1986–299 [printed]” ( BMNH); 1Ƥ (no. 22), “W. Malaysia: Pahang / Fraser’s Hill,ca 1300m /17–21.3.[19]93, light trap / Löbl&Calame, #14 [printed]” ( MNHG); 13 (no. 23), “ WEST MALAYSIA, PAHANG / 7.–10. 1. 1995 / FRASER’S HILL, 1500 m / 110 km N KUALA LUMPUR / lgt.S.BECVAR j. & s [printed]” ( BMNH); 633 (nos. 24–29), “ Malaysia: 10.V. / Fraser’s Hill / S.Snäll lgt. 1996 [printed]” ( SSCT); 533 (nos. 30–34), “ Malaysia Pahang Fraser’s / Hill 1200m 23–25.3 1992 light / B Gustafsson, H et H Hippa / G Sellerholm [printed]” ( NHRS); 1Ƥ (no. 35), “ MALAYSIA - Pahang / Banjaran Benom / Lata Jarom / 6.– 8.3.1997 / Ivo Jeniš leg. [printed]” ( NMPC); 13 (no. 36), same label data, but “Oliver Dulík leg. [printed]” ( NMPC); 233 (nos. 37–38) 2ƤƤ (nos. 39–40), “ MALAYSIA iii.1999 / Pahang & Kelantan prov. / Cameron Highlands. / lgt. local collector [printed]” ( NMPC); 1Ƥ (no. 41), “Weiden / Opf. [printed] / Cameron / Highlands [handwritten] / [on side:] STRÖHLE [printed]” ( VKCZ); 13 (no. 42), “ Malaysia / Pahang / Cameron Hids / 1000–1600ft / III– VII/1978 / C.M.Brandstetter [printed]” ( SMNS); 13 (no. 43), “ MALAYSIA / Tanah Rata / V. 1987. / leg. B. Molnár [handwritten]” ( HNHM); 13 (no. 44), “ MALAYSIA - Perak / Cameron Highlands / Tanah Rata / 13. – 16.3.1997 / Ivo Jeniš leg. [printed]” ( NMPC); 233 (nos. 45–46) 1Ƥ (no. 47), “ MALAYSIA W / CAMERON highl. / TANAH RATA env. / 23.– 26.3.2000 / Lgt. M. SNÍŽEK [printed]” ( NMPC); 13 (no. 48) 1 Ƥ (no. 49), “ MALAYSIA West, PAHANG / Cameron Highlands / TANAH RATA, 1200–1500m / 3.ii.– 19.ii. 2005 / Cechovsky Petr lgt [printed]” ( VKCZ); 13 (no. 50), “ MALAYSIA / TANAH RATA / 2. – 8. 4. 2008 / B.MAKOVSKÝ Lgt. [printed]” ( NMPC); 13 (no. 51), “ 28 VII 1991 U.V 750. / Rte TAPAH – TANAH RATA / PK 40 CAMERON HIGHLANDS / MALAYA Leg. J.HAXAIRE [printed]” ( NHMW); 13 (no. 52), “W Malaysia - Perak / road Tapah - Ringlet / 10km S of Ringlet mt. 900 / 14–19.IV.1999 / leg. A. Ballerio [printed]” ( NMPC); 333 (nos. 53–55), “ MALAYSIA W / CAMERON Highlands / RINGLET env. / 15.4.2000 / LGT. M. SNÍŽEK [printed]” ( NMPC); 13 (no. 56), “ MALAYSIA - PERAK Distr. 20.– 23.XI.2000 / Cameron Highlands / RINGLET env. / F. & L. Kantnerovi leg. [printed]” ( NMPC); 533 (nos. 57–61) 1Ƥ (no. 62), “ MALAYSIA W., PERAK / 40km SE of IPOH, 900 m / Banjaran Tai Wangsa / RINGLET, 29.iii.–15.iv. / 2004 Čechovsky Petr lgt. [printed]” ( NMPC, VKCZ); 13 (no. 63) 1Ƥ (no. 64), “ MALAYSIA –W; PERAK, ~ 900m; / 40 km SE IPOH; 4°25’N 101°23’E / Cameron Highlands; RINGLET; / M. Říha leg. 25.iv.–5.v.2001 [printed]” ( NMPC); 13 (no. 65), “ MALAYSIA, Pahang distr. / Cameron Highlands, 2001 / KAMPUNG KUALA BOH vill. env. / 4°27’N, 101°34’E, 850–1050m / R. Hergovits leg. 26. III.–3. IV. [printed]” ( NMPC); 13 (no. 66), “ MALAYSIA, Pahang distr. / Cameron Highlands, / KAMPUNG KUALA BOH vill. env., / N 4°27,9’, E 101°34,8’, 850–1050m, / 26. III.–3. IV. 2001 / K. Bucsek leg. [printed]” ( NMPC); 333 (nos. 67–69), “ MALAYSIA / Cameron / Highlands [printed]” ( ZSMC); 13 (no. 70), “W. Malaysia / Cameron / Highlands / 2000 ft. above / sea level. 18.2.[19]73 / C.C.Chua [handwritten] // Brit. Mus. / 1975-573 [handwritten]” ( BMNH); 13 (no. 71), same label data, but “5.3.[19]75” ( BMNH); 433 (nos. 72–75), “W – MALAYSIA / Cameron IV. / Highland 1987 [printed]” ( NHMW); 633 (nos. 76–81) 2ƤƤ (nos. 82–83), “ Cameron Highlands / MALAYSIA / late April 1987 [printed]” ( EUMC); 13 (no. 84), “Perak [handwritten] / Fry Coll. / 1905-100. [printed]” ( BMNH); 13 (no. 85), “64723 [handwritten] / Perak [printed] / Fry Coll. / 1905-100. [printed]” ( BMNH); 13 (no. 86), “ MALAYA [printed] / Perak / Gunong Kledang / 2500’ / 12 Feb 1959 [handwritten] / H. T. Pagden [printed] // C.I.E. COLL. / NO. [printed] 16488 [handwritten] // Eulichas sp. /? subocellata / Fairm. [handwritten] / R.D.Pope det. 1959 [printed] // Pres by / Com inst Ent / B M 1959-499 [printed]” ( BMNH); 13 (no. 87), W MALAYSIA, Perak distr., / 48 km NNE of IPOH, 2001 / FELDA LASAH vill. env., / 5°02’N, 101°12’E, 120–390 m / R. Hergovits leg. 13–21.III. [printed]” ( NMPC); 13 (no. 88), “W MALAYSIA, Perak distr., / FELDA LASAH vill. env., 48 km NNE of IPOH / N 5° 02,3’, E 101° 12,3’, 120–390 m, / 13.–21.III.2001 / P. Šomody leg. [printed]” ( NMPC); 1Ƥ (no. 89), “ MALAYSIA - Perak / Banjaran Bintang / Maxwell Hill (Taiping) / 18. – 19.2.1997 / Ivo Jeniš leg. [printed]” ( NMPC); 1Ƥ (no. 90), same label data, but “lgt. Oliver Ďulík [printed]” ( VKCZ); 13 (no. 91), “MALAYSIA-Perak / Bukit Larut / 23.– 25.2.2000 / K. Deneš jun. lgt. [printed]” ( VKCZ); 333 (nos. 92–94), “ WEST MALAYSIA, Perak / BUKIT LARUT (Maxwell Hill) / Taiping–Bintang Mts., (WGS84) / 04° 51’ 37’’N, 100° 47’ 56’’E / 9./ 13.7.2001, alt. 1100–1400m / lgt. R.Fouqué & H.Barlová [printed]” ( NMPC); 233 (nos. 95–96), “ Malaysia, Perak / Taiping, Bukit Larut 1036 m / 16.–17. 2. 1998 / V. Tichý lgt. [printed]” ( NKME); 13 (no. 97), “Taiping / Malaysia / 3.1975 [handwritten] // Eulichas / funebris / Westwood. / det. Stig Lundberg [handwritten]” ( NHRS); 13 (no. 98), “ Malaysia / Taiping / May 1987 [handwritten]” ( MCSN); 2ƤƤ (nos. 99–100), “ MALAYSIA / Taiping / X.1979 [handwritten]” ( MSCN); 1Ƥ (no. 101), “Taiping- Malaysia / VI-1982 [handwritten]” ( MSCN); 6ƤƤ (nos. 102–107), “ MALAYSIA / Taiping / V.1987 [handwritten]” ( MSCN, NMPC); 13 (no. 108), “ MALAYSIA - Kelantan / Banjaran Titi Wangsa / Kampong Lawa env. / 24. – 26.2.1997 / Ďulík & Jeniš leg. [printed]” ( NMPC); 633 (nos. 109–114) 1Ƥ (no. 115), “ MALAYSIA 10.–16.iv.1999 / Kelantan prov. / Kampong Raja env. / lgt. Mir. Janalík [printed]” ( NMPC, VKCZ); 13 (no. 116), same label data, but “lgt. Vít Kabourek [printed]” ( VKCZ); 433 (nos. 117–120), “ MALAY PENIN: [printed] / Selangor F.M.S. / Bukit Kutu 3457’ / April 1915 [handwritten] // Ex F.M.S. / Museum. / B.M. 1955-354. [printed]” ( BMNH); 13 (no. 121), “ MALAY PENIN: / Selangor. / Bukit Kutu [printed] / at light 3500 ft. / April 17th 1926 [handwritten] / H.M.Pendlebury. [printed] / [on reverse:] Ex. Coll: / F.M.S. Museum. [printed] // Ex F.M.S. / Museum. / B.M. 1955-354. [printed]” ( BMNH); 233 (nos. 122– 123), same label data, but “ April 20th ” ( BMNH); 13 (no. 124), “ Malaysia [handwritten]” ( MSCN); 13 (no. 125), “Pi Tam [Nakhon Si Thammarat prov., ca. 08°44’N 99°39’E] / S. Thailand / 8.VIII.1987 / M. Satô leg. [printed]” ( EUMC); 13 (no. 126) 3ƤƤ (nos. 127–129), “Saraburi [handwritten] / Thailand / leg. Sab. Steinke [printed] / [on side:] 1989 / XI. [handwritten]” ( NMPC, SMNS).

Additional material studied (doubtful localisation). 13, Philippines, Luzon, Sta Lucia Dolores Quezon, 500 m, at light, 25.vii.1991, J. Haxaire leg. ( NHMW).

Description. Habitus elongate, fusiform. Body colouring from brownish-red to brown. Pale part of setation consists of recumbent yellowish or yellow greyish setae forming indistinct ocellations on pronotum, and characteristic ocellations on elytra ( Fig. 14 ).

Measurements. Males: 26–33 mm (holotype 32 mm); females: 32–37 mm.

Head punctation consists of sparse, irregularly distributed moderately large setigerous punctures. Antenna long, slender, last antennomere rectangular, with apical elongation, 3.19–3.63 times as long as wide ( Fig. 35 ), its ventral side smooth.

Pronotum trapezoidal, ca. 2.01–2.04 times as wide as long. Sides almost regularly rounded ( Fig. 55 ). The disc with two indistinct rounded depressions. Punctation consists of sparse fine setigerous punctures on the disc, which become slightly coarser and denser laterally.

Elytra with numerous longitudinal rows of moderately large punctures, and very fine interstitial setigerous punctures.

Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally with distinct sinuation before apex.

Male. Aedeagus with phallobase of the same length as parameres. Parameres simple, broad, slightly attenuating to subapical hook. Their subbasal hook is missing, but the subapical hook is well developed and prominent. Median lobe lanceolate ( Fig. 75 ).

Female. Similar to male in habitus, but larger. Antenna shorter, and more slender.

Differential diagnosis. Eulichas robusta sp. nov. can be easily recognised from other Malayan species based on its large body size, regularly rounded sides of pronotum, and predominantly by the characteristic shape of aedeagus: Broad parameres with robust apical part, and distinctly lanceolate median lobe. However, the new species is very similar to E. gigantea known only from the female holotype. The only usable chatacter to distinguish between these species is the shape of pronotum, which is more rounded laterally in E. gigantea . Nevertheless, both species occur nearly 4.000 km from each other, and I do not expect them to be conspecific.

Collection circumstances. Collected at light.

Distribution. Known from number of localities in Malay Peninsula (Kelantan, Pahang, Perak, Selangor), and two localities in southern Thailand (Nakhon Si Thammarat, Saraburi).

Etymology. The Latin word for “robust” refers to generally large habitus of the species.

Figures

FIGURES 13 – 24. Habitus of Eulichas. 13 — E. oborili sp. nov. (holotype); 14 — E. robusta sp. nov. (holotype); 15 — E. sausai sp. nov. (holotype); 16 — E. serricornis (paratype, Felda Lasah); 17 — E. sikkimensis (Danda Pakhar); 18 — E. sundaensis sp. nov. (holotype); 19 — E. sundaensis sp. nov. Ƥ (Java); 20 — E. wewalkai sp. nov. (holotype); 21 — E. fasciolata (Crocker Range); 22 — E. jakli sp. nov. (holotype); 23 — E. subocellata (Tawau); 24 — E. villosa sp. nov. (paratype, Gunung Emas).

FIGURES 25 – 44. Male antennomeres IX – XI of Eulichas. 25 — E. alesbezdeki sp. nov.; 26 — E. bertiae; 27 — E. dudgeoni; 28 — E. jendeki; 29 — E. siamensis sp. nov.; 30 — E. similis sp. nov.; 31 — E. uniformis; 32 — E. baeri; 33 — E. incisicollis; 34 — E. oborili sp. nov.; 35 — E. robusta sp. nov.; 36 — E. sausai sp. nov.; 37 — E. serricornis sp. nov.; 38 — E. sikkimensis; 39 — E. sundaensis sp. nov.; 40 — E. wewalkai sp. nov.; 41 — E. fasciolata; 42 — E. jakli sp. nov.; 43 — E. subocellata; 44 — E. villosa sp. nov. Scale bar 1 mm.

FIGURES 55 – 64. Pronotum of Eulichas. 55 — E. robusta sp. nov.; 56 — E. sausai sp. nov.; 57 — E. serricornis sp. nov.; 58 — E. sikkimensis; 59 — E. sundaensis sp. nov.; 60 — E. wewalkai sp. nov.; 61 — E. fasciolata; 62 — E. jakli sp. nov.; 63 — E. subocellata; 64 — E. villosa sp. nov. Not in Scale.

FIGURES 72 – 77. Aedeagus of Eulichas in dorsal view. 72 — E. baeri; 73 — E. incisicollis; 74 — E. oborili sp. nov.; 75 — E. robusta sp. nov.; 76 — E. sausai sp. nov.; 77 — E. serricornis sp. nov. Scale bar 1 mm.