Rhyparus philippinensis Arrow, 1905

Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija & Medina, Milton Norman D., 2021, A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae), Acta Entomologica Musei Nationalis Pragae 61 (1), pp. 99-111 : 108-109

Rhyparus philippinensis Arrow, 1905

( Figs 6a–c , 12 , 18 , 24 , 30 a-b)

Type material. Sඒඇඍඒඉൾඌ: 1 J, labelled: ʻPhil. Isl.ʼ [round label] // ʻSyntype ʼ [round label] // ʻNHMUK 014189207ʼ ( BMNH); 1 ♀, labelled: ʻPhil. Islʼ [round label] // ʻ45 49ʼ [round label] // ʻSyntypeʼ [round label] // ʻNHMUK 014189209ʼ ( BMNH); 1 ♀, labelled: ʻPhil. Islʼ [round label] // ʻSyntypeʼ [round label] // ʻNHMUK 014189208ʼ ( BMNH); 1 ♀, labelled: ʻPhil. Islʼ [round label] // ʻSyntypeʼ [round label] // ʻNHMUK 013906231ʼ ( BMNH).

Additional material examined. PHILIPPINES: Lඎඓඈඇ: 1 ♀, Lagonoy, Camarines Sur, Bicol, vii.2018, local coll. ( CNCW); 1 ♀, Cagayan, Santa Ana, viii.2018, local coll. ( DUBC); 1 J, Barlig Mt. Province, xii.2018, local coll. ( LMCN). Mංඇൽൺඇൺඈ: 1 ♀, New Albay, Compostela Valley, vii.2018, local coll. ( CNCW); 1 J, Wao, Lanao del Sur, x.2018, local coll. ( CNCW); 1 ♀, Alamada, Lanao del Sur, x.2018, local coll. ( CNCW).

Diagnosis. Medium-sized species: body length 5.0– 6.5 mm ( Figs 6a–c ). Head as depicted in Fig. 12 ; epipharynx as in Fig. 18 . Intermediate lobes of pronotum as large as anterior one; median intercosta with dense punctation; second and third intercostae with dense punctation basally. All intercostae of elytra with two rows of medium-sized punctures. Internal and median protrusion of caudal bulbs connected ( Fig. 24 ); surface between external protrusion of caudal bulb of elytra and lateral margin of elytra distinctly sinuate – external protrusion more or less developed; however, median and internal ones always with the same proportions. Aedeagus as in Figs 30a–b .

Variability. Lateral lobes of pronotum slightly variable in size and proportions. Punctures of intercostae of pronotum and elytra can be proportionally variable in size between the specimens. Costae of pronotum and elytra are slightly variable in height and width.

Distribution. Philippines: Luzon, Mindanao.

Figures

Fig. 6. Habitus of Rhyparus philippinensis Arrow, 1905, syntypes: a – dorsal view; b – right dorsolateral view; c – right lateral view. Scale bar = 1.0 mm.

Figs 7–12. Head of Rhyparus species, frontal view: 7 – R. barclayi Masumoto, Ochi & Ho, 2019; 8 – R. flavohirtus sp. nov.; 9 – R. gracilis Arrow, 1905; 10 – R. helophoroides Fairmaire, 1893; 11 – R. mindanaoensis sp. nov.; 12 – R. philippinensis Arrow, 1905. Scale bar = 1.0 mm.

Figs 13–18. Epipharynx of Rhyparus species: 13 – R. barclayi Masumoto, Ochi & Ho, 2019; 14 – R. flavohirtus sp. nov.; 15 – R. gracilis Arrow, 1905; 16 – R. helophoroides Fairmaire, 1893; 17– R. mindanaoensis sp. nov.; 18 – R. philippinensis Arrow, 1905. Scale bar = 0.5 mm.

Figs 19–24. Elytra of Rhyparus species, oblique apico-dorsolateral view: 19 – R. barclayi Masumoto, Ochi & Ho, 2019; 20 – R. flavohirtus sp. nov.; 21 – R. gracilis Arrow, 1905; 22 – R. helophoroides Fairmaire, 1893; 23 – R. mindanaoensis sp. nov.; 24 – R. philippinensis Arrow, 1905. Scale bar = 1.0 mm.

Figs 25–30.Aedeagus of Rhyparus species, a – lateral view, b – dorsal view. 25 – R.barclayi Masumoto, Ochi & Ho, 2019; 26 – R. flavohirtus sp. nov.; 27 – R. gracilis Arrow, 1905; 28 – R. helophoroides Fairmaire, 1893; 29 – R.mindanaoensis sp. nov.; 30 – R.philippinensis Arrow, 1905. Scale bar = 0.5 mm.