Indonemoura clavata, Sivec & Stark, 2010

Sivec, Ignac & Stark, Bill P., 2010, Eleven New Species Of The Genus Indonemoura Baumann (Plecoptera: Nemouridae) From Thailand And Vietnam, Illiesia 6 (14), pp. 210-226 : 217-218

Indonemoura clavata sp. n.

( Figs. 20-24 )

Material examined. Holotype ♂ from Vietnam, Vinh Phu Province , Tam Dao, 800-1000 m, 19 May-13 June 1995, H. Malicky ( PMSL) . Additional material: Vietnam: Type locality, 19 May-13 June 1995, H. Malicky, 15♀ ( PMSL). Vinh Phu Province , Tam Dao, 23 March 1995, 1♀ ( ZMB) .

Adult habitus. General color dark brown. Head and wings dark brown; pronotum slightly paler with obscure rugosities. Legs, palpi and antennae uniformly dark brown.

Male. Forewing length 6.5 mm. Epiproct narrowed near midlength and slightly expanded subapically, apex with a shallow notch in dorsal aspect ( Fig. 20 ); ventrolateral aspect greatly expanded and armed along most of length with thick setal spines ( Fig. 21 ). Mesal lobe of paraprocts sharply bent near base and club shaped; outer lobe slender, heavily sclerotized, appressed to mesal lobe near apex, bearing a subapical knob and a bifurcate tip; inner lobe needlelike ( Figs. 22-23 ). Tergum 10 bearing a small median patch of sensilla; tergum 9 with a sparse patch of sensilla along posterior margin. Vesicle narrow, only slightly wider subapically ( Fig. 22 ); hypoproct slightly constricted near midlength between apex and posterior margin of sternum 9.

Putative female. Forewing length 8 mm. Subgenital plate scarcely produced beyond hind margin of sternum 8; posterior margin with a slight median emargination ( Fig. 24 ). Oval paragenital plates distinct.

Larva. Unknown.

Etymology. The species name refers to the club shaped mesal paraproct lobe.

Diagnosis. The epiproct of this species is similar to that of several species including I. auriformis Li & Yang ( Li & Yang 2008a), I. baishanzuensis Li & Yang , I. fujianensis Li & Yang , I. guangdongensis Li & Yang , I. hubeiensis Yang &Yang , I. macrolamellata (Wu) and I. yangi Li & Yang ( Li & Yang 2006; 2008c) and I. tricantha (described below). We formally recognize these species, distinguished on the basis of paraproct structures, particularly those of the outer lobes, as members of the Fujianensis complex. Within the complex only I. clavata has the mesal paraproct lobe club shaped and the outer lobe terminating in a pair of short, almost equally sized spines. Chinese members of the complex, except I. auriformis , are included in keys developed by Li & Yang (2006; 2008c).This new species and I. tricantha , the only regional members of the complex, are distinct from other Southeast Asian Indonemoura in having the ventral epiproct sclerite greatly produced into a wide, semicircular structure ( Fig. 21 ), typical of all species in the I. fujianensis complex.

Figures

Figs. 20-24. Indonemoura clavata. 20. Male terminalia, dorsal aspect. 21. Male terminalia, lateral aspect. 22. Male terminalia, ventral aspect. 23. Male paraprocts, caudal aspect. 24. Putative female terminalia, ventral aspect.