Tomosvaryella pilosiventris ( Becker, 1900 )

Motamedinia, Behnam, Skevington, Jeffrey H. & Kelso, Scott, 2021, Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species, Zootaxa 5002 (1), pp. 1-103: 57-59

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Tomosvaryella pilosiventris ( Becker, 1900 )


Tomosvaryella pilosiventris ( Becker, 1900)  

Figs 27B View FIGURE 27 , 31A–E View FIGURE 31 , 47E, 61

Tomosvaryella glabrum ( Adams, 1905)   .—syn. Kehlmaier et al. (2019): p. 54.

Tomosvaryella tecta De Meyer, 1993   .—syn. De Meyer et al. (2000): p. 148.

Diagnosis: This species can be recognized by the asymmetrical and irregular-shaped surstyli in dorsal view ( Fig. View FIGURE 31

31A); epandrium wider than long in dorsal view ( Fig. 31A View FIGURE 31 ); phallus trifid ( Fig. 31D–E View FIGURE 31 ). The surstyli shape is similar to T. subvirescens ( Loew, 1872)   . It differs from this species by the shape of left surstylus in dorsal view, wrinkled and wavy-shaped ( Fig. 31A View FIGURE 31 ) and triangular-shaped projections on the hind trochanter (Fig. 47E).

Specimens examined: IRAN: Kermanshah: Ghazanchi , 34°26’N, 47°00’E, 1304 m a.s.l., 1.vii.2015, M. Zardouei, Malaise trap, JSS51953 View Materials (1♂, CNC) GoogleMaps   ; North Khorasan: Ghazi , 37°29’N, 56°44’E, 1200 m a.s.l.,, B. Motamedinia, sweep net, JSS51961 View Materials (1♀, CNC) GoogleMaps   ; Sistan & Baluchestan: Iranshahr , 27°24’N, 60°50’E, 769 m a.s.l., 2–12.v.2016, M. Ghaforimoghadam, Malaise trap, JSS52165 View Materials (1♀, CNC) GoogleMaps   ; Zabol , 31°07’N, 61°28’E, 481 m a.s.l.,, H. Derafshan, sweep net, JSS51875 View Materials ‒6 (2♀, CNC) GoogleMaps   ; South Khorasan: Mohammadieh , 32°52’N, 59°01’E, 1419 m a.s.l., 3–17.vii.2016, B. Motamedinia, funnel Malaise trap, JSS51975 View Materials (1♀, CNC) GoogleMaps   ; Salmanefar- si, 32°30’N, 59°13’E,, B. Motamedinia, sweep net, JSS52028 View Materials (1♂, CNC) GoogleMaps   ; ISRAEL: Hof Rotem She- zaf, 32°46’N, 35°38’E, - 200m, 21.iii.2010, A. Freidberg, Malaise trap, JSS52064 View Materials ‒5 (2♂, TAU) GoogleMaps   ; Kinneret, Zee- lon, Beach , 32°50’N, 35°39’E, - 190 m a.s.l.,, A. Freidberg, JSS51737 View Materials ‒9 (3♂, TAU) GoogleMaps   ; Park haYarden, 32°50’N, 35°39’E, - 192 m a.s.l.,, L. Bodner, JSS43963 View Materials (1♀, TAU) GoogleMaps   ; RT. 90, nr. En Admon , 31°01’N, 35°22’E, 11.iv.1994, A. Freidberg & F. Kaplan, JSS51748 View Materials ‒9 (2♂, TAU) GoogleMaps   ; UNITED ARAB EMIRATES: Bithnah , 25°06’N, 56°08’E, 16.xi–26.xii.2006, A. van Harten, Malaise trap, JSS52242 View Materials (1♂, CNC) GoogleMaps   ; 19.x–16.xi.2006, A. van Harten, Malaise trap, CNCD11525 (1♂, CNC)   ; 4.vii–12.viii.2006, A. van Harten, CNCD159940 (1♂, CNC)   ; Wadi Maidaq , 25°11’N, 56°04’E, 26.xii.2006 – 20.ii.2007, A. van Harten, Malaise trap, JSS52252 View Materials (1♂, CNC) GoogleMaps   ;– 29.vii.2006, A. van Harten, Malaise trap, CNCD159304 (1♂, CNC)   ; Wadi Safad , 25°07’N, 56°11’E, 26.xii.2005 – 2.i.2006, A. van Harten, yellow & white pan traps, CNCD8965, CNCD8971 (2♂, CNC) GoogleMaps   ; Wadi Shawkah , 25°04’N, 56°01’E, 18.iii.2007, F. Menzel & A. Stark, sweep net, JSS52246 View Materials (1♂, EAD) GoogleMaps   ; Wadi Shawwkah , 25°04’N, 56°01’E, 31.x–27.xi.2006, A. van Harten, pan trap, CNCD159615 (1♂, CNC) GoogleMaps   ; Wadi Wurayah farm, 25°14’N, 56°11’E, 19.iv–19.v.2009, A. van Harten, Malaise trap, CNCD137547 (1♀, CNC) GoogleMaps   ; near ad-Dhaid, 25°13’N, 55°35’E, 16– 19.iii.2007, J. Btelka, pan trap, CNCD159474 (1♂, USNM) GoogleMaps   .

Distribution: Egypt, Iran, Israel, Kenya, South Africa, Spain (Canary Islands), Syria, Turkey, United Arab Emirates, West Bank, Zimbabwe ( Kehlmaier et al. 2019; Skevington 2020) ( Fig. 61 View FIGURE 61 ).

Notes: DNA barcodes of T. pilosiventris   overlap with T. subvirescens   (0.3–1.9% pairwise divergence). Despite this, the genitalia of these species are rather different. This is thus likely a case of recently diverged species, as seen in many other taxa (e.g. Skevington et al. 2007, Young et al. 2016, Motamedinia et al. 2020). There is always a possibility that it is a single species with polymorphic genitalia, so future genetic work may shed light on this issue.


Canadian National Collection of Insects, Arachnids, and Nematodes


Tel-Aviv University


Smithsonian Institution, National Museum of Natural History














Tomosvaryella pilosiventris ( Becker, 1900 )

Motamedinia, Behnam, Skevington, Jeffrey H. & Kelso, Scott 2021

Tomosvaryella glabrum ( Adams, 1905 )

Kehlmaier, C. & Gibbs, D. J. & Withers, P. 2019: 54

Tomosvaryella tecta

De Meyer, M. & Foldvari, M. & Baez, M. 2000: 148