Darevskia, Arribas, 1999

Arribas, Oscar, 2024, The osteology of the lacertid genus Darevskia Arribas, 1999 (Squamata, Lacertidae), Zootaxa 5510 (1), pp. 1-62 : 52-54

Darevskia hybrids

HYBRIDS between Darevskia (double entry by each parental species in alphabetic order).

(p)=parthenogenetic. (The result of the hybridization between a parthenogenetic female and a bisexual male is a triploid hybrid).

Hybridizations between bisexual species that gave rise to parthenogenetic taxa in the past have not been considered in the list (see main text to see these presumably parental species).

Interspecific hybrids (other than parthenogenetic origin ones) known:

* D. adjarica x D. derjugini (Darevsky 1967; Bischoff 1973; Orlova 1979; Murphy et al. 2000; Ciobanu et al. 2003) (1).

* D. alpina x D. brauneri (Darevsky 1967; Murphy et al 2000).

* D. alpina x D. caucasica (Darevsky 1967; Fu et al 1995; Murphy et al. 2000).

* D. alpina x D. mixta (Fu et al. 1997; Murphy et al. 2000).

* D. armeniaca (p) x D. nairensis (Bischoff 1973) .

* D. armeniaca (p) x D. portschinskii (Darevsky & Kulikowa 1961; Mertens 1964; Bischoff 1973).

* D. armeniaca (p) x D raddei (Darevsky & Kulikowa 1961 Mertens 1964) .

* D. armeniaca (p) x D. obscura (Darevsky 1967; Bischoff 1973; Darevsky et al. 1985).

* D. armeniaca (p) x D. o. macromaculata (Darevsky 1967; Bischoff 1973).

* D. armeniaca (p) x D. valentini (Darevsky 1966, 1967; Darevsky & Kulikowa 1961, 1962; Darevsky et al. 1973; Mertens 1964; Bischoff 1973; Darevsky et al. 1985; Freitas 2017).

* D. brauneri x D. alpina (Darevsky 1967; Murphy et al. 2000).

* D. brauneri x D. saxicola (Darevsky 1967; Murphy et al. 2000).

* D. brauneri brauneri x D. b. darevskii (2) (Darevsky 1967).

* D. b. darevskii x D. brauneri brauneri (2) (Darevsky 1967).

* D. caucasica x D. alpina (Darevsky 1967; Fu et al. 1995; Murphy et al. 2000).

* D. caucasica x D. daghestanica (Darevsky 1967; Fu et al. 1995; Murphy et al. 2000).

* D. caucasica x D. derjugini (Orlova 1979; Ciobanu et al. 2003).

* D. caucasica x D. saxicola (Darevsky 1967; Murphy et al. 2000).

* D. clarkorum x D. rudis (Darevsky & Vedmederja 1977; Ciobanu et al. 2003).

* D. daghestanica x D. caucasica (Darevsky 1967; Fu et al. 1995; Murphy et al. 2000).

* D. dahli x D. portschinskii (Darevsky & Kulikowa 1961; Mertens 1964; Darevsky 1967; Bischoff 1973).

* D. derjugini x D. adjarica (Darevsky 1967; Bischoff 1973; Orlova 1979; Murphy et al. 2000; Ciobanu et al. 2003)(1).

* D. derjugini x D. caucasica (Orlova 1979; Ciobanu et al. 2003).

* D. derjugini x D. mixta (Orlova 1979; Murphy et al. 2000; Ciobanu et al. 2003).

* D. derjugini x D. o. obscura (Darevsky & Kulikowa 1961; Mertens 1964; Bischoff 1973).

* D. derjugini x D. saxicola (Orlova 1979; Ciobanu et al. 2003).

* D. dryada x D. rudis (Darevsky & Tuniyev 1997; Ciobanu et al. 2003).

* D. mixta x D. alpina (Fu et al. 1997; Murphy et al. 2000).

* D. mixta x D. derjugini (Orlova 1979; Murphy et al. 2000; Ciobanu et al. 2003).

* D. nairensis x D. armeniaca (p) (Bischoff 1973).

* D. nairensis x D. unisexualis (p) (Bischoff 1973; Darevsky et al. 1985).

* D. parvula x D. rudis (Darevsky 1967; Murphy et al. 2000).

* D. portschinskii x D. armeniaca (p) (Darevsky & Kulikowa 1961; Mertens 1964; Bischoff 1973).

* D. portschinskii X D. dahli (Darevsky & Kulikowa 1961; Mertens 1964; Darevsky 1967; Bischoff 1973).

* D. portschinskii x D. raddei (Darevsky 1967; Murphy et al. 2000; Freitas 2017).

* D. portschinskii x D. rostombekowi (p) (Freitas 2017).

* D. raddei x D. armeniaca (p) (Darevsky & Kulikowa 1961; Mertens 1964).

* D. raddei x D. portschinskii (Darevsky 1967; Murphy et al. 2000; Freitas 2017).

* D. raddei x D. rostombekowi (p) (Darevsky et al. 1973, 1985, 1986).

* D. raddei x D. valentini (Darevsky & Kulikowa 1961) .

* D. rostombekowi (p) x D. portschinskii (Freitas 2017) .

* D. rostombekowi (p) x D. raddei (Darevsky et al. 1973, 1985, 1986).

* D. rudis x D. clarkorum (Darevsky & Vedmederja 1977; Murphy et al. 2000; Ciobanu et al. 2003).

* D. rudis x D. dryada (Darevsky & Tuniyev 1997; Ciobanu et al. 2003).

* D. rudis x D. parvula (Darevsky 1967; Murphy et al. 2000).

* D. rudis rudis x D. r.lantzicyreni (Darevsky 1967) .

* D. o. macromaculata x D. armeniaca (p) (Darevsky 1967; Bischoff 1973).

* D. o.macromaculata x D. o. obscura (Darevsky 1967) (3).

* D. o. obscura x D. armeniaca (p) (Darevsky 1967; Bischoff 1973; Darevsky et al. 1985).

* D. o. obscura x D. derjugini (Darevsky & Kulikowa 1961; Mertens 1964; Bischoff 1973).

* D. o. obscura x D. o. macromaculata (Darevsky 1967) (3).

* D. saxicola x D. brauneri (Darevsky 1967; Murphy et al. 2000).

* D. saxicola x D. caucasica (Darevsky 1967; Murphy et al. 2000).

* D. saxicola x D. derjugini (Orlova 1979; Ciobanu et al. 2003).

* D. unisexualis (p) x D. nairensis (Bischoff 1973; Darevsky et al. 1985).

* D. unisexualis (p) x D. valentini (Darevsky 1967; Bischoff 1973; Freitas 2017).

* D. valentini x D. armeniaca (p) (Darevsky 1966, 1967; Darevsky & Kulikowa 1961, 1962; Darevsky et al. 1973; Mertens 1964; Bischoff 1973; Darevsky et al. 1985; Freitas 2017).

* D. valentini x D. raddei (Darevsky & Kulikowa 1961) .

* D. valentini x D. unisexualis (p) (Darevsky 1967; Bischoff 1973; Freitas 2017).

* D. r. lantzicyreni x D. rudis rudis (Darevsky 1967) .

(1) By the localities concerned, the “ D. parvula ” of these hybridizations were in fact Georgian D. adjarica .

(2) Considered subspecies, or synonyms by Doronin et al. (2013)

(3) D. obscura is considered a species by Gabelaia et al. (2018) and Arribas et al. (2022). The strange and unexpected differences found by Mayer & Lutz (1989) could support this species status. However, even in this case, it remains to be resolved the complex internal relationships of the Great Caucasus forms (svanetica and chechenica) if they belong to rudis s. str, are conspecific with obscura , or should be considered independent with species status (see Arribas et al. 2013).