Cochlostoma (T.) gracile (Pfeiffer, 1849)

Zallot, Enrico, Kamchev, Panche, Schilthuizen, Menno, Fehér, Zoltán, Mattia, Willy De & Gittenberger, Edmund, 2024, Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae), European Journal of Taxonomy 927, pp. 1-163 : 26-28

Cochlostoma (T.) gracile (Pfeiffer, 1849)

Figs 15G , 21 (red dots), 22

Cyclostoma gracile Pfeiffer, 1846 –1850 ( 1846?): 191, pl. 26 figs 28–30.

Types

Not seen.

Other specimens

CROATIA • 1- Omiš 2 (topotypical); 43.4425° N, 16.7317° E; 2011; De Mattia leg.; EZ1093 • 1- Omiš 1 (topotypical); 43.4428° N, 16.7319° E; 1998; De Mattia leg.; WdM6667 • 1- Omiš 3 (topotypical); 43.4428° N, 16.7319° E; 1985; Pintér leg.; HNHM43203 .

Type locality

CROATIA • Split-Dalmatia, Omiš ; 43.4425° N, 16.7317° E.

Description

SHELL. Closely spaced riblets on last part of protoconch. Teleoconch spotless, with strong and well-spaced ribs. Lip is rather weak. Columellar lobe bent inwardly to cover umbilicus.

MEASUREMENTS. 2 ♀♀: whorls =8.1–8.6, H = 6.7–7.2 mm, H/W=2.66–2.89, roundness=0.16–0.17, ribs incl.= 57–57°, apert. incl.= 21–23°, ribs/mm 1 st wh.= 5–8, ribs/mm 4 th wh.=5–7.

FEMALE GENITAL ORGANS. As in C. (T.) stussineri .

Remarks

This species is considered widespread with several subspecies all along the Balkan Peninsula, reaching south Greece with C. (T.) gracile subaiorum Schütt, 1977 . Our findings, however, seem to indicate that it has in fact a limited range. Its p-distances from taxa currently considered as subspecies like stussineri and reitteri are 2.9 and 3.6%, respectively.

Figures

Fig. 15. Shell and female genitals of clade A samples with amplified 16S which share the same female genital set-up. A. NFS158, Tisovec, SLO (WdM-6748). B. Cochlostoma (T.) reitteri (Boettger, 1880), Kapela Pass, HR (WdM-6882). C. C. (T.) stussineri (Wagner, 1897), Rakow Skocjan, SLO (WdM-6918). D. NFS064, Terminillo V. Dell’Inferno, I (EZ-1067). E. C. (T.) patulum (Draparnaud, 1801), Cirque D’Archiane, FR (RMNH-117454). F. NFS153, 3 km SE of Oltari, HR (HNHM-100381). G. C. (T.) gracile gracile (Pfeiffer, 1849), Omiš 1, HR (WdM-6667). H. C. (T.) gracile gracillimum (Wagner, 1901), Vrbac Valley, BeH (Gr001).

Fig. 21. Distribution of the samples of Cochlostoma (T.) gracile (Pfeiffer, 1849): red dots= C. (T.) gracile gracile; green dots =C. (T.) gracile gracillimum (Wagner, 1901); cyan dots = C. (T.) gracile croaticum (Pfeiffer, 1870). A black center indicates samples with amplified 16S.