Boholdoyidae Kovalev

Lukashevich, Elena Dmitrievna & Ansorge, Jörg, 2024, First Upper Triassic Diptera (Insecta) from Germany, Zootaxa 5403 (1), pp. 115-129 : 122

Family Boholdoyidae Kovalev in Kalugina & Kovalev, 1985

Type genus. Boholdoyia Kovalev , 1985 (Novospasskoye, Transbaikalia, Russia, J 1–2 – K 1)

Composition. Besides the type and the new genus, the monotypic Daiamyi a Kovalev, 1990 from the Upper Jurassic or Lower Cretaceous of Daya, Transbaikalia, Russia.

Remarks. V. Kovalev described the family based on one male with badly preserved wings and the diagnosis of the family included a lot of body characters (Kovalev in Kalugina & Kovalev 1985). Later two males were described from other Mesozoic deposits and the venation of the family became clear ( Kovalev 1990). The new Upper Triassic isolated wings from Germany share the same pattern of venation. The following characters, visible in the Triassic wings, are diagnostic for the family (so they are not mentioned in the diagnosis of the new genus below): C extends along entire wing, strongly thickened at the anterior and distal margin, weak but distinct along the posterior margin, Sc weak, shorter than half of the wing, R 1 ends about wing midlength, Rs stem long, Rs with three long free branches, R 2+3 begins proximal to r-m and ends in the wing margin far from R 1, R 4+5 fork several times longer than distal portion of R 4+5 stem, R 5 ends in the wing apex, basal cells closed; ba longer and narrower than bp, ends near wing midlength, three medial veins, M 1+2 fork distal to r-m.