Caecum insculptum Carpenter, 1857

Raines, Bret K., 2020, A Rosetta Stone for eastern Pacific Caecidae (Gastropoda: Caenogastropoda), Zootaxa 4827 (1), pp. 1-146 : 54-56

publication ID	https://doi.org/ 10.11646/zootaxa.4827.1.1
publication LSID	lsid:zoobank.org:pub:2CE528C2-DE07-4155-B37D-BE2C342A3901
DOI	https://doi.org/10.5281/zenodo.4457763
persistent identifier	https://treatment.plazi.org/id/03A3DF49-FFD7-FFB6-FF44-FB8AFB2B397E
treatment provided by	Plazi
scientific name	Caecum insculptum Carpenter, 1857
status

Treatment

Caecum insculptum Carpenter, 1857 View in CoL

Figures 36 View FIGURE 36 A–F, 37A–H, 38A–B, 127I

Caecum (Elephantulum) insculptum Carpenter, 1857: 315 View in CoL , 2 syntypes NHMUK 1857.6 About NHMUK .4.1514, Mazatlan.

Caecum (Elephantulum) insculptum View in CoL — Carpenter, 1858: 420.

Caecum insculptum View in CoL — Carpenter, 1860: 32; Palmer, 1951: 43; Brann, 1966: pl. 32, fig. 360; Raines & Pizzini, 2009: 120, figs. 4d, 5d, 7a–c.

Caecum mirabile de Folin, 1867a: 45 View in CoL , 12 syntypes MNHN-IM-2000-4593, Panama. (syn. nov.).

Elephantulum insculptum View in CoL — Keen, 1968: 419, figs. 89a–b; Keen, 1971: 398, fig. 478; Shasky, 1984: 27; Skoglund, 1992: 29.

Fartulum (Elephantulum) insculptum — Draper, 1985: 4, fig. 3.

Brochina insculptum — Lightfoot, 1993b: 76, fig. 21.

Material examined. In addition to the type material the following lots were examined. NHMLAC: LACM 1965 About LACM - 28.34 About LACM , Barra de Navidad , Jalisco, Mexico, intertidal, 3 sh ; LACM 1935 About LACM - 90.10 About LACM , Secas Islands , Chiriqui Prov., Panama, 45 m, 1 sh ; LACM 1971 About LACM - 83.56 About LACM , Los Arcos, Banderas Bay , Jalisco, Mexico, intertidal to 5 m, 13 sh ; LACM 1972 About LACM - 81.22 About LACM , S of Bocapan, Tumbes Dept. , Peru, intertidal, 1 sh ; LACM 1970 About LACM - 11.56 About LACM , Ancon Point, Santa Elena , Guayas Prov., Ecuador, intertidal, 40 sh ; LACM 1970 About LACM - 12.43 About LACM , Ancon Point, Santa Elena , Guayas Prov., Ecuador, intertidal, 15 sh ; LACM 1972 About LACM - 42.68 About LACM , Ballena Point, Ballena Bay , Puntarenas Prov., Costa Rica, 3–16 m, 21 sh ; LACM 1972 About LACM - 53.34 About LACM , Herradura Bay , Puntarenas Prov., Costa Rica, 23 m, 12 sh ; LACM 1972 About LACM - 46.65 About LACM , Alcatraz Island , Tortugas Islands, Puntarenas Prov., Costa Rica, 2–8 m, 28 sh ; LACM 1972 About LACM - 30.29 About LACM , Santa Elena Point, Guanacaste Prov., Costa Rica, 13–16 m, 12 sh ; LACM 1972 About LACM - 38.33 About LACM , Penca Point, Guanacaste Prov., Costa Rica, 8–13 m, 3 sh ; LACM 1934 About LACM - 133.29 About LACM , Tangola-Tangola Bay , Oaxaca, Mexico, 5–6 m, 1 sh ; LACM 1972 About LACM - 57.44 About LACM , Quepos, Puntarenas Prov., Costa Rica, 24 m, 11 sh ; LACM 1972 About LACM - 58.63 About LACM , Quepos, Puntarenas Prov., Costa Rica, 10–24 m, 23 sh ; LACM 1968 About LACM - 41.89 About LACM , Cuastecomate Bay , Jalisco, Mexico, 5–21 m, 6 sh ; LACM 1968 About LACM - 45.53 About LACM , Cuastecomate Bay , Jalisco, Mexico, 3–6 m, 1 sh ; LACM 1966 About LACM - 15.32 About LACM , El Tule, Canelo Bay , Baja California, Mexico, intertidal, 2 sh ; LACM 1946 About LACM - 10.17 About LACM , Creston Island , Mazatlan, Sinaloa, Mexico, intertidal, 3 sh ; LACM 1935 About LACM - 136.3 About LACM , Secas Islands , Chiriqui Prov., Panama, 22 m, 1 sh ; LACM 1967 About LACM - 15.17 About LACM , Banderas Bay , Jalisco, Mexico, 10–21 m, 16 sh . SBMNH: SBMNH 452507 About SBMNH , Playa Tamarindo, Guanacaste, Costa Rica, 5–15 m, 4 sh ; SBMNH 127147 About SBMNH , Chacala, Nayarit, Mexico, 3–5 m, 1 sh ; SBMNH 452488 About SBMNH , La Puntilla, Salinas , Guayas, Ecuador, intertidal, 2 sh ; SBMNH 619528 About SBMNH , Bahia Ballena , Puntarenas, Costa Rica, 13–15 m, 6 sh ; SBMNH 128135 About SBMNH , Isla Lobos, Mazatlan , Sinaloa, Mexico, 6 sh ; SBMNH 620796 About SBMNH , Islas Ladrones, Chiriquí, Panama, 10–25 m, 1 sh ; SBMNH 620799 About SBMNH , Bahia Tenacatita , Cabo Tamarino, Jalisco, Mexico, 8–10 m, 1 sh ; SBMNH 620806 About SBMNH , Isla Pajaro [Mogo Mogo], Panama Prov. , Panama, intertidal, 3 sh ; SBMNH 621369 About SBMNH , Bahia Banderas , Nayarit, Mexico, 6 sh ; SBMNH 621531 About SBMNH , Barra de Navidad , Jalisco, Mexico, 3–45 m, 1 sh ; SBMNH 621533 About SBMNH , Bahia Herradura , Puntarenas, Costa Rica, intertidal, 1 sh ; SBMNH 621558 About SBMNH , Playa Olas Atlas, Sinaloa, Mexico, 10–25 m, 5 sh ; SBMNH 622373 About SBMNH , Punta Mirador, Puerto Lopez , Manabi, Ecuador, intertidal, 1 sh ; SBMNH 622381 About SBMNH , Punta Ancon, W side, Guayas, Ecuador, intertidal, 20 sh. BCD: Navidad Bar , Colima, Mexico, 16–32 m, 3 sh ; off Piedras Blancas, Nayarit, Mexico, 18 m, 10 sh .

Original diagnosis. “ C. ( Elephantulum ) t. juniore vitrea, diaphana, adulta solida; liris validis quadratis et interstitiis profundis ornata; aperturam versus haud tumente; septo ungulato, subprominente, apice obtuso, paululum dextrorsum verso, margine laterali subrecto; operculo?” Carpenter (1857: 315).

Description. Protoconch unknown. Teleoconch I short, sculptured with narrow longitudinal ribs and weak striae, crossed by fine axial growth lines ( Fig. 37C View FIGURE 37 ), transition to teleoconch II marked by axial incised line ( Fig. 37A View FIGURE 37 ). Teleoconch II about 4.5 times length of teleoconch I, sculptured with identical characters as previous stage ( Fig. 37C View FIGURE 37 ), transition to teleoconch III marked by axial incised line ( Fig. 37A View FIGURE 37 ). Teleoconch III sculptured with low, flattened, longitudinal ribs and weak striae, crossed by axial growth lines ( Fig. 37D View FIGURE 37 ), transition to teleoconch IV marked by rapid increase in diameter ( Fig. 37B View FIGURE 37 ). Teleoconch IV sculptured with rounded longitudinal ribs and weak striae ( Fig. 37F View FIGURE 37 ), transition to teleoconch V marked by suture ( Fig. 38A View FIGURE 38 ). Teleoconch V (last growth stage) large for genus [Tol 2.61–3.10 mm], tubular, thick, robust, mildly to moderately and regularly arched [Larc: 0.90–1.03 mm; Arc: 0.22–0.25 mm], subcylindrical, with slight increase in diameter from posterior to aperture, semi-translucent white ( Figs. 36 View FIGURE 36 A–B). Longitudinal sculpture comprised of 15–20 strong, regularly spaced ribs ( Fig. 37H View FIGURE 37 ). Longitudinal interspaces wider than ribs ( Figs. 37 View FIGURE 37 G–H). Surface glossy, with fine, continuous, longitudinal striae in interspaces of longitudinal ridges ( Fig. 37G View FIGURE 37 ). Posterior [Dpe: 0.49–0.55 mm] with thick edge, squared shoulder ( Fig. 36C View FIGURE 36 ). Septum mucronate, inflated ( Fig. 36C View FIGURE 36 ). Mucro heel-like, positioned on dorsal margin ( Fig. 36C View FIGURE 36 ). Aperture [Da: 0.60–0.74 mm] thick, slightly flared ( Fig. 38H View FIGURE 38 ). No varix. Lip smooth, with deflected peristome ( Fig. 36C View FIGURE 36 ). Periostracum thin, smooth, light tan ( Fig. 36A View FIGURE 36 ). Operculum thin, light brown; exterior surface with 6–7 concentric rings ( Figs. 36 View FIGURE 36 D–E). Conceptual reconstruction of growth stages shown in Fig. 38A View FIGURE 38 .

Distribution and Habitat. From Saladita Bay, Guaymas, Mexico to Tumbes, Peru. Intertidal to 45 m. Uncommon subtidally on sand and rubble substrates.

Remarks. The type material of Caecum insculptum is comprised of one subadult and one adult (NHMUK 1857.6.4.1514, Fig. 36F View FIGURE 36 ). Raines and Pizzini (2009: 120) made reference to a “ Caecum insculptum ” complex, which represents a group of caecids with longitudinal lines and grooves. Of all the eastern Pacific species, C. insculptum has the most well-developed longitudinal grooves. Raines and Pizzini (2009: 116) also suggested C. mirabile and C. mirificum de Folin, 1867 were synonyms. However, further examination suggests that the type material of C. mirabile (MNHN-IM-2000-4593, Fig. 36G View FIGURE 36 ) is actually conspecific with C. insculptum .

Kingdom	Animalia
Phylum	Mollusca
Class	Gastropoda
Order	Littorinimorpha
Family	Caecidae
Genus	Caecum