Systaria bregibec, Jäger, 2018

Jäger, Peter, 2018, On the genus Systaria (Araneae: Clubionidae) in Southeast Asia: new species from caves and forests, Zootaxa 4504 (4), pp. 524-544 : 533-535

Systaria bregibec spec. nov.

Figs 27–28 , 45–46

Type material: CAMBODIA: Kep Province: Holotype: male, Rabbit Island , 10°26’11.45''N, 104°19'12.98''E, 21 m elevation, disturbed secondary forest, forest floor and foliage, at night, P. Jäger leg. by hand, 9.7.2017 ( SMF).

Etymology. The species name is a combination of three Latin words: “brevis”, meaning “short”, referring to the short embolus, “gibber”, meaning “hump”, referring to the distinct ventral hump on the male palpal tibia, and “beccus”, meaning “beak”, referring to the beak-shaped distal part of the RTA in retrolateral view ( Figs 27–28 ). All three character states are characteristic for the species group with currently three species included (see diagnosis and discussion); term in apposition.

Diagnosis. Small spiders with body length of males: 9.1. Male copulatory organs similar to S. insolita Dankittipakul & Singtripop 2011 and S. hainanensis Zhang, Fu & Zhu 2009 in having a short embolus, a beakshaped (double-headed) RTA tip and a distinct ventral hump at the palpal tibia, but can be distinguished by both processes of distal RTA pointed in retrolateral view (blunt in S. hainanensis ) and of similar size (dorsal process distinctly more prominent in S. insolita ) ( Figs 27–28 ).

Description. Male (holotype): prosoma length 4.5, prosoma width 3.1, anterior width of prosoma 2.1, opisthosoma length 4.6, opisthosoma width 2.1. Eyes: AME 0.29, ALE 0.25, PME 0.20, PLE 0.26, AME–AME 0.16, AME–ALE 0.06, PME–PME 0.36, PME–PLE 0.24, AME–PME 0.18, ALE–PLE 0.02. Cheliceral furrow with 3 promarginal and 2 retromarginal teeth. Spination: Palp: femur p001, d012, patella–tarsus with bristles, not counted in spination; femur I p0011, d111, r110 (small), II p0111, d111, r110 (small), III p111, d111, r111, IV p1101, d111, r111; patella I–IV 000; tibia I p01, v 220, II p01, v220(2120), III–IV p11, d010, r11, v222; metatarsus II v100, III p112, r112, v202, IV p112, r122, v222. Leg formula: 4123. Measurements of palp and legs: Palp 4.4 (1.8, 0.7, 0.7, -, 1.2), I 13.9 (3.9, 1.7, 3.6, 3.0, 1.7), II 13.3 (3.7, 1.7, 3.4, 2.9, 1.6), III 11.6 (3.3, 1.4, 2.6, 2.9, 1.4), IV 16.6 (4.5, 1.7, 3.9, 4.6, 1.9). Palp as in diagnosis ( Figs 27–28 ). Cymbium two times longer than tibia. RTA arising distally from tibia with its disto-dorsal margin straight, disto-ventrad. Embolus arising in 9-o’clock-position from tegulum, slightly bent, with broad base, its tip situated close to tip of membranous conductor in 12-o’clockposition. Broad part of spermophor visible retrolaterally and proximally, S-shaped narrow part visible proximally to embolus base. Colouration. Deep yellowish- to reddish-brown without pattern. Fovea black, chelicerae, labium, gnathocoxae deep reddish-brown, sternum marginally darker brown. Opisthosoma dorsally with one pair of black spots and some black transverse striae in front of spinnerets. Dark setae around spinnerets

Female: unknown.

Distribution. Known only from the type locality ( Figs 45–46 ).

Natural history. The specimen has been found on a slope in a degraded secondary forest, with dry and weak leaf litter layer.

Figures

FIGURE 45. Records of Systaria spp. in SE Asia described in this paper. Triangles—species recorded from caves, circlesspecies recorded from forest.

FIGURE 46. All known records of the genus Systaria (World Spider Catalog 2018) including undescribed species mentioned in Deeleman-Reinhold (2001) according to habitats. Triangles—species recorded from caves, circles—species recorded from forest. See discussion.

FIGURES 27–32. Systaria spp., copulatory organs. 27–28 Systaria bregibec spec. nov., holotype male from Cambodia, Rabbit Island. 29–30 Systaria bifidops spec. nov., holotype male from Malaysia, Bukit Fraser. 31–32 Systaria panay spec. nov., holotype female from Philippines, Panay Island. 27–30 Left palp (27, 29 ventral, 28, 30 retrolateral). 31 Epigyne, ventral. 32 Vulva, dorsal. Abbreviations: BP—bifid process of RTA, CO—copulatory opening, LO—loop of copulatory ducts, TPproximal projection of tegulum, VH—ventral hump of palpal tibia.