Cinetomorpha Simon, 1892

Ott, Ricardo, Ubick, Darrell, Bonaldo, Alexandre B., Brescovit, Antonio D. & Harvey, Mark S., 2019, A revision of the new world goblin spider genus Cinetomorpha Simon, 1892 revalidated from Gamasomorpha Karsch, 1881 (Araneae, Oonopidae, Oonopinae), Zootaxa 4641 (1), pp. 1-152: 8-10

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Cinetomorpha Simon, 1892


Cinetomorpha Simon, 1892   , revalidated

Cinetomorpha Simon, 1892: 558   (type species by monotypy Cinetomorpha simplex, Simon, 1892   ); Simon 1893b: 442. Yumates Chamberlin, 1924: 595   (type species by original designation Yumates nesophila Chamberlin, 1924   ). New synonymy. Lucetia Dumitresco & Georgesco, 1983: 109   (type species by monotypy Lucetia distincta Dumitresco & Georgesco, 1983   ).

New synonymy.

Diagnosis. Species of Cinetomorpha   resemble those of Grymeus   and Sicariomorpha   by the presence of a relative large, female internal genitalia at the level of the epigastric scutum which is not partially or totally fused with the ventral scutum ( Figs 1D, E View FIGURES 1 ), with rounded, small receptaculum ( Figs 1D, E View FIGURES 1 ; 2E, G View FIGURES 2 ; 4B View FIGURES 4 ; 34H; 81H; 112H; See also Ott et al., 2015, figs. 15̅16), but can be distinguished from these genera by the much more elaborate distal portion of the male embolic complex, with the embolus bearing a sclerotized distal process and longer embolar prolateral process ( Figs 2C View FIGURES 2 , 33A View FIGURES 33 ̅B, 70L, 108D̅G). Cinetomorpha   species also differ by the presence of a slit-like ectal extensions of posterior spiracles in both sexes ( Fig. 2A View FIGURES 2 ), similar to the ones found in species of Dysderina   complex (see Abrahim et al., 2012, figs 93, 127), but whose species differ in having leg spines.

Description: CEPHALOTHORAX. Carapace without any pattern, anteriorly narrowed to 0.49 times its maximum width or less, with rounded posterolateral corners, posterolateral edge without pits, posterior margin not bulging below posterior rim, anterolateral corners without extension or projections, posterolateral surface without spikes, thorax without depressions, fovea absent, without radiating rows of pits; lateral margin undulate, rebordered; plumose setae near posterior margin of pars thoracica absent; non-marginal pars cephalica setae light, needle-like, present in u-shaped row; non-marginal pars thoracica setae absent; marginal setae light, needle-like ( Figs 3C View FIGURES 3 , 70A View FIGURES 70 , 86C View FIGURES 86 , 105E View FIGURES 105 ). Clypeus margin unmodified, vertical in lateral view, median projection absent; setae present, light, needle-like ( Figs 70B View FIGURES 70 , 73E View FIGURES 73 ). Chilum absent. Eyes six, well developed, ALE circular, PME oval, PLE circular; posterior eye row recurved from above; ALE-PLE separated by less than ALE radius, PME touching ( Figs 6C View FIGURES 6 , 68E View FIGURES 68 ). Sternum as long as wide, uniform, anterior margin continuous groove (anterior pouch), generally conspicuous in males ( Figs 5G View FIGURES 5 , 6B View FIGURES 6 , 52B View FIGURES 52 , 108B View FIGURES 108 ), absent in females, median concavity absent, with radial furrows between coxae I- II, II-III, III-IV, radial furrow opposite coxae III absent, without pits, sickle-shaped structures absent, anterior corner unmodified, lateral margin with infra-coxal grooves and anterior and posterior openings, distance between coxae approximately equal, extensions of pre-coxal triangles absent, lateral margins unmodified, without posterior hump; setae sparse, light, needle-like, evenly scattered, without hair tufts. Mouthparts: Chelicerae straight; promargin and retromargin generally without teeth (in males of C. chicote   , one retromarginal tooth, Fig. 18G View FIGURES 18 ); fangs directed medially, shape normal, without prominent basal process, tip unmodified; setae light, needle-like, densest medially; paturon distal region unmodified, posterior surface unmodified, promargin unmodified, inner margin unmodified; in some species of the groups simplex   and puberula   , male paturon with enlarged mesal setae ( Figs 6A View FIGURES 6 ̅D). Labium rectangular, not fused to sternum, same as sternum in sclerotization; with 3-5 setae on anterior margin, subdistal portion with unmodified setae. Male endites slightly excavated distally, with shallow distal groove female endites unmodified; serrula present in single row ( Figs 108B View FIGURES 108 ̅C), anteromedian tip unmodified, posteromedian part unmod- ified, same as sternum in sclerotization. Abdomen: ovoid, without long posterior extension, inter-scutal membrane rows of small sclerotized platelets absent; dorsum soft portions without color pattern, dorsal scutum covering full length of abdomen, no soft tissue visible from above, not fused to epigastric scutum, anterior half without projecting denticles ( Figs 3B, D View FIGURES 3 ; 4A View FIGURES 4 ; 34A, E). Epigastric scutum strongly sclerotized, surrounding pedicel, slightly protruding, small lateral sclerites absent. Male anterior spiracles forming an additional mesal secondary opening, usually on an indent under a fold at the posterior border of the epigastric scutum ( Figs 5F View FIGURES 5 , 37H View FIGURES 37 ), rarely as almost independent openings in the surface of the epigastric plate ( Figs 2A View FIGURES 2 , 31G View FIGURES 31 ̅H, 33D). Female epigastric scutum without lateral joints. Post-epigastric scutum strongly sclerotized, covering nearly full length of abdomen, fused to epigastric scutum in males, not fused to epigastric scutum and without posteriorly directed lateral apodemes in females, anterior margin unmodified. Spinneret scutum present, forming incomplete ring. Supra-anal scutum absent. Dorsum with setae present, light, needle-like. Epigastric area setae uniform, light, needle-like. Post-epigastric area setae present, light, needle-like. Spinneret scutum with fringe of needle-like setae. Dense patch of setae anterior to spinnerets absent. Interscutal membrane with setae. Colulus represented only by setae. Legs: without color pattern; femur IV not thickened, same size as femora I-III, patella plus tibia I shorter than carapace, tibia I unmodified, tibia IV specialized hairs on ventral apex present, tibia IV ventral scopula absent, metatarsi I and II meso-apical comb absent, metatarsi III and IV weak ventral scopula absent. Leg spines absent. Female palp without claws, spines absent; tarsus unmodified, patella without prolateral row of ridges ( Fig. 4G View FIGURES 4 ). Genitalia: male epigastric region with sperm pore small; furrow without omega-shaped insertions, without setae. Male palp normal size, not strongly sclerotized, right and left palps symmetrical, prolateral excavation absent; trochanter normal size, unmodified; femur normal size, two or more times as long as trochanter, without posteriorly rounded lateral dilation, attaching basally to patella; patella shorter than femur, not enlarged, without prolateral row of ridges, setae unmodified ( Figs 6E View FIGURES 6 ; 39A, B View FIGURES 39 ); cymbium yellow, ovoid in dorsal view, fused with bulb at base, plumose setae absent, without stout setae, without distal patch of setae; bulb yellow, 1 to 1.5 times as long as cymbium, stout and spherical. Embolic complex distally darker; prolaterally curved, composed of three contiguous parts: a two-branched embolus and a basally inserted conductor ( Figs 2C View FIGURES 2 , 37 View FIGURES 37 A-D, 70I-L). Conductor of variable shape, usually entire ( Figs 2C View FIGURES 2 , 70L View FIGURES 70 , 32G View FIGURES 32 , 33A View FIGURES 33 ), rarely subdivided distally ( C. chicote   sp. nov.; Figs 18H, I View FIGURES 18 ). Embolus with dorsal process bearing sub-distal ejaculatory opening, distally attenuated, and prolateral process which may be slightly longer ( Fig. 2C View FIGURES 2 ) to much longer ( C. chicote   sp. nov., Fig. 18H View FIGURES 18 ) than dorsal process, usually with a translucent dorsodistal region ( Figs 6H View FIGURES 6 , 43H View FIGURES 43 ). Female epigynum with internal genitalia positioned anteriorly to the level of the epigastric scutum and not fused to any scutum, variable in shape, lateral apodemes present, usually well developed; receptaculum much narrower than genitalia width, round to oval in shaped; anterior receptacle inconspicuous; paddle-like sclerite conspicuous ( Figs 2E, G View FIGURES 2 ) or not ( Figs 105H View FIGURES 105 , 109H View FIGURES 109 ); globular apendix positioned anteriorly to internal genitalia, usually short and twisted, assuming an asymmetrical form ( Figs 2E, G View FIGURES 2 ).

Composition. The genus Cinetomorpha   contains 41 species, of which 31 are newly described.

Distribution. Species of Cinetomorpha   are currently known only from the Americas.








Arachnida (awaiting allocation)


Arachnida (awaiting allocation)


Cinetomorpha Simon, 1892

Ott, Ricardo, Ubick, Darrell, Bonaldo, Alexandre B., Brescovit, Antonio D. & Harvey, Mark S. 2019


Dumitresco, M. & Georgesco, M. 1983: 109
Chamberlin, R. V. 1924: 595
Simon, E. 1893: 442
Simon, E. 1892: 558