Coelichneumon lividusus ( Uchida, 1925 )

Riedel, Matthias & Watanabe, Kyohei, 2021, The genus Coelichneumon Thomson in Japan (Hymenoptera, Ichneumonidae, Ichneumoninae), Zootaxa 4948 (4), pp. 501-545 : 524-526

Coelichneumon lividusus ( Uchida, 1925)

Hadrojoppa lividusa Uchida, 1925: 448 , ♂.

Hadrojoppa lividusa var. areolata Uchida, 1925: 448 , ♂, synonymized by Townes et al. (1965).

Hadrojoppa tibialis Uchida, 1925: 452 , ♀, synonymized by Townes et al. (1965).

Holotype of H. lividusa : (♂) “[Maruyama] 10.10.29; [red] Type; Hadrojoppa lividusa sp. n.; [red] Hadrojoppa lividusa Uchida type” (Sapporo), type studied. Missing: tip of right flagellum.

Holotype of H. lividusa var. areolata : (♂) “ Hadrojoppa lividusa var. areolatus n. var.; [red] Type; 3/ VI.18/ Sapporo Matsumura; [red] Hadrojoppa lividusa var. areolata Uchida —type” (Sapporo), type studied. Missing: left flagellum, left fore wing.

Holotype of H. tibialis : (♀) “[Tokyo]/ Japan T.Uchida; [red] Type; Coelichneumon tibialis sp.n. det Matsumura; [red] Coelichneumon tibialis Uchida –type” ( Sapporo ), type studied. Missing: right fore tibia and tarsus, right 2 nd to 5 th mid tarsomeres, right 3 rd to 5 th hind tarsomeres, left 2 nd to 5 th mid tarsomeres. Metasoma broken off, glued on label.

Non-type material: JAPAN: 1 ♂, Tokyo, Oume City, Mt. Otakeyama , 1.vi.2008, H. Katahira leg. ; 1 ♂, Fukui Pref., Oono City, Arashimadake , 6.vi.1982, H. Kurokawa leg. ; 1 ♂, Fukushima Pref., Hinoemato Vil. , 19.vii.– 1.viii.2004, H. Makihara leg. , MT; 1 ♀, Saitama Pref., Niiza City, Heirinji , 11.ix.1985 ; 1 ♂, Tokyo, Oume City, Mt. Mitake-san , 1.vi.2008, M. Gunji leg. ; 1 ♀, Saitama Pref., Yarii Town , 3.xi.2013, T. Nambu leg. ; 1 ♀, Nagano Pref., Outaki Vil., Mt. Outake-san, Hakkaisan , 1780–1859 m alt., 4.viii.2017 .

Description: ♀: Body length 18‒20 mm. Flagellum strongly lanceolate, with 46‒47 flagellomeres; 1 st flagellomere (without anellus) 1.7‒1.9× longer than wide, 2 nd flagellomere 2.2× longer than wide, combined length c.0.56× eye length, 10 th flagellomere square, flagellum beyond the middle attenuated and strongly widened, widest flagellomeres 2.0‒2.2× wider than long. Temple moderately and roundly narrowed behind the eye. OED 1.2× and OOD 1.0× ocellar diameter. Frons with transverse rugae, face densely punctate and finely granulate. Clypeus with dense or partly scattered punctures, smooth and shining. MI 0.55‒0.65. Gena laterally c.1.0× as wide as eye, with very scattered punctures ventrally. Genal carina reaching hypostomal carina far from mandibular base.

Mesosoma covered with pale hairs. Notaulus impressed in frontal 1/4 of mesoscutum. Mesoscutum with dense punctures, finely granulate and slightly opaque. Mesopleuron with coarse punctures, speculum large, smooth and shining; metapleuron with coarse punctures, partly striate; juxtacoxal carina strong. Scutellum moderately elevated, punctate, about as long as wide, without lateral carina, with scattered punctures. Propodeum rounded; area superomedia slightly elevated, heart-shaped, about as long as wide or longer than wide, with finely rugose or smooth surface, anterior transversal carina slightly caudal to its middle. Area externa with scattered punctures and some fine rugae. Hind coxa densely punctate, with strong brownish scopa (length c.1/4 to 2/5 of coxal length). Hind femur 3.3‒3.5× longer than wide, with dense punctures. Tarsomeres distinctly widened; 3 rd mid tarsomere 1.2‒1.4× longer than wide. Areolet pentagonal, frontal distance between veins 2rs-m and 3rs-m 2‒3× their width; vein 2m-cu in its middle. Vein 1cu-a postfurcal by 2× its width.

Metasoma strongly oxypygous. Lateral field of postpetiolus punctate, c.0.5× as wide as middle field, the latter with c.18 rather coarse striae. 2 nd tergite as long as wide to 1.1× wider than long. Thyridium moderately oblique, 0.9‒1.2× wider than the interval; interval narrower than median field of postpetiolus; 2 nd to 4 th tergites finely and densely punctate and aciculate medially; 5 th tergite with superficial punctures, ± shining. 7 th tergite 1.2× wider than long. Seen from dorsal, ovipositor sheath c.0.5× as long as 7 th tergite.

Color: Black. Metasoma black, with distinct bluish shine. Stripes on flagellomeres 8/9‒13/14, sometimes spot on mandibular base, stripe of frontal orbit confluent with triangular spot on vertex, collare and hind edge of pronotum, sometimes small posterolateral spots of scutellum, spots on all trochanters, subbasal rings of all tibiae, and external stripes on tarsomeres yellow. Wings slightly infuscate; pterostigma reddish-brown to blackish.

♂: Body length 17‒24 mm. Flagellum with 40‒46 flagellomeres, 1 st flagellomere 2.1‒2.5× longer than wide. Tyloids on flagellomeres 6/8‒16/18, long-oval, centrally situated, maximally 0.6× as long as the flagellomeres. Temple strongly and roundly narrowed behind eye. OED 0.9‒1.3× and OOL 1.0× ocellar diameter. Frons with transverse rugae. Face densely punctate. Clypeus and gena with rather dense punctures. Gena c.0.9× as wide as eye. MI 0.2‒0.4.

Notaulus distinct in frontal 1/3 of mesoscutum. Mesoscutum punctate and granulate, opaque. Mesopleuron with rather fine punctures; metapleuron with coarse dense punctures; juxtacoxal carina present. Scutellum moderately elevated and slightly longer than wide, without lateral carina, with rather dense punctures. Area superomedia ± elevated above surrounding areas, heart-shaped, ± wider than long, smooth with some fine puncures; anterior transversal carina in its caudal third. Area externa punctate, with some fine striae. Area petiolaris with coarse transverse rugae. Fore metatarsus with apical tooth. Hind femur with dense strong punctures, 3.7‒4.3× longer than wide.

Lateral field of postpetiolus 0.6‒0.7× as wide as the median field, the latter with c.15 strong striae. Thyridium almost transverse, c.1.0× as wide as the interval; interval narrower than median field of postpetiolus. 2 nd tergite slightly 1.0‒1.1 × as long as wide. 2 nd to 4 th tergites densely punctate and aciculate medially; 5 th tergite with superficial punctures and some aciculation basally.

Color: Black. Metasoma with weak bluish shine. Palps, mandible except teeth, clypeus, face, frontal orbit usually confluent with stripe on vertex, ventral outer orbit, scapus ventrally, collare and upper margin of pronotum, spot on tegula, subtegular ridge, small spot on mesopleuron in front of epicnemial carina, sides or dumb-bell shaped spot of scutellum, frontal spots on fore and mid coxae, all trochanters and trochantelli, subbasal rings of all tibiae, frontal stripes on fore and mid femora and tibiae, fore and mid tarsi completely, and external stripes on 1 st to 4 th hind tarsomeres yellow. Wings slightly yellowish or brownish infuscate, pterostigma dark reddish-brown.

Distribution: Japan (Hokkaido, Honshu and Shikoku) ( Uchida 1925, Konishi & Yamamoto 2000).