Pachyrhinus elegans (Couper 1865)

Benzel, Joseph & Bright, Donald E., 2021, A revision of the genus Pachyrhinus Sch ӧnherr 1823 (Coleoptera: Curculionidae Entiminae) in the Nearctic Region, Zootaxa 4999 (6), pp. 501-533: 509-516

publication ID

https://doi.org/10.11646/zootaxa.4999.6.1

publication LSID

lsid:zoobank.org:pub:2355BCA5-4260-4CDF-8545-8FD854D3E6A5

persistent identifier

http://treatment.plazi.org/id/039F9144-FFEC-F87F-40AA-F9C6FA811FFF

treatment provided by

Plazi

scientific name

Pachyrhinus elegans (Couper 1865)
status

 

Pachyrhinus elegans (Couper 1865)  

Polydrusus elegans Couper 1865a: 63   ; Becker 1974:70.

Scythropus elegans: Horn 1876: 116   ; Chittenden 1890: 168; Fall 1901: 309; Blatchley and Leng 1916: 131; Hatch 1971: 274; Furniss and Carolin 1977: 336.

Pachyrhinus elegans: O’Brien & Wibmer and Wibmer 1982:43   ; McNamara 1991: 330; Downie and Arnett 1996: 1485; Majka et al. 2007; Bright and Bouchard 2008: 159.

Scythropus lateralis Casey 1888: 276   ; Fall 1901:309. NEW SYNONYMY.

Pachyrhinus lateralis: O’Brien & Wibmer and Wibmer 1982: 43   .

Scythropus miscix Fall 1901: 309   . NEW SYNONYMY.

Pachyrhinus miscix: O’Brien & Wibmer and Wibmer 1982: 43   .

Diagnosis. Members of P. elegans   may be distinguished from those of other Nearctic Pachyrhinus species   by the non-mottled scale pattern on the elytra ( Figs. 39–40 View FIGURES 39–44 ) as opposed to the mottled scale patterns in P. californicus   and P. cinereus   . It is the only Nearctic Pachyrhinus species   to have an iridescent green coloration, though this feature is variable ( Fig. 37 View FIGURES 33–38 ). The aedeagus of P. elegans   ( Figs. 41–43 View FIGURES 39–44 ) is less arcuate than those of P. californicus   and P. cinereus   . It also bears a blunt tip rather than the long, tapered tip of other species. The spermatheca in P. elegans   ( Fig. 44 View FIGURES 39–44 ) is less tightly coiled making it C-shaped rather than comma-shaped as in P. californicus   .

Description (male). Length: 4.5–8.0 mm, average 6.0 mm ( Figs. 33-38 View FIGURES 33–38 ); color black, red on legs and antennae ( Fig. 34 View FIGURES 33–38 ); vestiture composed of elongate, appressed, contiguous, mostly non-overlapping scales, short, recurved and long, erect setae sparsely arranged throughout body; scales ( Fig. 6 View FIGURES 6–9 ) 2.0× long as wide, with rounded tip bearing small spines, green colored ( Fig. 10 View FIGURES 10–15 ); scales lighter colored on underside ( Fig. 35 View FIGURES 33–38 ) and in bands on 1st, and 7th through 10th striae ( Fig. 34 View FIGURES 33–38 ). Antennomers III to VII conical, subequal in length ( Fig. 19 View FIGURES 16–19 ); elytra uniformly covered in dense scales ( Fig. 39 View FIGURES 39–44 ); scales uniform in color along strial rows; rows II through VI iridescent green in color, other rows lighter ( Fig. 40 View FIGURES 39–44 ); appressed setae arranged in rows along mid region of interstriae, longitudinally placed at every third interstrial puncture. Aedeagus mesally curved ( Fig. 41 View FIGURES 39–44 ). Lateral margins gradually curved, ending in point covered by long fine setae ( Figs. 42–43 View FIGURES 39–44 ). Ostium elongate, ovate, arcuate, laterally emarginated, terminating in blunt point with small indentation at tip.

Description (female). Similar to male; spermatheca C-shaped ( Fig. 44 View FIGURES 39–44 ), ramus elongate, subconical; cornu elongate, recurved, apically narrowed.

Variation. Coloration in this species is highly variable ( Figs. 45-53 View FIGURES 45–53 ), commonly copper ( Figs. 45–48 View FIGURES 45–53 ) or iridescent green ( Figs. 49–50 View FIGURES 45–53 ), with blue ( Figs. 51–53 View FIGURES 45–53 ) abundant in northern Rocky Mountains, British Columbia and the northwestern United States; integument ranging from black to orange, corresponding with scale color; funicular antennomeres either subequal in size or decreasing in size to 5 th antennomere, then increasing in size.

Distribution ( Fig. 54 View FIGURE 54 ). Pachyrhinus elegans   occurs commonly throughout the coniferous forests of temperate North America, ranging from New Brunswick to Pennsylvania in the east and British Columbia and Alberta to northern California and Colorado in the west. In collections studied, no specimens occurred in central regions of North America between Indiana and the Front Range of Colorado. Specimens throughout the range are morphologically very similar with the exception of variation of scale color.

Material Examined (803 specimens). Type Material, Lectotype, designated by Becker (1974), is in the collection of Laval University, Quebec City, Quebec, and was examined and photographed ( Figs. 33–38 View FIGURES 33–38 ). P. lateralis   holotype, USA, California, Lake Co. T. L. Casey ( USNM). P. miscix   holotype, USA, California, Sacramento Co. H. C. Fall. (MCZ).

CANADA, Alberta, Banff StP. VI-10-1897 / Hubbard & Schwartz ( USNM: 3 males, 6 females)   ; 1922-V-25 /C. B. D. Garrett ( CNCI 1 male, 2 female)   ; Burmis. VI-15-1952 / K. C. Herrman ( CNCI: 1 male, 1 female)   ; Crow’s Nest. VI-9-1930 / J. H. Post ( CNCI: 2 female)   ; VI-17-1962 / K. C. Herrman ( CNCI: 1 female)   . Waterton. VI-8-1962 / W. R. M. Mason ( CNCI: 2 male)   ; VII-16-1970 / R. A. Kelton ( CNCI: 1 male)  

British Columbia, Aspen Grove. V-14-1934 / ponderosa pine, H. B. Leech ( USNM: 1 male, 1 female)   ; V-25- 1934 / ponderosa pine, ( USNM: 1 male)   ; Bagneg L. V-9-1976 / B. F. & J. L. Carr ( CNCI: 1 male, 2 female)   ; Chopaka. VI-24-1923 / R. Glendenning ( CNCI: 1 female)   ; Corbin , VII-11-1961 / B. F. & J. L. Carr ( CNCI: 1 male)   ; Cranbook. V-12-1922 / C. B. D. Garrett ( CNCI: 1 male, 7 female)   ; Cranston. V-8-1958 / Douglas fir, H. & A. Howden ( CNCI: 4 male, 9 female)   ; Creston. V-12-1956 / G. S. Smith ( CNCI: 2 males, 2 female)   ; Elko. V-7-1958 / Douglas fir, H. & A. Howden ( CNCI: 1 male)   ; Falkland. V-18-1944 / H. B. Leech ( UCRC: 1 male, 1 female)   ; Fife. VI-6-1959 / Douglas fir, H. & A. Howden ( CNCI: 1 female)   ; Fry Cr. below Bridge, Purcell Mtns. Wilderness Pk. V-28-2010 / R. W. Baumann ( CWOB: 1 male)   ; Goldbridge. VI-29-1942 / H. B. Leech ( UCRC: 1 female)   ; Golden , VI-27-1976 / B. F. & J. L. Carr ( CNCI: 1 female)   ; Kaslo. IV-23-1929 / ( CNCI: 2 male)   ; V-10-1939 / ( CNCI: 2 males, 1 female)   ; II-7/ R. P. Curie ( CWOB: 1 female)   ; Kuskanoo. V-12-1958 / Douglas fir, H. & A. Howden ( CNCI: 1 male)   ; Oliver ; V- 23-1958 / Douglas fir, H. & A. Howden ( CNCI: 1 male)   ; V- 27-1959 / L. A. Kelton ( CNCI: 1 female)   ; V-27-1959 / R. E. Leech ( CNCI: 1 female)   ; Osoyos , V-28-1934 / G. S. Walley ( CNCI: 1 female)   ; V-30-1958 / Douglas fir, H. & A. Howden ( CNCI: 2 female)   ; Radium. VI-4-1953 / B. F. & J. L. Carr ( CNCI: 2 male, 1 female)   ; Salmon Arm. VI-29- 1923 / E. R. Buckell ( CNCI: 1 female)   ; V-29-1928 / ( CNCI: 1 female)   ; III-24-1929 / H. B. Leech ( USNM: 1 female)   ; IV-24-1933 ( USNM: 1 male)   ; V-11-1933 ( USNM: 1 female)   ; IV-4-1934; ( USNM: 14 male, 13 female)   ; Salmo. VI- 27-1942 / H. B. Leech ( UCRC: 1 female)   ; VI-9-1968 / Campbell & Smetana, ( CNCI: 1 male)   ; Sanca Cr. Hwy. 3A, S of Sanca. VI-1-2010 / R. W. Baumann ( CWOB: 1 female)   . Sirdar. IV-12-1980 / I. Askevold ( CWOB: 3 males, 2  

females); Skookumchuck , V-30-1984 / B. F. & J. L. Carr ( CNCI: 1 male)   ; Summit Cr. Hwy. 3, Blazed Cr. rest area. VI-2-2010 / R. W. Baumann ( CWOB: 1 female)   ; Terrance. VII-5-1960 / ( CNCI: 1 female)   ; VI-8-1960 / W. W. Moss ( CNCI: 1 female)   ; VI-16-1960 / R. J. Pilfrey ( CNCI: 1 male)   ; Vernon. VI-24-1923 / E. P. Venables ( CNCI: 1 male)   ; IV-17-1934 / H. B. Leech ( USNM: 1 female)   ; Wardner. V-19-1927 / ( CNCI: 1 female)   ; Windmire. IV-23-1955 / B. F. & J. L. Carr, ( CNCI: 16 male, 3 females, 3 unknown)   .

New Brunswick, Charlo. VIII-24-1943 / ( CNCI: 1 female); Kouchibouguac NP. V-24-V-3- 1977 / S. J. Miller ( CNCI: 3 female)   .

Nova Scotia, Kentville , V-23-VI-24-1923 / R. P. Gorham ( CNCI 4 males); Lunenburg, V-5-1969 / P. Ward ( CNCI: 1 male)   .

Ontario, Ad & Lennox. June-25-1950 / J. F Brimley ( CNCI 1 male, 2 females)   ; Arkell. IV-30/ H. H. Holmes ( USNM: 1 female)   Belleville. ( CUIC: 2 males, 2 females)   ; Bells Corners , V-5-1957 / E. Macdougall ( CNCI: 1 female)   ; VI-19-1963 / D. D. Munroe ( CNCI: 1 female)   ; Constance Bay , 1935-VI-10 / W. J. Brown ( CNCI, 1 male)   ; Hastings co. VII-10-1938 / Brimely ( CNCI: 3 males, 2 female)   ; Marmora , IV-18-1942 / white pine, J. F. McAlpine ( CNCI: 6 male, 7 female)   ; Pettit. Hubbard and Schwartz ( USNM: 2 males, 3 females)   ; Trenton. J. D. Evans ( USNM: 2 males, 4 females)   ; W. Robinson ( USNM: 3 females)   ; Toronto. J. R. Ciew ( CUIC: 2 males, 1 female, 4 undetermined)   .

Quebec, Aylmer, V-9-1932 / W. J. Brown. ( CNCI: 1 male. 1 female); Val de Bois , VI-17-1973 / R. Sexton ( CNCI: 2 female)   .

USA, California, Amador Co., Tragedy Spring, Hwy. 88. VI-21-2006 / S. M. Clark & M. H. Goodman ( CWOB: 1 female)   ; El Dorado Co., S fk. American R., Hwy. 50, Riverton. III-21-1985 / R. W. Baumann ( CWOB: 1 male)   ; Flume, vic. of Pacific House. V-26-1989 / R. D. Gordon ( USNM: 15 males, 1 female)   ; Glenn Co., Plaskett Sta. VI-30-1967 / W. Gagne ( CWOB: 1 female)   ; Mariposa Co., VII-4-1952 / J. H. McLeod ( CNCI: 5 males, 9 female)   ; Modoc Co., Soup Springs. V-22-2007 / B. C. Kondratieff, R. W. Baumann ( CSUC: 1 female)   ; Mono Co., 3 mi N of Mammoth Lakes. D. G. Kissinger ( USNM: 5 males, 7 females)   ; Napa Co., Mt. St. Helena. V-30-1915 / F. E. Blaisdell ( CLR: 1 female)   ; Placer Co., Dutch Flat. III-9-1909 / ( UCRC: 1 male)   ; San Diego Co., Alpine. VII-1934 ( CNCI: 1 male, 3 female)   . Shasta Co., Hat Cr., Hwy 89, Honn campground. V-25-2007 / B. C. Kondratieff, R. W. Baumann ( CSUC: 2 females)   ; Manzanita L. VI-16-1959 / Kelton & Madge ( CNCI: 1 male)   ; Trinity Co., V-20-1934 / F. R. Platt ( UCRC: 1 male)   ; Tulare Co., Giant Forest . 1904/ Wickham ( USNM: 2 males, 2 females)   .

Colorado, Arapahoe Co., Arapaho Pk. VII-29-1968 / L. A. Kelton ( CNCI: 1 female)   ; Boulder Co., Eldora. VI-19-1982 / ponderosa pine, G. H. Nelson ( CWOB: 1 male)   ; Left Hand Canyon. IV-19-2005 / B. C. Kondratieff. ( CSUC: 1 male)   ; Nederland. V-27-1961 / W. R. M. Mason. ( CNCI: 1 female)   ; Clear Cr. Co., Argentine Rd. Wickham. ( USNM: 1 female)   ; Jefferson Co., Lookout Mtn. VII-3-1927 / A. Nicolay ( USNM: 1 female)   ; V-25/ VI-14- 2010 / CO dept. AG. ( CSUC: 1 female)   ; Larimer Co., 31 km. W Ft. Collins. IV-25-1972 / dwarf mistletoe, R. B. Penfield ( USNM: 1 female)   ; V-2-1972 / dwarf mistletoe, R. B. Penfield ( USNM: 1 female)   ; 2 mi. S, NW of Mt. Margaret trail, Red Feather Lakes area , CR74E 1 mi. off Parvin Lake. V-28-2010 / ponderosa pine, D. Leatherman. ( CSUC: 1 male, 10 females)   ; Estes Pk. VI-19-1961 / B. H. Poole ( CNCI: 4 males, 8 females)   ; Park Co., N Fk. IV-16- 1926 / H. G. Dyar ( USNM: 1 female)   ; Sada Cr. Pike NF, VII-19-1931 / ( USNM: 1 female)   ; Sagauche Co., Hayden Pass. VII-29-1968 / E. C. Becker ( CNCI: 3 females)   ; Ward Co., nr. Ward. VI-2/ VI-9-1933 / H. G. and H. E. Rodeck ( CWOB: 1 male)   .

Idaho, Benewah Co., St. Maries. IV-30-1963 / ponderosa pine ( USNM: 1 female)   ; Blaine Co., Pettit L. VII- 13-1968 / B. F. & J. L. Carr ( CNCI: 1 male, 1 female)   ; Boise Co., Boise NF. V-18-1954 / ponderosa pine ( USNM: 1 male, 1 female)   ; Idaho City. VI-26-1958 / Alpine fir, W. E. Cole (1 female)   ; Deadwood R. at Jct. S fk. Payette R. W of Loman. IV-23-1985 / R. W. Baumann & Nelson ( CWOB: 1 male)   ; Boise NF, N43.5637° W115.3976°. V-13- 2015 / White Pine, J. S. Benzel (5 females) GoogleMaps   ; Greyback Gulch , N43.4823° W115.5158°. V-13-2015 / ponderosa, J. S. Benzel   (8 males, 7 females) GoogleMaps   ; N43.4590° W115.5520°. V-13-2015 / Douglas fir, J. S. Benzel (12 males, 8 females); Bonner Co., Sand Point. V-23-1899 / U. S. Hopkins ( USNM: 1 females)   ; Bonner Co., Coeur d’Alene. IV-22-1920 / ponderosa pine, J. O. Evenden ( USNM: 4 males, 7 females)   ; Bonner Co., Sandpoint. VI-8-1950 / N. M. Downie ( CWOB: 2 males, 4 females, 1 undetermined); VI-11-1954 / N. M. Downie ( USNM: 2 females)   ; VI-10/ VII-10- 1971, ( CWOB: 1 male, 6 females)   ; VI-19-1973, ( CWOB: 1 female)   ; VII-15-1969 ( CWOB: 2 male)   ; VI-19-1973 ( USNM: 1 male, 5 females)   ; VII-9-1974 / N. M. Downie ( CWOB: 1 female)   ; Clark Co., Fairfield. VII-6-1960 / D. R. Smith ( USNM: 1 male, 1 female)   ; Idaho Co., Lochsa R., hwy. 12, Wendover campground. IV-28-1992 / R. W. Baumann & Zenger ( USNM: 1 female)   ; Latah Co., Moscow. V-20-1928 / J. E. Shull ( USNM: 1 male)   ; IV-20-1929 / H. A. Waters ( USNM: 2 females)   ; IV-29-1950 / N. M. Downie ( UCRC: 1 male, 1 undetermined)’ Big Sand Cr. V- 7-1975 / R. Haswell ( USNM: 8 males, 15 females)   ; Shoshone Co., Cedar Mtn., Moscow. IV-15-1934 / M. C. Lane ( UCRC: 6 males, 3 females, 1 undetermined); Shoshone Co. , Cathedral Peak , 11 mi SW of Heron   Montana. VII-2- 1962 / western white pine. D. G. Hellin ( USNM: 3 males, 1 female)   ; Valley Co., Krassel. V-23-1959 / M. M. Furniss ( USNM: 2 males, 6 females)   .

Maine, Cumberland Co., Douglas Hall. V-7/ V-8-1916 / A. S. Nicolay ( USNM: 1 male, 3 females); Oxford Co. , Porter. VI-12/ A. E. Brower ( USNM: 7 males, 6 females)   .

Massachusetts, Essex Co., Lawrence. VI-1864 / white pine (1 male, 1 female)   ; N of Sangus. VI-1906 / H. M. Russel (1 female)   ; Lawrence. ( CUIC: 4 undetermined); Franklin Co., Charlemont. IV/ ( USNM: 2 female)   ; Middlesex Co., Tyngsborough. V-17-1933 / ( CUIC: 4 undetermined); Worcester Co. , Southborough. V-8-1927 / C. A. Frost, ( CUIC: 1 male)   ; Winchandon , 1915-V-30 / C. A. Frost ( CNCI, 1 male)   .

Montana, Beaverhead Co., above Wise R. V-30-1933 / L. J. Liu & R. W. Baumann ( CWOB: 3 males, 6 females)   ; Gallatin Co., V-13-1902 / R. W. Cooley ( USNM: 1 female)   ; Gallatin Co., Bozeman. VI-6-1912 / R. W. Cooley ( USNM: 1 female)   . Granite Co., Sliderock Mtn. 16 mi SE of Clinton. V-18-1910 / whitebark pine, D. G. Fellen ( USNM: 1 male: 1 undetermined)   ; Flint Cr. Mtns., Racetrack Pk. VI-30-1961 / S. M. Sutton ( CWOB: 1 female)   ; Bonner. V-20-1958 / Western Larch , R. E. Denton ( USNM: 2 males, 3 females)   ; Lake Co., N of Polson. VI-22-1977 / Douglas fir, S. Kohler ( USNM: 1 female)   ; Lewis & Clark Co., Blackfoot R., Hwy. 279. V-21-1993 / R. W. Baumann & L. J. Liu ( CWOB: 2 males, 2 females)   ; Helena. IV-14/ V-5-1978 / Hubbard & Schwartz ( USNM: 2 males, 2 females)   ; Missoula. IV-20-1914 / ponderosa pine, J. Brunner ( USNM: 2 males, 1 female, 1 undetermined)   ; nr. Frenchtown. VI-13-1977 / Douglas fir, S. Kohler ( USNM: 2 females)   ; Musselshell Co., Roundup. VI-3-1982 / B. F. & J. L. Carr ( CNCI: 10 males, 1 female)   ; Ravalli Co., Gird Cr. IV-23-1933 / W. L. Jellison ( USNM: 1 female)   ; Burch Cr. IV-8-1934 / W. L. Jellison ( USNM: 1 female)   Sleeping Child Cyn. V-3-1937 / W. L. Jellison ( USNM: 1 female)   ; Gird Cr. IV-27-1941 / W. L. Jellison ( USNM: 2 females)   ; Bass Cr. Florence. IV-22-1979 / grand fir, S. Koeler ( USNM: 1 male, 1 females)   .

New Hampshire, Cheshire Co., Mohadrock. G. G. Milling ( USNM: 1 male)   ; Grafton Co., Hanover. ( USNM: 1 male)   , Rockingham Co., Hampton. I-14/ S. A. Shaw ( USNM: 1 male, 1 female)   , Stafford Co., Durham. ( USNM: 2 females)   , Sullivan Co., Claremont. VI / ( USNM: 7 males, 10 females)   .

New Jersey, [no data] ( USNM: 2 females)   .

New York, Erie Co., Buffalo. ( USNM: 2 females)   ; Essex Co., Cold R., High Peaks Wild Area, Saranac R. V-11-2006 / L. Myers ( CSUC: 1 male, 1 female)   ; Tompkins Co., Ithaca. V-5-1895 / Chitenden ( USNM: 6 males, 7 females, 1 undetermined)   ; IV-26-1915 / ( CUIC: 1 male)   ; 1937/ J. C. Bradley ( CUIC: 6 males, 3 females)   ; W Danby. V-30-1915 / ( CUIC: 1 female)   ; Westchester Co., Rye L. VI-3-1934 / H. L. McIntyre ( USNM: 2 female)   .

Ohio, [no data] ( USNM: 1 female)   .

Oregon, Baker Co., Baker. VI-2-1939 / J. H. Baker ( USNM: 4 males, 9 females)   ; Pine Cr. V-8-1945 / J. H. Baker ( USNM: 1 female)   . VI-14-1945 ( USNM: 1 female)   ; Sumpter , VI-15-1984 / B. F. & J. L. Carr ( CNCI: 1 male, 1 female)   ; Crook Co., Ochoco NF. V-27-1933 / M. C. Lane ( USNM: 6 males, 7 females)   ; Deschutes Co., 20 mi. S of Bend. V-27-1964 / K. Goeden ( CWOB: 1 male, 1 females)   ; Douglas Co., 15 mi W of Diamond L. VI-29-1971 / A. E. Lewis ( CWOB: 1 male)   ; Grant Co. Austin, V-25-1984 / B. F. & J. L. Carr ( CNCI: 1 female)   ; Prairie city, V-28-1984 / B. F. & J. L. Carr ( CNCI: 1 female)   ; Jackson Co., 10 mi. S of Ashland. IV-16-1959 / K. Goeden ( USNM: 1 female)   ; Squaw L. V-19-1962 / J. Schuh ( USNM: 1 male)   ; Hwy. 230, 14 mi N of Union Cr. VII-1-1971 ( CWOB: 1 female)   ; Klamath Co., Ft. Klamath. V-26-1933 / McLane ( USNM: 2 males, 1 female)   Lake Co., Lakeview. VI-28-1984 / B. F. & J. L. Carr ( CNCI: 1 male, 1 female)   ; Squaw Butte Cr. Hwy 140 E Lakeview. V-20-2013 / B. C. Kondratieff ( CSUC: 7 males, 6 females)   ; Lincoln Co., Newport. V-3-1965 / lodge pole pine, K. Goeden ( CWOB: 2 females)   ; Umatilla Co., Battle Mtn. V-27-1984 / B. F. & J. L. Carr ( CNCI: 3 males, 1 female)   ; Lehman Springs. VI-14-1984 / ( CNCI: 1 female)   ; Meacham. V-13-1933 / M. C. Lane ( USNM: 42 males, 31 females)   ; Union Co., Blue Mtns. VI- 11-1940 / C. A. Frost ( USNM: 4 males, 2 females)   . Wallowa Co., Enterprise. VI-13-1984 / B. F. & J. L. Carr ( CNCI: 2 male)  

Pennsylvania, Franklin Co., Mt. Alto. V-6/ J. N. Knull ( USNM: 2 males, 2 females)   ; 1931-IV-11/V-25/ A.C. Davis ( CNCI: 8 females, 10 males)   ; Warren Co., Bear L. ( USNM: 1 female)   .

Utah, Cache Co., Logan Cyn. VI-6-1933 / G. F. Knowlton ( USNM: 1 male); Duchesne Co. , Argyle Cr. Argyle Cyn. VI-15-1999 / R. W. Baumann ( CWOB: 1 male, 1 female)   .

Washington, Kittitas Co., Easton. V-18-1931 / ponderosa pine, W. W. Baker ( USNM: 3 males, 2 females)   ; Kittitas Co., Easton. / A. Koebele ( USNM: 5 males, 3 females)   ; Okanagan Co., Okanogan NF., Early Winters RS. V-30-1966 / D. S. Horning Jr. ( CWOB: 1 female)   ; Pierce Co., Mt. Rainier NP, Glacier Bridge. V-30-1997 / B. C. Kondratieff. ( CSUC: 1 male)   ; Spanaway. May-30-1925 / ponderosa pine, W. W. Baker (1 male, 1 female)   ; Walla Walla co., Walla Walla. May-1933 / M. C. Lane (4 males, 7 females)   ; Yakima Co., Tampico. May-30-1925 / M. C. Lane (1 female)   .

Wyoming, Teton Co., SE entrance, Yellowstone NP. VI-19-1947 / J. H. Baker ( USNM: 1 female)   .

Biology. Main hosts are white pine, western white pine, and ponderosa pine ( Pinus strobus   L., P. monticola D. Don.   and P. ponderosa Douglas ex C. Lawson   ). It is also found on a variety of other pines and Douglas fir ( Bright and Bouchard 2008).

Remarks. Two species of Pachyrhinus   , P. lateralis   and P. miscix   , have holotypes which are morphologically identical to confirmed specimens of P. elegans   . Pachyrhinus lateralis   ( Figs. 55-60 View FIGURES 55–60 ) was described from a single specimen taken at Lake County, California. It was differentiated from P. elegans   by Casey (1888) who cited a more transverse and depressed prothorax and tumid elytral humeri ( Fig. 55 View FIGURES 55–60 ). An examination of the holotype revealed the left side of the pronotum is cracked and dislocated, resulting in the appearance of a more depressed prothorax ( Fig. 56 View FIGURES 55–60 ). The elytral humeri were within the individual variation found in P. elegans   .

The holotype of Pachyrhinus miscix   ( Figs. 61-66 View FIGURES 61–66 ) is almost identical to that of P. lateralis   . Pachyrhinus miscix   was differentiated from P. elegans   by Fall (1901) citing a less arcuate front and middle tibia in the male ( Fig. 62 View FIGURES 61–66 ), a longer fourth funicular joint, and a more transverse thorax and hirsute abdomen ( Fig. 63 View FIGURES 61–66 ). Comparisons of the holotype and of other specimens identified as P. miscix   with P. elegans   indicate that these features fall within the range of individual variation for P. elegans   . The more arcuate tibia is a sexual feature common to the males of all Pachyrhinus species   and the relative sizes of the funicular joints were variable even within series of the material examined. The more hirsute abdomen is variable with females and more abraded specimens having less hirsute ventrites.

USNM

Smithsonian Institution, National Museum of Natural History

CNCI

Canadian National Collection Insects

UCRC

University of California, Riverside

CUIC

Cornell University Insect Collection

CSUC

California State University, Chico, Vertebrate Museum

CLR

All Saint's Church

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Curculionidae

Genus

Pachyrhinus

Loc

Pachyrhinus elegans (Couper 1865)

Benzel, Joseph & Bright, Donald E. 2021
2021
Loc

Pachyrhinus elegans: O’Brien & Wibmer and Wibmer 1982:43

Bright, D. E. & Bouchard, P. 2008: 159
Downie, N. M. & Arnett, R. H. 1996: 1485
McNamara, J. 1991: 330
O'Brien, C. W. & Wibmer, G. J. 1982: 43
1982
Loc

Pachyrhinus lateralis: O’Brien & Wibmer and Wibmer 1982: 43

O'Brien, C. W. & Wibmer, G. J. 1982: 43
1982
Loc

Pachyrhinus miscix: O’Brien & Wibmer and Wibmer 1982: 43

O'Brien, C. W. & Wibmer, G. J. 1982: 43
1982
Loc

Scythropus lateralis

Fall, H. C. 1901: 309
1901
Loc

Scythropus miscix

Fall, H. C. 1901: 309
1901
Loc

Scythropus elegans:

Furniss, R. L. & Carolin, V. M. 1977: 336
Hatch, M. H. 1971: 274
Blatchley, W. S. & Leng, C. W. 1916: 131
Fall, H. C. 1901: 309
Horn, G. H. 1876: 116
1876
Loc

Polydrusus elegans

Becker, E. C. 1974: 70
Couper, W. 1865: 63
1865