Eperua oleifera Ducke var. campestris Ducke (1932: 728)

Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V. & Mansano, Vidal F., 2023, A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae), Phytotaxa 617 (1), pp. 1-127 : 76-80

14b. Eperua oleifera Ducke var. campestris Ducke (1932: 728)

( Figures 14 , 43 , 45 )

Type:— BRAZIL: Amazonas : Borba , Rio Madeira, campo arenoso, 17 January 1930, Ducke W. A. s.n. RB23291 (holotype RB sheet I [00539605]!, sheet II[00585835]!, & sheet III[00585836]!; isotypes NY [00004384] image!, P [00835938] image!, R [000028783]!, U [0003232] image!, US [00001150]!).

Treelet, or shrub 1.5–9.0 m tall. Blades: lower pair base rounded, subcordate to cordate, middle pair base rounded, subcordate to cordate, upper pair base obtuse, rounded, subcordate to cordate, tertiary veins very conspicuous, prominent in the adaxial surface, rarely not prominent.

Phenology:— Flowering mainly in April, from June to September, and in November, with a flowering peak in August; fruiting from May to August, in October, and November.

Distribution:— Southeast of Amazonas state in the Madeira River basin ( Brazil).

Habitat:— White sand campina and campinarana, in upland and fooldplains areas, at ca. 100 m elev.

Occurrence in protected areas:— Unknown.

Etymology:— The specific epithet relates to its habitat in the campinas.

Vernacular names:— copaiba jacaré (Pessoal do Proj. Radam 13234).

Uses:— Unknown.

Taxonomic notes:— Eperua oleifera var. campestris differs from the type variety by being a treelet or shrub (vs. tall tree in var. oleifera ) with the lower and middle pair of leaflets having a rounded, cordate to subcordate base (vs. rounded to obtuse, rarely cordate or subcordate, in var. oleifera ), and very conspicuous tertiary veins (vs. conspicuous to inconspicuous in var. oleifera ). Eperua oleifera var. campestris presents strigose to tomentose inflorescence (vs. tomentose, indumentum tending to be denser, in var. oleifera ), and it is the only reproductive difference, justifying its permanency as a variety.

Selected specimens:— BRAZIL. Amazonas, Apuí, rodovia Transamazônica, igarapé Coca-cola, 7°26’5”S 61°2’23”W, 18 August 2009, Assunção P.A.C.L. & Carvalho F.A. de 1118 ( INPA ). Borba, rio Madeira, 09 November 1935, Ducke W.A. 13 (K). Manaus, rio Negro basin September 1948, Schultes R.E. & López F. 10337 (IAN, NY, US); 5 Km S of Borba, 4°23’0”S 59°35’0”W, 26 June 1983, Zarucchi J.L. et al. 2842 (HFSL, INPA , K, MG, MO, NY, RB, UFACPZ, US); Axinim, basin of rio Abacaxis, ca. 4km from mouth of igarapé Surubim, 4°31’0”S 58°34’0”W, 04 July 1983, Zarucchi J.L. et al. 2941 ( INPA , K, MG, MO, NY, RB, UFACPZ, US). Manicoré, BR 230, rod. Transamazônica, a 243 Km de Humaitá, 7°40’0.1”S 61°10’0”W, 24 April 1985, Cid-Ferreira C.A. 5791 (F, HRCB, INPA , K, MG, MO, RB, US). Maués, campina Socoró, 4°17’2”S 58°34’40”W, 07 August 2015, Demarchi L.O. et al. 529 ( INPA ). Novo Aripuan„, BR 230, rod. Transamazônica entre os Kms 350 e 400 ao leste de Humaitá, 30 April 1985, Cid-Ferreira C.A. 5913 (F, HRCB, INPA , K, MG, MO, NY, RB, US); Nova Prainha, 23 July 1976, Mota C.D.A. da & Monteiro O.P. s.n. INPA 60692 ( INPA ). Nova Olinda do Nortes, rio Abacaxis, Igarapé do Socoró, campina Socoró, 4°17’2”S 58°34’40”W, 07 August 2015, Holanda A.S.S. et al. 799 ( INPA ). Rio de Janeiro (cultivated), Rio de Janeiro, Arboreto do Jardim Botânico do Rio de Janeiro, nº 317, canteiro A, séc. XXV, 25 April 1985, Carvalho I.R. & Ricardo C.C. s.n. RB243379 ( INPA, RB ).

Figures

FIGURE 14. Geographic distribution of Eperua purpurea, E. oleifera var. campestris, and E. oleifera var. oleifera.

FIGURE 43. Eperua oleifera var. campestris: A. habit; B. bifurcate trunk; C. leaf, adaxial surface, and detail of the leaflet base; D. leaf, abaxial surface; E. flowers; F. fruits. Photos, F. Farroñay.

FIGURE 45. Eperua oleifera. A. fruiting branchlet, var. oleifera.; B. flowering branchlet, var. campestris; C adaxial petal, var. campestris; D. base of the stamens joined in a diadelphous sheath, var. campestris; E. anther and filament, var. campestris; F. carpel, var. campestris; G. stigma, var. campestris. Specimens, A: Zarucchi J.L. et al. 2935; B: Zarucchi J.L. et al. 2842; C–E: Schultes R.E. & López F. 10337. Illustrated by Carlos Alvarez.