Ommatius (Pygommatius) strigiatus, Scarbrough & Marascia, 2003

Scarbrough, Aubrey G. & Marascia, Claudio G., 2003, Revision of Ommatius Wiedemann (Diptera: Asilidae). IV. Pygommatius subgen. nov. with twenty-five Afrotropical species, Zootaxa 228 (1), pp. 1-94 : 56-57

Ommatius (Pygommatius) strigiatus , sp. n.

Figs. 20 , 27 , 37 , 47 , 210­217

Male. Black. Length, body 8.1­9.5 mm; wing 5.7­6.7. Head: Yellowish­gray to gray tomentose, white setose. Face with sparse, short setae dorsally, longer and slightly more abundant setae ventrally; 4 pencil­like white bristles ventrally and 6 thin, black bristles present; FHWR 1.0:5.9­1.0:6.1. Frons dull brown to brown­yellow tomentose. Antenna with pedicel swollen, wider and slightly longer than flagellum. Ocellar tubercle with 2 posterior setae as long as 3 antennal segments combined. Occiput with 4­5 black postocular bristles dorsally, tip of longest bristle about third distance toward ocellar tubercle.

Thorax: Mesonotum subshiny, mostly brown tomentose, grooves, sides, and posterior brown­yellow to yellowish­gray; setae mostly black, sparse, scattered anteriorly; white setae abundant laterally, especially long on postpronotum and between rows of dorsocentral bristles posteriorly; 4­5 lateral and dorsocentral black bristles present. Scutellum brown­yellow tomentose dorsally, gray apically with scattered, long, white setae; 2 black marginal bristles and a shallow preapical groove present. Pleuron black, mostly gray tomentose, with scattered, white setae and bristles; anepisternum yellowish­gray. Halter yellow.

Wing ( Fig. 20 ): Apical and posterior margins narrowly dense microtrichose. Cell r 4 uniformly narrow beyond base to wing margin. Apex of cell m 3 perpendicular to long axis of wing, just beyond crossvein r­m.

Leg: Coxae and trochanters black, former gray tomentose with white setae. Femora mostly yellow; fore and middle femora anteriorly mostly black; hind femur with apical half black, at least dorsally. Fore and middle femora basally with 4­6 thick, yellow bristles, the basal bristle unusually long, about half as long as fore femur ( Fig. 27 ), each bristle thereafter shorter and thinner; middle femur with bristles much thinner, shorter than on fore femur. Hind femur ( Fig. 37 ) narrow basoventrally, widest on basal one­fifth; 1 short setigerous bristle and 6­8 long, thin, yellow anteroventral setae present; 5­7 short, peglike, black bristles basoposteriorly and 2 long, brown, bristles present preapically; HFWLR 1.0:4.4­1.0:5.0. Tibiae yellow with dark apex, fore tibia sometimes entirely yellow; lateral bristles yellow. Hind tibia ( Fig. 47 ) shorter than hind tarsus; apically, a short flange terminates in a raised anterior spur and a small tubercle present, each with a black bristle. Tarsi mostly black except basal tarsomeres mostly yellow; bristles black except 1 fore tarsal bristle yellow; basal tarsomere of middle tarsus laterally with row of short, closely spaced, bristles.

Abdomen: Constricted apically, segments 6­8 about half as wide as segment 2. Ground color black with apical margins of most segments brownish­yellow; tomentum mostly brown dorsally, lighter brownish­gray laterally and ventrally; mostly white setose, tergites black setose medially. Sternite 4 laterally with 6­7 white bristles; sternite 5 with row of very short bristles, each very light in color and with a large socket, the row ends apically with a small oval patch of 7­8 very short black and white bristles; sternite 6 with numerous white bristles, those of sternite 8 apically contrastingly long.

Terminalia ( Figs. 210­214 ): Ventral lamellae basally with ear­like process; process dense setose. Epandrium 2 branched; dorsal branch apically narrow, podiform; ventral branch forked apically. Gonocoxite with internal flange fused with external surface, not projecting posteriorly. Aedeagal sheath projecting above distiphallus as a flat, grooved, oval plate. Hypandrium with extremely narrow, pointed, drooping apex; surface with minute grooves and ridges.

Female. Differs from male as follows. Length, body 6.7­11.5 mm; wing 5.9­7.7 mm. Head: FHWR 5.6­1.0:6.7. Leg: Fore and middle femora basally with only 3­4 long, thin setae. Hind femur anteroventrally with 4­5 bristly setae, longest as great as width of femora medially; HFWLR 1.0:5.9­1.0:6.0. Middle tarsus anteriorly with only 3 short bristles. Abdomen: Bristle pattern of sternites 4­6 absent. Cercus longer than tergite 9. Terminalia ( Figs. 215­217 ): Three spermathecae present; duct basal. Apical margin of sternite 8 with notch admedially.

SPECIMENS EXAMINED. Holotype ♂, allotype ♀, SOUTH AFRICA: N­W Province / Pilanesberg National Park / Bakiburg 25 o 20’40”S 27 o 03’25”E 1170 m Camp / 12­ 19.xi.1999 / JGH Londt ( NMSA) . Paratypes: BOTSWANA: 1 ♂, Serowe / Farmer’s Brigade / x. 1991 SE2226BD / Per Forchammer / Malaise Trap ( CMNH) ; 2 ♂, Serowe # 56 / Farmer's Brigade / 22° 25 S: 26° 14' E 1000 m / 28.xi.1990 / D. Forchhammer / malaise trap ( NMSA) . SOUTH AFRICA: 4 ♂, 3 ♀, same data as holotype ( NMSA) .

Distribution. A species captured in November at 1000 meters in Botswana and South Africa.

Etymology. Latin, strigiatus , for grooves and ridges, referring to the surface of the hypandrium.

Remarks. In addition to the characters in the key, the uniformly narrow cell r 4 ( Fig. 20 ) and short tibia ( Fig. 47 ) further characterize O. strigiatus .

Figures

FIGURES 11­20. Right wing of male Ommatius (Pygommatius) spp. 11. O. imaginus, sp. n..; 12. O. limbus, sp. n..; 13. O. litoreus sp. n.; 14. O. lulua, sp. n.; 15. O. magnipes, sp. n.; 16. O. ornatipes Engel; 17. O. pectinus sp. n.; 18. O. porticus, sp. n.; 19. O. renudus, sp. n.; 20. O. strigiatus, sp. n.

FIGURES 21­28. Right wing of male Ommatius (Pygommatius) spp. 21. O. talus, sp. n.; 22. O. vultus, sp. n.; 23­28. Right fore femur of male Ommatius (Pygommatius) spp. showing arrangement of anteroventral bristles. 23. O. brevicornis Curran; 24. O. grossus, sp. n.; 25. O. limbus, sp. n.; 26. O. litoreus, sp. n.; 27. O. strigiatus, sp. n..; 28. O. vultus, sp. n.

FIGURES 29­38. Male Ommatius (Pygommatius) spp. 29. O. jaculator Walker, middle femur. 30­ 38. Hind femur, 30. O. brevicornis Curran, anterior view; 31. O. comosus, sp. n., anterior view; 32. O. dasypogon Oldroyd, anterior view; 33. O. jaculator Walker, anterior view; 34. O. limbus, sp. n., with coxa, anterior view; 35. O. litoreus, sp. n., anterior view; 36. O. neglectus Bromley, anterior view; 37. O. strigiatus, sp. n., posterior view; 38. O. vultus, sp. n., posterior view.

FIGURES 39­48. Left hind tibia of male Ommatius (Pygommatius) spp. 39. O. comosus, sp. n.; 40. O. fluvius, sp. n.; 41. O. imaginus, sp. n.; 42. O. limbus, sp. n., anterior; 43. O. litoreus, sp. n.; 44. O. neglectus Bromley; 45. O. porticus, sp. n.; 46. O. renudus, sp. n.; 47. O. strigiatus, sp. n.; 48. O. vultus, sp. n., inner surface.

FIGURES 210­217. Male terminalia of Ommatius strigiatus, sp. n. 210­212. Lateral, dorsal and ventral views; 213. Gonostylus; 214. Aedeagus, lateral view. Female terminalia. 215. Spermatheca; 216. Genital fork; 217. Sternite 8, ventral view.