Eperua glabriflora ( Ducke 1940: 27 ) R.S. Cowan (1975: 31)

Fortes, Elenice A., Dos, Ilka P., Steege, Hans Ter, Aymard, Gerardo, Secco, Ricardo De S., Martins-Da-Silva, Regina Célia V. & Mansano, Vidal F., 2023, A taxonomic revision of the genus Eperua (Leguminosae, Detarioideae, Detarieae), Phytotaxa 617 (1), pp. 1-127 : 50-51

8. Eperua glabriflora ( Ducke 1940: 27) R.S. Cowan (1975: 31)

≡ Eperua bijuga Martius ex Bentham (1870: 226) var. glabriflora Ducke (1940: 27) .

Type (designated by Cowan 1975): — BRAZIL. Amazonas : Estrada do Aleixo, 3 April 1936, Ducke W. A. 158 (lectotype US[00001143]!, isolectotypes A not seen, F [0057801] image!, K [000555105] image!, MO [954335] image!, NY [00004376] image!, R sheet I [000054528]!, & sheet II[ R000054528 a]!) .

Tree 3.0– 37.5 m tall. Trunk 10.0– 54.3 cm in diameter, bark dark-yellow-brown, yellowish, brownish, lenticellate to slightly scaled. Stipules 1.3–3.7 × 0.7–2.7 mm, free, non-foliaceous, caducous to persistent. Leaves 2(3)-jugate; petioles 0.9–6.4 cm long; rachis 1.7–15.5 cm long; petiolules 4.0– 13.8 mm long; blades 6.2–23.0 × 2.7–17.0 cm, coriaceous, glabrous, epunctate, rarely pellucid-punctate, not discolorous, rarely discolorous, equilateral, lower pair elliptic, sometimes ovate, rarely broadly ovate, smaller, middle pair elliptic, upper pair elliptic, rarely broadly elliptic, obovate or ovate, apex acuminate, mucronate, base symmetrical, lower pair base obtuse to rounded, rarely subcordate to cordate, middle pair base obtuse to subcordate, upper pair base obtuse, sometimes rounded, rarely subcordate or cuneate, margin revolute, sometimes flat to slightly revolute at the base, secondary venation with two intramarginal veins, vein closer to the margin continuous, main vein straight, depressed on the adaxial surface, tertiary veins conspicuous, areoles smaller, concave. Inflorescences terminal or axillary, sometimes on ramiflorous, panicle, rarely raceme, erect, glabrous, 3.0– 14.9 cm long, lateral racemes 1.0–6.4(–10.6) cm long, alternate and distichous, sometimes spirally arranged; bracts 4.7–5.3 × 5.6–7.2 mm, broadly ovate, concave, apex gland present, glabrous, caducous; bracteoles 5.3–10.3 × 3.1–6.5 mm, ovate, rarely broadly ovate, cucullate, apex gland present, glabrous, caducous, attached to the lower middle portion of the pedicels; pedicel (2.0–)3.0– 9.3 mm long, 0.9–3.4 mm in diameter, not twisted, glabrous; buds 0.7–1.7 cm long, 0.4–0.9 cm in diameter, glabrous. Flowers: hypanthium (1.2–)2.0– 4.2 mm long, (1.2–)2.1–7.0 mm in diameter, cup-shaped, inequilateral, glabrous; sepal 0.8–2.0 × 0.3–1.1 cm, elliptic to oblong, unequal, the outer sepals more robust, the dorsal one larger, cucullate, apex gland present on the outer ones, purple, sometimes pale-purple to pink-reddish, or greenish, glabrous, pellucid-punctate, inner sepals scarious marginally; adaxial petal 2.4–5.6 × 2.1–6.7 cm, flabellate, tubular, apex rounded, cripsed, base attenuate, purple, lilac to red or purple, red, greenish to white, glabrous; petalodia 0.4–3.5 × 0.2–0.8 mm, white to pink with inner white; stamens inserted, joined basally in a diadelphous sheath, dorsal one free, longer filaments 2.0– 4.2 cm long, shorter filaments 1.4–2.8 cm long, sheath inequilateral, shorter side 2.7–10.0 mm long, longer side 4.4–13.6 mm long, sheath tomentose, pubescent to puberulous, free filaments glabrous, brownish-yellow, shorter stamens anthers 1.1–2.8 × 0.5–1.8 mm, rectangular, longer stamens anthers 1.7–3.6 × 0.6–2.2 mm, rectangular; ovary 3.3–7.5 × 1.7–4.4 mm, obovate, glabrous, fovoate or tomentose, brownish-yellow, 2–4 ovules, stipe 3.1–7.7 mm long, glabrous or tomentose, style 1.9–3.5 cm long, stigma obtuse to capitate. Legumes 3.5–15.4 × 5.2–13.8 cm, stipe 0.9–2.6 cm long, obovate to orbicular, sometimes oblongelliptic, apex obtuse to truncate, sometimes rounded, dorsal margin thickened, brown to dark-brown, veins absent, rarely with some inconspicuous transversal veins, glabrous, foveate to rugose. Seeds 1 per fruit, 4.3–10.8 × 3.8–7.7 cm, oblate, orbicular to obovate, brown to dark-brown.

Conservation status:— Categorized as Least Concern (LC) according to the IUCN criteria ( IUCN 2012, IUCN Standards and Petitions Committee 2022). The estimated extent of occurrence for the species is 52,541.927 km 2 and the estimated area of occupancy is 228.000 km 2.

Eperua glabriflora has two varieties: E. glabriflora var. glabriflora and E. glabriflora var. gynopubescens .