Ommatius (Pygommatius) imaginus, Scarbrough & Marascia, 2003

Scarbrough, Aubrey G. & Marascia, Claudio G., 2003, Revision of Ommatius Wiedemann (Diptera: Asilidae). IV. Pygommatius subgen. nov. with twenty-five Afrotropical species, Zootaxa 228 (1), pp. 1-94 : 37-38

Ommatius (Pygommatius) imaginus , sp. n.

Figs. 11 , 41 , 139­146

Male. As O. comosus except as follows. Length, body 11.3­13.3 mm; wing 8.7­9.0 mm. Head: FHWR 1.09.8­1.0:11.5. Antenna brown­yellow to yellow, pedicel and scape darkest dorsally. Occiput with 9 brown postocular bristles, extending as a row to mid­lateral margin of head.

Thorax: Pronotum brown. Pleuron mostly yellow, anepisternum, katepisternite, anterior anepimeron, and meron at least brown. Katatergal bristles yellow.

Wing as O. comosus except apex of cell m 3 truncate ( Fig. 11 ).

Leg: Hind trochanter with 2­3 short brown bristles. Fore femur (see O. brevicornis Fig. 24 ) with 9 long, stout ventral bristles. Hind femur (see O. comosus , Fig. 31 ) with base entirely, dorsobasal third, slightly more than basal half anteriorly and posteriorly yellow, brown otherwise; 9­10 long, thin, brown, anteroventral setae present, most aligned basally in a row to brown marking anteriorly, 4­5 clustered at base of row; one short, brown, ventrobasal bristle present; a row of 7­8 short, thin to thick, bristles present posteriorly; abundant, mostly wavy, setae ventrally and posteriorly, that on apical half longest; HFWLR 1.0:4.0:1.0:4.4. Fore and middle tibiae yellow with thin, yellow bristles, longest bristle three­fourths as fore or middle tibia. Hind tibia ( Fig 41 ). Fore tarsus with 1 yellow bristle; remaining tarsomeres of all tarsi brown.

Abdomen: Margins of most segments yellow, those on terga 1­4 laterally wide; sterna 1­4 yellow to brown­yellow. Tergum 3 with 3­4 brown bristles in apical corner; tergum 4 with dense yellow setae laterally, bordered dorsally with several brown bristles. sternum 3 with only sparse short wavy setae medially. Sternum 4 with long, wavy, dense, yellow setae; sternum 5 bare, sternum 6 with an admedial stripe of mostly brown, erect setae, those basally yellow. Sternum 7 with 1 stout brown bristle in apical corner; sternum 8 with brown bristles apically.

Terminalia ( Figs. 139­143 ): As O. comosus except dorsal branch wider and ventral branch without median spine. Gonostylus large, podiform.

Female. Differs from male as follows. Length, body 9.7­11.7 mm; wing 7.5­9.0 mm. Head: Face yellow tomentose, 6 thin brown bristles and thick, yellow, ventral bristles present; scape sometimes yellow; 2­3 brown postocular bristles present: 1.0:11.2­11.8. Thorax: Pronotum reddish­yellow or brown. Pleuron with 0­7 brown katatergal bristles. Leg: Normal, without unusual setae and bristles of male. Femora brown­yellow to brown on apical third to half except ventrally on fore and middle femora. Middle and hind femora with most or all ventral bristles brown; hind femur with abundant, bristly brown setae posteroventrally. Fore and middle tibiae with narrow apex light brown, bristles unusually long, thin, mostly yellow. HFWLR 1.0:5.7­1.0:5.8. Hind tibia normal. Fore tarsus with mostly or only brown bristles. Abdomen: Wide lateral margin of tergites 1­6 yellow; sternites 1­2 entirely or mostly yellow; margins of remaining sternites narrowly yellow. Most tergites and sternites with brown bristles apically; most sternites with 3­4 brown bristles laterally. Tergite 9 short, membranous, as a thin line dorsomedially.

Terminalia ( Figs. 144­146 ): Three spermathecae present; duct basal. Sternite 8 apically triangular, sides distinctly arched, membranous medially with surface folded as a low, arched, ridge in dry specimens; apical margin rounded with a trumpet­like process anteriorly.

SPECIMENS EXAMINED. Holotype ♂, CAMEROON: 50 km N. Kribi / Rt. 87 / 30.xi.1987 / A. Freidberg ( NMSA) . Paratype: CAMEROON: 1 ♂, Bambalang / Off Rt. N11 / 35 Km E. Bamenda / 18, 21.xi.1987 / Amnon Freidberg ( NMSA) ; 1 ♀, Kribi , / 11.vii.1974 ( ZMAN) ; 1 ♀, Prov. Sud / 21 km SE Kribi / E­slope Mt. Elephant / 2°48.2N’ 10°01.9’E / 10.ii.1994 alt. 140 m / J. J. Wieringer ( ZMAN) ; Douala / 26­7­1975 / Cameroun Litt / J. A. W. Lucas ( ZMAN) . CONGO; 1 ♂, Museum Paris / Congo Ogooué / Sam Kita / R. Ellenberger 1910 ( MNHN) ; 1 ♀, R. P. Congo / Dimonika / Mayombe v.1979 / J. David ( MNHN) ; 1 ♀, Seve / Elacis / guineensis / (Palmacés) / 26.x.73 ( MNHN) ; 1 ♀, Congo Museum Paris / Congo Français / Ogoouè / N. Gomo / P. Ellenberger 1910 ( NMHN) .

Distribution. A forest species captured during October to July in Cameroon and the Republic of Congo.

Etymology. Latin imaginus , for ‘likeness’ or ‘image of, referring to its similarity to O. comosus .

Remarks. Ommatius imaginus differs from O. comosus in the sparse setae on sternite 3, fringe of yellow bristles bordered dorsally with brown bristles, and the combined characters of the male terminalia ( Figs. 139­143 ), especially the absence of an apical spine on the epandrium and the larger podiform gonostylus. Females differ from those of O. comosus in that the spermathecal duct arise directly at the base of the spermatheca ( Fig. 144 ) and the triangular apical margin of sternite 8 are slightly convex ( Fig. 146 ).

Figures

FIGURES 11­20. Right wing of male Ommatius (Pygommatius) spp. 11. O. imaginus, sp. n..; 12. O. limbus, sp. n..; 13. O. litoreus sp. n.; 14. O. lulua, sp. n.; 15. O. magnipes, sp. n.; 16. O. ornatipes Engel; 17. O. pectinus sp. n.; 18. O. porticus, sp. n.; 19. O. renudus, sp. n.; 20. O. strigiatus, sp. n.

FIGURES 39­48. Left hind tibia of male Ommatius (Pygommatius) spp. 39. O. comosus, sp. n.; 40. O. fluvius, sp. n.; 41. O. imaginus, sp. n.; 42. O. limbus, sp. n., anterior; 43. O. litoreus, sp. n.; 44. O. neglectus Bromley; 45. O. porticus, sp. n.; 46. O. renudus, sp. n.; 47. O. strigiatus, sp. n.; 48. O. vultus, sp. n., inner surface.

FIGURES 139­146. Male terminalia of Ommatius imaginus, sp. n 139­141. Lateral, dorsal and ventral views; 142. Gonostylus; 143. Aedeagus, lateral view. Female terminalia. 144. Spermatheca; 145. Genital fork; 146. Sternite 8, ventral view.

FIGURES 21­28. Right wing of male Ommatius (Pygommatius) spp. 21. O. talus, sp. n.; 22. O. vultus, sp. n.; 23­28. Right fore femur of male Ommatius (Pygommatius) spp. showing arrangement of anteroventral bristles. 23. O. brevicornis Curran; 24. O. grossus, sp. n.; 25. O. limbus, sp. n.; 26. O. litoreus, sp. n.; 27. O. strigiatus, sp. n..; 28. O. vultus, sp. n.

FIGURES 29­38. Male Ommatius (Pygommatius) spp. 29. O. jaculator Walker, middle femur. 30­ 38. Hind femur, 30. O. brevicornis Curran, anterior view; 31. O. comosus, sp. n., anterior view; 32. O. dasypogon Oldroyd, anterior view; 33. O. jaculator Walker, anterior view; 34. O. limbus, sp. n., with coxa, anterior view; 35. O. litoreus, sp. n., anterior view; 36. O. neglectus Bromley, anterior view; 37. O. strigiatus, sp. n., posterior view; 38. O. vultus, sp. n., posterior view.