Harpactea bilecenoglui, Kunt & Özkütük, 2023

Kunt, Kadir Boğaç & Özkütük, Recep Sulhi, 2023, New data on the Harpacteinae of Turkey (Araneae, Dysderidae), Zootaxa 5375 (3), pp. 379-408 : 389-393

Harpactea bilecenoglui sp. nov.

Figs 24–36

Material examined: Holotype 1 ♂ ( ETZM), Alanya Dist. , Alanya Castle (36°32’18.38”N 31°59’39.86”E), asl c. 125 m, 04 Dec 2011, leg. Kadir Boğac Kunt. Paratypes 5 ♂, 16 ♀, 12J ( ETZM), same data as holotype — 8 ♀, 8 ♂ ( ETZM), Alanya Dist. , TaŞatan Plateau (36°38’26.79”N 32°4’2.79”E), asl c. 1170 m, 07 Nov 2012, leg. Kadir Boğac Kunt — 13 ♂, 13 ♀, 3J ( ETZM), Alanya Dist. , Alanya Castle (36°32’18.38”N 31°59’39.86”E), asl c. 125 m, 07 Jan 2013, leg. Kadir Boğac Kunt — 1 ♀ ( ETZM), Alanya Dist. , TaŞatan Plateau, Banlıca Area (36°40’10.98”N 32° 9’15.06”E), asl c. 1017 m, 1 Apr 2016, leg. Kadir Boğac Kunt.

Etymology: The specific epithet is a patronym honoring an old friend of the first author, Turkish ichthyologist, Murat Bilecenoğlu.

Diagnosis: Males of H. bilecenoglui sp. nov. resemble those of H. elvericii sp. nov., H. isaurica Brignoli, 1978 and H. sanctaeinsulae Brignoli, 1978 by the spherical, bulging bulb lacking conductor and spiniform embolus ( Figs 27–30 , 40–41 , 44–45 ). Bulbs of H. bilecenoglui sp. nov. and H. sanctaeinsulae are distinguished by the widened first half of the spiniform embolus of H. bilecenoglui sp. nov. compared with H. sanctaeinsulae .

Female of H. bilecenoglui sp. nov. resemble those of H. elvericii sp. nov., H. isaurica and H. sanctaeinsulae by the short spermatheca, with basal half enlarged, oval-shaped, spermathecal keel expands frontaly into a square like structure ( Fig. 33 ).

Description: Measurements: [Holotype ♂ / Paratypes ♀] TL 3.50/4.05–4.65; AL 1.80/2.05–2.40; CL 1.70/2.00–2.25; CW 1.25/1.60–1.70; Clh 0.04/0.03–0.04; AEd 0.10/0.11–0.12; iAE 0.03/0.02–0.03; PLEd 0.08/0.12–0.12; PMEd 0.05/0.08–0.09; ChF 0.23/0.33–0.40; ChG 0.12/0.25–0.30; ChL 0.60/0.90–1.00.

[Paratype ♂] TL 4.25; AL 2.25; CL 2.00; CW 1.60; Clh 0.03; AEd 0.11; iAE 0.02; PLEd 0.12; PMEd 0.09; ChF 0.29; ChG 0.23; ChL 0.75.

Medium sized harpacteine spiders. Carapace brown in both sexes. No obvious tonal difference between head and thoracic regions.

Margins of the carapace are comparatively darker than in general. Fovea longitudinal, dark brown and prominent ( Figs 24–25 ). AE, PLE and PME arranged annularly, close to each other. AEs not in contact with each other. Sternum, labium, gnatocoxae light brown. Sternum lighter than carapace. Tips of the labium and gnatocoxae with brown setae. Chelicerae brown. Surface of the chelicerae medially covered with protuberances bearing short blackish setae. Cheliceral groove with four teeth, two on promarginal and two on retromarginal. The teeth on the promarginal are stronger than those on the retromarginal. The promarginal teeth at the base of the cheliceral groove are relatively larger than the one following it. There is a very small amount of space between the two. The position of the first retromarginal tooth is exactly in the centre of the promarginal teeth. Both retromarginal teeth are of the same size, with a gap between them almost five times their basal length ( Fig. 26 ). Abdomen cylindrical; yellowishcream, covered with brownish setae. Legs reddish brown. Leg formula: VI, I, II, III. Leg spination and measurements given in Tables 7 and 8.

Palp ( Figs 27–30 ): Palpal tarsus longer than tibia. Tegulum swollen, asymmetrical, oval or spherical, depending on perspective. Coloration of bulb lighter than palpal segments. Embolus black, massive, spear-shaped; connected with tegulum posterior-ventrally. Conductor and median apophysis absent.

Vulva ( Figs 31–36 ): Anterior arc of vulva strongly sclerotized. The shape of the spermathecal keel varies between individuals ( Figs 31 – 36 ). However, it is generally quadrangular ( Figs 33–34 ). The centre of the spermatheca bulging to the sides ( Fig. 32 ). Variation between individuals, but usually ovalish or hexagonal. Anterior margin of the anterior arc short, circular, ring-shaped ( Fig. 33 ). The roundishstructure and the transversal bar appear to be interconnected. The posterior diverticulum is usually membranous and distinct, although its width may vary, it is typically equal to its length.. ( Figs 31, 33 – 34 ).

Figures

FIGURES 24–26. Harpactea bilecenoglui sp. nov. 24. Habitus, male 25. Ditto, female 26. Chelicerae, ventral view (red dots, promarginal; white dots, retromarginal teeth).

FIGURES 27–30. Harpactea bilecenoglui sp. nov. 27. Bulb (right), retrolateral view 28–30. Bulb (left), prolateral view. Abbreviations: E, Embolus T, Tegulum.

FIGURES 31–36. Harpactea bilecenoglui sp. nov. 31–35. Vulva, dorsal view. 36. Ditto, ventral view, after maceration. Abbreviations: AA, anterior margin of the anterior arch; AC, anterior arc; PD, posterior diverticulum; RS, roundishstructure; S, spermatheca; SK, spermathecal keel; TB, transversal bar.

FIGURES 40–43. Harpactea elvericii sp. nov. 40. Bulb (left), prolateral view 41. Bulb (right), retrolateral view 42. Vulva, dorsal view 43. Ditto, nearly dorsal view. Abbrevations: AA, anterior margin of the anterior arch; AC, anterior arc; E, embolus; PD, posterior diverticulum; S, spermatheca; SK, spermathecal keel; T, tegulum; TB, transversal bar.

FIGURES 44–45. Harpactea sanctaeinsulae, holotype 39. Bulb, prolateral view 40. Ditto, retrolateral view.