Dicotyledoneae DE CANDOLLE 1819

Halamski, Adam T. & Kvaček, Jiří, 2013, The Type Specimen Of Debeya (Dewalquea) Haldemiana Rediscovered, Acta Musei Nationalis Pragae Series B 69 (1 - 2), pp. 83-86 : 83-84

Class Dicotyledoneae DE CANDOLLE 1819

Subclass, order, and family incertae sedis

Informal group Debeya sensu KRASSILOV, LEWY, NEVO et SILANTIEVA 2005

1873

1880

Text-figs 1A–G

Dewalquea haldemiana DEBEY ex SAPORTA et MARION ( Araliophyllum haldemianum DEBEY in litt.) p. 60; pl. 7, figs 1–2.

Dewalquea haldemiana DEBEY ex SAPORTA et MARION var. latifolia HOSIUS et VON DER MARCK p. 173; pl 35, fig. 114, pl. 34, fig. 115.

1880

1889

2013

Dewalquea haldemiana DEBEY ex SAPORTA et MARION var. angustifolia p. 173; pl 33, figs 116, 117, pl. 34, figs 118-121.

Dewalquea haldemiana (DEBEY ex SAPORTA et MARION) ROEMER , p. 143, pl. 12, figs 2, 3, 4

Debeya haldemiana (DEBEY ex SAPORTA et MARION) KNOBLOCH ; Halamski p. 422; figs 7B, C, 8C, 9C, D, F, 10C [ubi syn.].

L e c t o t y p e d e s i g n a t e d h e r e: MGUWr 7536p (text-figs 1A–G).

T y p e l o c a l i t y: Haldem, Westphalia, Germany.

T y p e h o r i z o n: Campanian, Late Cretaceous.

E m e n d e d d i a g n o s i s. Leaf compound, pedate, petiolate, with 6–7 petiolulate to subsessile linear leaflets; primary venation pinnate, secondary venation brochidodromous; leaflet apices acuminate; tertiary venation ramified.

D e s c r i p t i o n. The lectotype shows a compound, pedate, petiolate leaf composed of six leaflets (text-fig. 1 A). Petiole 28 mm long and 2.5 mm wide; inflated proximally, 3.7 mm wide. Petiolules very short (text-fig. C), bases of the leaflets decurrent; sometimes blades begin so closely to the insertion point of the petiolule that the leaflet is subsessile. Leaflets are linear, the median one 210 mm long, the lateral ones 200 mm long, up to 13 mm wide; margins entire, parallel in the median region; apex acuminate. Venation is pinnate consisting of strong primaries. Secondaries are brochidodromous, subopposite, excurrent, originating at an angle of 40–70 o, first running towards the margin, then subparallel to it, at least looping and joining the higher veins at 1/8 of the blade width (text-figs 1E-G); secondary spacing slightly decreasing or increasing distally depending on the leaflet considered. Intersecondaries are absent. Tertiary veins are poorly preserved, apparently forming a ramified pattern .

Figures

Text-fig. 1. Debeya (Dewalquea) haldemiana (DEBEY ex SAPORTA et MARION 1873) HALAMSKI 2013. Lectotype MGUWr 7536p; Haldem, Westphalia, Germany; Campanian. Scale bars 10 mm. A. General view of the specimen. B. Enlargement of the acuminate apex of a leaflet. C. Enlargement of the basal part with the petiole and the petiolules. D, E, F, G. Enlargements of the venation.