Kury, Adriano B., 2015, Opiliones are no longer the same — on suprafamilial groups in harvestmen (Arthropoda: Arachnida), Zootaxa 3925 (3), pp. 301-340 : 324-328

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Phylogeny of Cyphophthalmi

The mite-harvestmen ( Cyphophthalmi ) are here presented in more detail, separated from the rest of the Opiliones . In Figs. 20 View FIGURE 20 to 23, are presented relevant hypotheses found in the literature concerning the branching pattern of the Cyphophthalmi . The six following genera are here used to represent the major groups of Cyphophthalmi as currently understood, because they are the oldest in each family: Parogovia Hansen, 1921 (for Neogoveidae Shear, 1980 ), Ogovea Roewer, 1923 (for Ogoveidae Shear, 1980 ), Pettalus Thorell, 1876 (for Pettalidae Shear, 1980 ), Siro Latreille, 1796 (for Sironidae C.L. Koch, 1839 ), Stylocellus Westwood, 1874 (for Stylocellidae Hansen and Sørensen, 1904 ) and Troglosiro Juberthie, 1979 (for Troglosironidae Shear, 1993 ).

Hypothesis C 1 ( Fig. 20 View FIGURE 20 ): Shear (1980) published the first cladistic analysis of the Cyphophthalmi , unfolding the group into five families, which resulted in a symmetrical arrangement. Shear called his two major clades Tropicophthalmi and Temperophthalmi. Later, Shear (1993) added a sixth family, Troglosironidae .

Hypothesis C 2 ( Fig. 21 View FIGURE 21 ): Giribet & Boyer (2002) attempted both a morphological and a molecular analyses, which turned out very unresolved and it is not shown here. With a much expanded matrix, Boyer et al. (2007) made an important molecular analysis of the Cyphophthalmi , but failed to include Ogoveidae because of a lack of material for molecular study. This asymmetrical tree has Stylocellidae as sister group to the rest.

Hypothesis C 3 ( Fig. 22 View FIGURE 22 ): Clouse et. al. (2010) analyzed the Stylocellidae , including also the other Cyphophthalmi families. They used molecular data, and also did not include Ogoveidae . It was substantially different from Boyer’s analysis. They have not proposed any superfamilial names, but did recover the clade later known as Boreophthalmi.

Hypothesis C 4 ( Fig. 23 View FIGURE 23 ): Giribet et al. (2011) finally obtained material of Ogoveidae for molecular study. The result showed Pettalidae as sister group of the rest as in Clouse et al. (2010) and some novel groups. They recovered only one of Shear’s clades, the Ogoveoidea. Three new infraordinal names were proposed.