Raveniola, Zonstein, 1987
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publication ID |
https://doi.org/ 10.5852/ejt.2024.967.2699 |
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publication LSID |
lsid:zoobank.org:pub:C08B8027-50CC-417E-BCD4-5183B9FF6738 |
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DOI |
https://doi.org/10.5281/zenodo.14034714 |
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persistent identifier |
https://treatment.plazi.org/id/03A8B723-5510-FF86-FDB4-E2E2FAC5CA08 |
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treatment provided by |
Plazi |
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scientific name |
Raveniola |
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Raveniola View in CoL virgata species group
Diagnostic characters
Less numerous cuspules confined to probasal edge of maxilla ( Figs 220–228 View Figs 220–228 , 247–255 View Figs 247–255 ). PLS small, lackung spigots or entirely absent; apical segment of PLS mostly triangular or digitiform in some species ( Figs 485–486 View Figs 475–486 , 597–618 View Figs 593–601 View Figs 602–610 View Figs 611–618 ). Males: tibiae and metatarsi I–II without modified hairs ( Figs 278–289 View Figs 274–281 View Figs 282–289 , 304– 309 View Figs 290–309 ); cymbium very short (as in Figs 370–378 View Figs 364–378 ); embolus gradually tapering and only slightly curved subapically, always without subapical keel ( Figs 439–465 View Figs 439–447 View Figs 448–456 View Figs 457–465 ). Females: spermathecae always with trunk, narrow Y- or F-shaped ( Figs 534–554 View Figs 522–536 View Figs 537–554 ).
Species included
Raveniola ferghanensis ( Zonstein, 1984) View in CoL , R. kirgizica sp. nov., R. kopetdaghensis ( Fet, 1984) View in CoL , R. mikhailovi Zonstein, 2021 View in CoL , R. nenilini sp. nov., R. ovchinnikovi sp. nov., R. tarabaevi sp. nov., R. virgata ( Simon, 1891) View in CoL and R. vulpina sp. nov.
Key to the species of the virgata View in CoL group
Males
Males of Raveniola tarabaevi sp. nov. are unknown.
1. Small to rudimentary PMS present ( Figs 598, 601–602, 605–606, 609 View Figs 593–601 View Figs 602–610 , 612, 614, 616) .................. 2
– PMS absent ( Fig. 604 View Figs 602–610 ) ......................................................................... R. kopetdaghensis ( Fet, 1984) View in CoL
2. Embolus as long as tegulum or only slightly exceeds the latter in length ( Figs 441–442, 445–465 View Figs 439–447 View Figs 448–456 View Figs 457–465 ) ... 3
– Embolus considerably longer than tegulum ( Figs 439–440 View Figs 439–447 ).......... R. ferghanensis ( Zonstein, 1984) View in CoL
3. Cephalothorax, palps and legs medium to dark reddish brown; dorsal abdomen brownish with darker chevron-like pattern ( Figs 20, 23–25 View Figs 19–27 , 28–35 View Figs 28–35 , 75–81 View Figs 69–75 View Figs 76–81 ); apical segment of PLS shorter (see Figs 597– 598 View Figs 593–601 , 602, 605, 614, 616).................................................................................................................... 4
– Cephalothorax, palps and legs pale yellowish orange; entire abdomen uniformly greyish white ( Figs 26–27 View Figs 19–27 ); apical segment of PLS longer ( Fig. 612 View Figs 611–618 ) ............................... R. ovchinnikovi sp. nov.
4. Embolus almost straight and slightly curved only close to apex ( Figs 448 View Figs 448–456 , 459–461, 465 View Figs 457–465 ) ............. 5
– Embolus very gently twisted ( Figs 445 View Figs 439–447 , 450–452, 454 View Figs 448–456 ).................................................................... 6
5. Palpal tibia 4.0–4.2 times as long as wide; cymbium densely spinose ( Figs 377–378 View Figs 364–378 ). Tegulum noticeably longer than wide ( Figs 459–465 View Figs 457–465 ) ......................................................... R. vulpina sp. nov.
– Palpal tibia 3.4–3.5 times as long as wide; cymbium with 4–5 dorsal spines ( Fig. 374 View Figs 364–378 ). Tegulum almost as long as wide ( Figs 448–449 View Figs 448–456 ).................................................................. R. nenilini sp. nov.
6. Palpal tibia 3.4–3.9 times as long as wide; cymbium with 4–6 dorsal spines ( Figs 373, 376 View Figs 364–378 ). Tibia I 3.9–4.3 times as long as wide; metatarsus I straight or very gently curved ( Figs 282, 286–287 View Figs 282–289 ) .... 7
– Palpal tibia 4.3 times as long as wide; cymbium densely spinose ( Fig. 371 View Figs 364–378 ). Tibia I 4.6 times as long as wide; metatarsus I clearly curved ( Fig. 280 View Figs 274–281 ) ................................................... R. kirgizica sp. nov.
7. Tibia I longer:4.3–4.4 times as long as wide ( Fig.282 View Figs 282–289 ).Palpal tibia longer:3.8–3.9 times as long as wide ( Fig. 373 View Figs 364–378 ). Proximal part of embolus less tapered ( Figs 445–447 View Figs 439–447 ) ..... R. mikhailovi Zonstein, 2021 View in CoL
– Tibia I stouter: 3.9–4.1 times as long as wide ( Figs 286–287 View Figs 282–289 ). Palpal tibia shorter: 3.4–3.6 times as long as wide ( Fig. 376 View Figs 364–378 ). Proximal part of embolus more tapered ( Figs 454–458 View Figs 448–456 View Figs 457–465 ) ............................. .................................................................................................................... R. virgata ( Simon, 1891) View in CoL
Females
Adult females of Raveniola kopetdaghensis View in CoL and R. ovchinnikovi sp. nov. are unknown.
1. Cephalic portion of carapace lesser elevated over thoracic part, fovea narrower ( Figs 129–135 View Figs 127–135 ). Spermathecae with outer and inner branches subequal in length, or outer branch longer than inner one ( Figs 537–554 View Figs 537–554 ) ............................................................................................................................ 2
– Cephalic portion of carapace more elevated over thoracic part, fovea broader( Fig. 128 View Figs 127–135 ). Spermathecae with outer branch much shorter than inner one ( Figs 534–536 View Figs 522–536 ) ......................................................... ........................................................................................................ R. ferghanensis ( Zonstein, 1984) View in CoL
2. Tarsus II with entire or narrowly divided scopula. The distance between paired spermathecae a maximum of four times as long as their length, usually this distance considerably shorter ( Figs 538– 554 View Figs 537–554 ). .................................................................................................................................................. 3
– Tarsus II with widely divided scopula. The distance between paired spermathecae about five times as long as their length ( Fig. 537 View Figs 537–554 ) ......................................................................... R. kirgizica sp. nov.
3. Palpal tarsus with entire or narrowly divided scopula. Outer and inner branches of spermathecae nearly subequal in length; inner branch either evenly narrow along the entire length or dilated subapically ( Figs 538–543, 547–554 View Figs 537–554 )................................................................................................ 4
– Palpal tarsus with widely divided scopula. Outer branch of spermathecae noticeably longer than inner one, or (if not so) the latter subapically flattened, narrowed or constricted ( Figs 544– 546 View Figs 537–554 ) ................................................................................................................... R. tarabaevi sp. nov.
4. Abdomen dorsally either with intermittent dark median stripe ( Figs 56–57, 60 View Figs 54–62 ) or without this contrasting detail ( Figs 61–62 View Figs 54–62 ). Spermathecae shorter, with stouter lateral diverticula ( Figs 538– 540, 547–554 View Figs 537–554 ) .................................................................................................................................. 5
– Abdomen dorsally with almost entire dark median stripe (as in Fig. 58 View Figs 54–62 ). Spermathecae longer, with thinner lateral diverticula ( Figs 541–543 View Figs 537–554 ) ............................................................. R. nenilini sp. nov.
5. Abdomen without contrasting median stripe, but with diffuse or reduced chevrons ( Figs 61–62 View Figs 54–62 ). Lateral diverticulum dilated and flattened subapically ( Figs 551–554 View Figs 537–554 ). PME smaller, apical segment of PLS longer ( Figs 617–618 View Figs 611–618 ) ............................................................................... R. vulpina sp. nov.
– Abdomen with intermittent dark median stripe and chevron-like pattern ( Figs 56, 57, 60 View Figs 54–62 ). Lateral diverticulum differs in shape ( Figs 538–540, 547–550 View Figs 537–554 ). PME larger, apical segment of PLS shorter ( Figs 607–608 View Figs 602–610 , 615)........................................................................................................................... 6
6. PTC I–II and III–IV with 6–7 and 7–9 teeth on each margin, respectively. Spermathecae shorter ( Figs 538–540 View Figs 537–554 )...................................................................................... R. mikhailovi Zonstein, 2021 View in CoL
– PTC I–II and III–IV with 5 and 4–6 teeth on each margin, respectively. Spermathecae longer ( Figs 547–550 View Figs 537–554 )............................................................................................. R. virgata ( Simon, 1891) View in CoL
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