Tomosvaryella geniculata ( Meigen, 1824 )

Motamedinia, Behnam, Skevington, Jeffrey H. & Kelso, Scott, 2021, Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species, Zootaxa 5002 (1), pp. 1-103: 32-33

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Tomosvaryella geniculata ( Meigen, 1824 )


Tomosvaryella geniculata ( Meigen, 1824)  

Figs 16A‒E View FIGURE 16 , 53 View FIGURE 53

Pipunculus geniculatus Meigen, 1824   .—syn. Földvári & De Meyer (1999): p. 331.

Diagnosis: This species can be recognized by the shape of surstyli in dorsal view, both surstyli slightly rectangular shaped, broader at the base and tips ( Fig. 16A View FIGURE 16 ), tip of both surstyli flattened apically in lateral view ( Fig. 16D–E View FIGURE 16 ); epandrium wider than long ( Fig. 16A View FIGURE 16 ); phallus trifid, one of ejaculatory ducts bearing some small spines in lateral view ( Fig. 16E View FIGURE 16 ); gonopods equal in height ( Fig. 16B View FIGURE 16 ).

Specimens examined: IRAN: Kermanshah: Ghazanchi , 34°26’N, 47°00’E, 1304 m a.s.l.,, M. Zardouei, funnel Malaise trap, JSS51824 View Materials (1♂, CNC) GoogleMaps   ; Harsin , 34°19’N, 47°26’E, 1324 m a.s.l., 22.vii.2016, M. Zardouei, funnel Malaise trap, JSS51944 View Materials (1♂, CNC) GoogleMaps   ; Sarpolezahab , 34°28’N, 45°49’E, 456 m a.s.l., 14.v.2016, M. Zardouei, Malaise trap, JSS52230 View Materials (1♀, CNC) GoogleMaps   ; South Khorasan: Ark , 33°01’N, 58°41’E, 1605 m a.s.l., 30.v–, B. Motamedinia, Malaise trap, JSS51997 View Materials –8 (1♂, 1♀, USNM) GoogleMaps   ; Ghiok , 32°47’N, 59°07’E, 1917 m a.s.l.,, B. Motamedinia, sweep net, JSS52001 View Materials (1♂, CNC) GoogleMaps   ; Jafarieh , 32°48’N, 59°14’E, 1760 m a.s.l., 21.iv–5.v.2016, B. Mo- tamedinia, Malaise trap, JSS52007 View Materials (1♀, CNC) GoogleMaps   ; Mohammadieh , 32°52’N, 59°01’E, 1419 m a.s.l., 14–30.vii.2016, B. Motamedinia, funnel Malaise trap, JSS52051 View Materials (1♂, CNC) GoogleMaps   ; 3–17.vii.2016, B. Motamedinia, Malaise trap; funnel Malaise trap, JSS52041 View Materials –2, JSS52045 View Materials –6, JSS52048, JSS51966 –7, JSS51968 View Materials –72 (11♂, 1♀, CNC)   ; 5–23.ix.2016, B. Motamedinia, Malaise trap, JSS52025 View Materials (1♂, CNC)   ; ISRAEL: Nahal Eshharim , 30°28’N, 34°35’E, 790 m a.s.l., 2.v.1995, I. Yarom, Malaise trap, JSS51791 View Materials (1♂, TAU) GoogleMaps   .

Distribution: Austria, Belgium, Bulgaria, Croatia, Cyprus, Czech Republic, Denmark, Finland, France, Greece, Germany, Great Britain, Hungary, Iran Israel, Italy, Latvia, Macedonia, Mongolia, Morocco, Netherlands, Poland, Portugal, Romania, Russia, Slovakia, Spain, Sweden, Switzerland ( Skevington 2020) ( Fig. 53 View FIGURE 53 ).

Note: This species is genetically most similar to T. emaratensis   sp. nov., differing by 9.7% (pairwise divergence) (Supplementary Table 1).


Canadian National Collection of Insects, Arachnids, and Nematodes


Smithsonian Institution, National Museum of Natural History


Tel-Aviv University














Tomosvaryella geniculata ( Meigen, 1824 )

Motamedinia, Behnam, Skevington, Jeffrey H. & Kelso, Scott 2021

Pipunculus geniculatus

Foldvari, M. & De Meyer, M. 1999: 331