Satyrites reynesii,

Jong, Rienk De, 2017, Fossil butterflies, calibration points and the molecular clock (Lepidoptera: Papilionoidea), Zootaxa 4270 (1), pp. 1-63: 36-37

publication ID 10.5281/zenodo.583183

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Satyrites reynesii


reynesii  . Satyrites reynesii Scudder, 1872 

Nymphalidae  : Satyrinae  .

France, Bouches-du-Rhône, Aix-en-Provence; late Oligocene –early Miocene.

Depository: MVMF (holotype).

Published figures: Leestmans (1983: Figs 9–10); Scudder (1875: Pl. I Figs 2View FIGURE 2, 5View FIGURE 5).

The original description ( Scudder 1872) is repeated by Scudder (1875), who erected a new genus for the fossil: Lethites  . Kozlov (1988) accepted Scudder’s genus switch. The poorly preserved specimen bears only the forewings, which yield some usable characters. Subcosta basally strongly swollen; radial formula 1, 2, 3+(4+5), cell closed. A swollen subcosta is typical, but not universal, for Satyrinae  ( Nymphalidae  ), but this character occurs in some genera of the Biblidinae  as well ( Biblis  , Ergolis  , Eurytela  , Hypanis  , Vila  ). In Nymphalinae  with swollen Sc, the cell is usually open or at least, the ldc is weak. In the fossil ldc seems to be fairly strong. Thus, it is plausible to consider the fossil a member of the Satyrinae  . According to Scudder it is similar to the recent genera Lethe  and Cercyonis  , but it is not clear why it should be more similar to these genera than to any other satyrine genus. The similarity is not an indication of relationship, since Lethe  and Cercyonis  are only remotely related ( Wahlberg et al. 2009). Moreover, in Lethe  there are no basally swollen veins in the forewing ( Ackery et al. 1999).

Neorinopsis, Pseudoneorina  , Neorinella  and Satyrites  have been classified, without comment, as fossil genera belonging to the subtribe Lethina  of the tribe Satyrini  by Wahlberg & Peña (2015), but these choices are not supported by apomorphies. For further details, see the discussion under Neorinella garciae  and Neorinopsis sepulta  .