Cistus × cyprius var. pictus Pau & Font Quer

Nualart, Neus, Soriano, Ignasi, Prieto, David Pérez & Ibáñez, Neus, 2021, Catalogue and typification of the Moroccan taxa published by Carlos Pau, Phytotaxa 519 (1), pp. 1-94 : 28-52

Cistus × cyprius var. pictus Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 402 (1928), in sched., nom. nud.

= Cistus × cyprius Lam. [ Cistus ladanifer L. × C. laurifolius L.]

Note:—Hybrid between Cistus ladanifer var. salicifolius Mill. and C. laurifolius var. atlanticus Pit. published without a description.

2,3 Cistus ladanifer subsp. mauritianus Pau & Sennen in Sennen, Diagn. Nouv.: 178 (1936b)

Ind. loc.:—“Hab. Maroc: Djebel Kerker; dj. Tendri; Atlas Rifain à Targuist; Tizi-Ifri; Beni-Seddat, etc., 800–1500 m.”

Lectotype (designated by Soriano 2008: 30):—Beni-Bu-Yah djebel Kerker , 1000 m, 14 June 1933, Sennen 8671 & Mauricio (BC-Sennen 832209! [https://www.ibb.csic.es/herbari/ JPEG / BC832209 .jpg])

Notes:— The lectotype chosen by Soriano (2008) bears a handwritten label indicating djebel Kerker and also a label of Sennen’s exsiccata nº 8671 with a mention to all the localities indicated in the protologue. Other specimens bearing this exsiccata’s label exist, but they cannot be considered as isolectotypes, because (1) no specific locality is indicated ( BC 139542 !, BCN 14429 [photo!], BC-Sennen 832208!, MA 159621 !), or (2) they are from other localities cited in the protologue (BC-Sennen 832207!, MA 79684 !, MA 159622 !) .

1,3 Cistus ladanifer var. tangerinus Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 283 (1924a)

= Cistus ladanifer subsp. mauritianus Pau & Sennen

Ind. loc.:—“Yebel Ouebir, en sus laderas bajas y mirando al mar”

Holotype:— Tanger, Jebel Kebir , April 1921, Pau s.n. ( MA 79686 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00079686])

Note:—The specimen has three revision labels: C. ladanifer var. petiolatus Maire by Demoly in 1990, C. ladanifer subsp. africanus Dans. in 1993, and C. ladanifer subsp. mauritianus by Soriano in 1996 (all are synonyms).

1,2,3 Cistus riphienensis Pau in Bol. Soc. Iber. Ci. Nat. 31: 97 (1934), pro hybr.

=§ Cistus creticus subsp. eriocephalus (Viv.) Greuter & Burdet

Ind. loc.:—“KER-KER (Beni-Bu-Yahi), entre los 600 y 1400 m. Sennen y Mauricio ”

Lectotype (designated here):— Maroc: Beni-Bu-Yahi, dj. Kerker , 600–900 m, 14 June 1933, Sennen & Mauricio 8679 (lectotype MA 79282 ! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00079282]; isolectotypes BC-Sennen 832182!, BCN 39830 !)

Notes:—The nothotaxon was published as a hybrid between Cistus sennenianus Pau and C. villosus L. Original material was distributed in Sennen’s exsiccata Plantes d’Espagne 1933 and labelled as C. sennenianus . The selected lectotype was labelled also by Pau as Cistus villosus “ var. rifeniensis ”. The BC specimen was identified by Soriano in 1994 as C. creticus subsp. eriocephalus , although in APD (2020) the accepted name is C. creticus L. The subspecies of C. creticus differ by the absence or presence of glandular trichomes, but also by the shape, size, colour, and texture of the seeds ( Lo Bianco et al. 2017).

1,2,3 Cistus sennenianus Pau in Cavanillesia 5: 175 (1932)

= Cistus creticus subsp. eriocephalus (Viv.) Greuter & Burdet

Ind. loc.:—“este año en el Rif”

Lectotype (designated here):— Maroc: Beni-Bu-Yahi, Montes Idbiren , 21 June 1932, Sennen & Mauricio 8353 (lectotype MA 79281 ! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00079281]; isolectotypes BC 139560 !, BCN 39829 !, MA 159538 , MPU 008001 [photo!])

Notes:—The original material was distributed in Sennen’s exsiccata Plantes d’Espagne 1932. The BC specimen was identified as Cistus creticus subsp. eriocephalus by Soriano in 1995. The selected lectotype includes several original labels handwritten by Pau.

1,3 Convolvulus arvensis var. biflorus Pau in Font Quer, Iter Marocc. 1930 nº 525 (1932), in sched. = Convolvulus arvensis L. Ind. loc.:—“Hab. in arvis argillosis, pr. Rgaia (Beni Mzuar), ad 50 m. alt.; 4 julii” Lectotype (designated here):—In arvis argillosis, pr. Rgaia (Beni Mzuar), 50 m, 4 July 1930, Font Quer 525- 1930 (lectotype BC 98023! [https://www.ibb.csic.es/herbari/ JPEG / BC 98023.jpg]; isolectotypes BC 809008 !, BCN 73387 [photo!], GDA 5732 [photo!], MA 94345!, MPU 006711 [photo!], S 11-38117 [photo!]) Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimens were identified by Menemen in 1995 as C. arvensis . Among the two sheets of Herb. BC, the selected lectotype is the one with the most complete plants.

1,3 Convolvulus carmeli Pau & Sennen in Sennen, Pl. Espagne 1932 n° 8455 (1932–1933), in sched., nom. nud.

= Convolvulus valentinus Cav.

Notes:— Sennen & Mauricio (1934: 79) published this name as Convolvulus charmeli without indicating authors nor description. In APD (2020) this name is recorded as “ charmeli ”.

1,3 Convolvulus suffruticosus var. melillense Pau in Ann. Sci. Acad. Polytecn. Porto 6: 99 (1911)

= Convolvulus valentinus Cav.

Ind. loc.:—“ Melilla et Zeluan”

Holotype:— Melilla, ribazadas del río, April 1910, Pau s.n. ( MA 94101 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00094101])

Notes:— Carine & Robba (2010) and Wood et al. (2015) have indicated “ holotypes ” for this taxon.Two inaccuracies should be noted in Carine & Robba (2010): (1) the name of the taxon, cited as “ Convolvulus valentinus var. melillensis Pau ”, and (2) the transcription of the locality label, “Molilk ribanadas del rio” (which makes no sense) instead of “Melilla ribazadas del río” [=Melilla banks of the river]. Furthermore, Wood et al. (2015) indicated as holotype the specimen “ MOROCCO, Ain Tellout, Henry 6-462 (holotype MA!)”, previously referenced by Carine & Robba (2010) as the holotype of C. suffruticosus f. angustifolius Maire.

1,3 Convolvulus suffruticosus var. melliflorus Pau in Ann. Sci. Acad. Polytecn. Porto 6: 99 (1911)

≡ Convolvulus supinus var. melliflorus (Pau) Carine & Robba

Ind. loc.:—“Zeluan. (Abril, Mayo 1910)”

Lectotype (designated by Nualart et al. 2018):—Zeluan, 2 May 1910, Pau s.n. (lectotype MA 94100 ! [http:// colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00094100]; isolectotype MA 94102 !)

1,3 Convolvulus valentinus var. transfretanus Pau & Font Quer in Font Quer, Iter Marocc. 1929 nº 356 (1930a), in sched.

= Convolvulus valentinus Cav.

Ind. loc.:—“Hab. in collibus calc., pr. Hassi Berkan (Ulad Setut), 600 m. alt.; 30 majii”

Lectotype (designated here):—In collibus calc., pr. Hassi Berkan (Ulad Setut), 600 m, 30 May 1929, Font Quer 356-1929 (lectotype BC 98101 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98101 .jpg]; isolectotypes BCN 72509 [photo!], GDA 5794 [photo!], MA 94061 !, MPU 006661 [photo!], S 11-38447 [photo!])

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was revised by Menemen as C. valentinus . Sa’ad (1967) and Carine & Robba (2010) considered this variety as a nomen nudum. However, a description is included in the label and since the publication is earlier than 1953, the name is effectively published in schedulis (Art. 30.7, Turland et al. 2018).

1,2,3 Convolvulus vidalii Pau in Vidal, Bol. Soc. Esp. Hist. Nat. 21: 279 (1921)

Ind. loc.:—“Xauen, 14-V”

Lectotype (designated by Sa’ad 1967: 239):—Xauen, 14 May 1921, Vidal s.n. (lectotype MA 94375 ! [http:// colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00094375]; isolectotype BC 41894 !)

1,2,3 Coronilla arenivaga Pau in Font Quer, Iter Marocc. 1930 nº 363 (1932), in sched.

= Coronilla repanda (Poir.) Guss. subsp. repanda

Ind. loc.:—“Hab. in arenosis, pr. El Araix; 31 martii”

Lectotype (designated here):— In arenosis, pr. El Araix, 31 March 1930, Font Quer 363-1930 (lectotype BC

97990! [https://www.ibb.csic.es/herbari/JPEG/BC97990.jpg]; isolectotypes BCN 73130 [photo!], GDA 33599 [photo!], MA 68418!, MPU 006738 [photo!], S 10-2437 [photo!], Z 000022594 [photo!])

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The S and BC specimens were identified as C. repanda subsp. repanda by Lassen in 1984 and Díaz Lifante in 1996, respectively.

1 Cotyledon mucizonia f. glaberrima Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 327 (1924a)

= Sedum mucizonia (Ortega) Raym. -Hamet

Ind. loc.:—“Busemlal (M. Martínez); Samsa; Beni Hozmar”

Lectotype (designated here):—Samsa: Busemblal, May, Pau s.n. (MA 52031! [http://colecciones.rjb.csic.es/ #cardAdv.php?CatalogNumber=MA-01-00052031])

Notes:— Two syntypes are conserved in Herb. MA, one from Busemblal ( MA 52031 ) and another from Beni Hozmar ( MA 52030 !). The selected lectotype is the specimen with more samples. Both were identified as Sedum mucizonia by Velayos in 1995 .

1,3 Cotyledon mucizonia var. parviflora Pau in Font Quer, Iter Marocc. 1930 nº 268 (1932), in sched.

= Sedum mucizonia (Ortega) Raym. -Hamet

Ind. loc.:—“Hab. in rupibus umbrosis montis Bu-Zeitun (Beni Hosmar), ad 400 m. alt.; 15 junii”

Lectotype (designated here):—In rupibus umbrosis montis Bu-Zeitun (Beni Hosmar), 400 m, 15 June 1930, Font Quer 268-1930 (lectotype BC 830604 ! [https://www.ibb.csic.es/herbari/ JPEG / BC830604 .jpg]; isolectotypes BCN 73025 [photo!], GDA 27736 [photo!], MA 52036 !)

Notes:—The MA specimen was labelled as type by Byalt in 2005 and the GDA as syntype by Vizoso in 2013. The MA and BC specimens were identified as Sedum mucizonia by Velayos in 1995 and Rutherford in 1996, respectively.

1,3 Crepis taraxacifolia var. tangerina Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 351 (1924a)

=§ Crepis salzmannii Babc.

Ind. loc.:—“ Tánger ”

Lectotype (designated by Talavera et al. 2013: 237):— Tanger, April 1921, Pau s.n. (lectotype MA 140593! [http:// colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00140593]; isolectotype MA 140515 [photo!]*) Notes:— Talavera Solís et al. (2013) mention MA 140515 as isolectotype. Babcock (1947: 840) and APD (2020) synonymized the variety of Pau under Crepis vesicaria subsp. stellata (Ball) Babc. However, Talavera Solís et al. (2013) consider C. salzmannii as the accepted name based on the length of the beak and the achenes of the MA specimens. There is a BC specimen (BC 103263!) identified as Crepis vesicaria subsp. stellata by Babcock in 1941; in a label Babcock indicated that there were two types of achenes: the marginal ones of 4 mm long and not beaked, and the inner ones of 4. 5 mm long and attenuate into a short beak. Heteromorphic achenes are a diagnostic character for C. vesicaria subsp. stellata , therefore the BC sheet can not be considered as an isolectotype of C. salzmannii , because it belongs to a taxon different from that of the lectotype. It should be noted that Pau (1924a) described this variety only based on the glabrescent leaves and the farinose capitula.

1,3 Cynosurus echinatus var. tangerinus Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 398 (1924a)

= Cynosurus echinatus L.

Ind. loc.:—“ Tánger ”

Holotype:— Tanger, April 1921, Pau s.n. ( MA 10926 !)

Notes:—Pau labelled the specimen as “ var. breviaristatus Pau ”, but in the publication he used the name var. tangerinus with the description “ Glumae et glumerulae [sphalm. glumellulae] breviter aristatae ”.

1,3 Cytisus monspeliensis var. tangerinus Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 309 (1924a)

= Teline monspessulana ( L.) K.Koch

Ind. loc.:—“Yebel Ouebir”

Holotype:—Djebel Kebir, April 1921, Pau s.n. ( MA 60678 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00060678])

Note:—The specimen was identified as Teline monspessulana by Talavera in 1996.

1,2,3 Dactylis juncinella var. maroccana Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 60 (1928), in sched.

= Dactylis glomerata subsp. juncinella (Bory) Stebbins & D.Zohary

Ind. loc.:—“Hab. in cedretis montis Iguermalez (Atlante rhiphaeo), 1950 m. alt.”

Lectotype (designated here):—In cedretis montis Iguermalez (Atlante rhiphaeo), 1950 m, 20 July 1927, Font Quer 60-1927 (lectotype BC 68957 ! [https://www.ibb.csic.es/herbari/ JPEG / BC68957 .jpg]; isolectotypes GDA 30466 [photo!], MA 10650 !, BCN 15738 [photo!], BCN 73971 [photo!])

Notes:— The GDA specimen and the BCN 73971 specimen were labelled as syntypes, the first one by Vizoso in 2013. The BC specimen was identified as D. glomerata subsp. hispanica f. capitellata (Link) Maire & Weiller by Pyke in 2005; according to Maire (1955: 68) this form is a synonym of Dactylis juncinella Bory .

1,3 Danaa gigantea Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 330 (1924a)

≡ Petroselinum crispum subsp. giganteum (Pau) Dobignard

Ind. loc.:—“Peñascos de Gorgues, a 1.000 metros de altura”

Lectotype (designated here):—Beni Hozmar, peñascos de los Gorgues, 900– 100 m, 8 May 1921, Pau s.n. (lectotype MA 85946 ! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00085946]; isolectotype MA 85945 !)

Notes:—Previously typified by Dobignard (2009: 48) on a specimen collected by Font Quer in Beni Hozmar in 1930 (some years after the publication of this variety in 1924) and distributed in the exsiccata Iter Maroccanum 1930 nº 480 ( MPU 006724 [photo!] and MPU 006725 [photo!]). This typification has to be superseded according to Art. 9.19 ( Turland et al. 2018), because original material exists: plants collected in 1921 by Pau. The selected lectotype is a specimen with developed basal leaves and numerous flowers.

1,3 Daucus carota var. herculeus Pau ex Maire in Bull. Soc. Hist. Nat. Afrique N. 24(7): 214 (1933)

= Daucus carota subsp. maximus (Desf.) Ball

Ind. loc.:—“Hab. in arvis argillosis Imperii Maroccani occidentalis prope Lixum (Font Quer)”

Lectotype (designated by Martínez-Flores & Crespo 2019: 1362):— Pr. El Araix, l. Txumix, 40 m, fl. 29 May 1930, fruct. 22 June 1930, Font Quer 494-1930 (lectotype BC 98160 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98160 .jpg]; isolectotypes BCN 73359 [photo!], GDA 40447 [photo!], MA 89134 !, MPU 006719 [photo!], MPU 006720 [photo!], MPU 006721 [photo!], S 07-7174 [photo!], S-G 10216 [photo!], Z 000028669 [photo!], Z 000028670 [photo!])

Notes:—Pau proposed this name at the species rank as “ Daucus herculeus Pau ” in the specimen distributed in Iter Maroccanum 1930 nº 494. However, the indication in the label “ D. Gingidio affinis ” does not meet the requirement of Art. 38.1(a) for a “description or diagnosis” ( Turland et al. 2018), and therefore this name should be considered a nomen nudum. Some years later, Maire (1933) provided a description to this taxon and subordinated it to D. carota subsp. maximus in rank of variety. It should be noted that the locality indicated by Maire “Lixum” is a site of an ancient city located in Tchemmich hill (variant name of “Txumix” used by Font Quer in the label). The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as D. carota subsp. maximus by Pyke in 2015.

3 Daucus gingidium var. africanus Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 462 (1928), in sched.

= Daucus carota subsp. hispanicus (Gouan) Thell.

Ind. loc.:—“ Hab. in rupibus maritimis Marsa Quebira (Bocoya), 20 m. alt.; fl. 19 majii, fruct. 23 julii” Lectotype (designated here):—In rupibus maritimis Marsa Quebira (Bocoya), 20 m, fl. 19 May 1927, fruct. 23 July 1927, Font Quer 462-1927 (lectotype BC 24351 ! [https://www.ibb.csic.es/herbari/ JPEG / BC24351 .jpg]; isolectotypes BCN 18038 [photo!], GDA 40444 [photo!], MA 89122 !, MPU 006280 [photo!], MPU 006281 [photo!])

Notes:—The GDA and BCN specimens were labelled as syntypes, the first one by Vizoso in 2013. The MPU 006281 specimen was identified as D. carota subsp. gummifer (Syme) Hook.f. by Okeke in 1976, and the BC specimen as D. carota subsp. hispanicus by Pyke in 2015 (both names are synonyms).

1,3 Daucus minusculus Pau in Bol. Soc. Esp. Hist. Nat. 14: 428 (1914)

≡ Pseudorlaya minuscula (Pau) M.Laínz

Ind. loc.:—“alrededores de Larache … Sr. Pérez Camarero, durante la primavera pasada [1914]”

Lectotype (designated by Sáenz de Rivas 1974: 194):—Larache, 1914, Pérez Camarero 31 (MA 89307! [http:// colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00089307])

Notes:—The specimen was labelled as holotype by Sáenz de Rivas in 1974, who also identified it as Pseudorlaya minuscula . Sáenz de Rivas (1974) and APD (2020) cite this taxon as “Pau ex Font Quer”, but the description was made only by Pau although the author of the paper is Font Quer, as indicated in Nualart et al. (2018).

1 Delphinium pentagynum f. exagynum Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 273 (1924a), nom. nud.

= Delphinium pentagynum Lam.

Notes:—There is no description in the protologue, as already indicated by Carrasco (1975).

1 Deschampsia flexuosa [unranked] minorifolia Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 42 (1928), in sched., nom. nud.

= Deschampsia flexuosa (L.) Trin.

1 Deschampsia flexuosa [unranked] rigidifolia Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 41 (1928), in sched., nom. nud.

= Deschampsia flexuosa (L.) Trin.

1,2,3 Deschampsia stricta var. longiseta Pau & Font Quer in Font Quer, Mem. Acad. Ci. Artes Barcelona 22(18): 345

(1931b)

=§ Deschampsia flexuosa (L.) Trin. subsp. flexuosa

Ind. loc.:—“Djebel-er-rgel, de Beni Mezouar, cerca de Rgaia, en las laderas síliceas de este monte y a 500 m de alt.”

Lectotype (designated here):— In collibus arenaceis montis Dj. Er-Rgel, c. R’gaia (Beni Mezouar), 500 m, 5 July 1930, Font Quer 45-1930 (lectotype BC 136375 ! [https://www.ibb.csic.es/herbari/ JPEG / BC136375 .jpg]; isolectotypes B 10 0167336 [photo!]*, BCN 72671 !, GDA 30482 [photo!], MA 144931 !, MPU 006787 [photo!], SANT 7143 [photo!])

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as D. flexuosa by Ruiz in 1995, and the MA specimen as D. stricta Hack. by Buschmann in 1950 (both names are synonyms). El Oualidi et al. (2012) mention the B specimen as type. According to APD (2020) the accepted name is D. flexuosa subsp. iberica Rivas Mart. , but other authors consider this subspecies endemic to the Central Iberian mountain range and accept for Morocco only the typical subspecies ( Fennane 2014e). According to the description of both subspecies by Rivas Martínez (1964) and based on examination of the lectotype, this variety should be synonymized under D. flexuosa subsp. flexuosa due to the scabrous ligules and the absence of milky white innovation leaves.

1,2,3 Dianthus brachyanthus var. maroccanus Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 197 (1928), in sched.

= Dianthus brachyanthus Boiss.

Ind. loc.:—“Hab. in lapidosis cacuminis montis Yebel Lerz d. (Atlante rhiphaeo), solo siliceo, 2200 m. alt., 12 junii”

Lectotype (designated here):—In lapidosis cacuminis montis Yebel Lerz d. (Atlante rhiphaeo), 2200 m, 12 June 1927, Font Quer 197-1927 (lectotype BC 9731 ![https://www.ibb.csic.es/herbari/ JPEG / BC9731 .jpg]; isolectotypes B 10 1180138 [photo!], BCN 17294 [photo!], G 00424974 [photo!], GDA 1738 [photo!], MA 33786 !, MPU 006341 [photo!])

Notes:— The GDA specimen was labelled as syntype by Vizoso in 2013 and the B specimen as isotype by Vogt & Bernhard in 2018. The BC specimen was identified as D. brachyanthus by Romo in 1996 .

1 Dianthus laricifolius f. africanus Pau in Bol. Soc. Ibér. Ci. Nat. 31: 99 (1934), nom. nud.

= Dianthus lusitanus Brot.

Notes:— Pau (1934) cites this form, without a description, from Ker-Ker collected by Sennen and Mauricio in 14 June 1933. The specimen MA 33455 [photo!] matches this data and is handwritten by Pau as var. africanus . Its woody base is a character of D. lusitanus .

Dianthus laricifolius f. maroccanus Pau & Sennen in Sennen, Pl. Espagne 1933 nº 8713 (1933–1934), in sched., nom. nud.

= Dianthus anticarius Boiss. & Reut.

Notes:—The BC specimen corresponding to Sennen’s exsiccata Plantes d’Espagne 1933 nº 8713 (BC 139277!) was identified as D. anticarius by Romo in 1995. On the label, there is neither description nor diagnosis, only the indication of a new form for Morocco.

1 Dianthus ripheniensis Pau , in sched. (MA 33510 [photo!] [http://colecciones.rjb.csic.es/#cardAdv.

php?CatalogNumber=MA-01-00033510]), nom. nud.

= Dianthus sylvestris subsp. longibracteatus (Maire) Greuter & Burdet

Notes:—The name was handwritten by Pau in a label of a specimen collected by Sennen and Mauricio in 14 June 1933 from Ker-Ker. The shape and size of calyx and epicalyx match those of D. sylvestris subsp. longibracteatus .

1,2,3 Dianthus siculus var. lanceolatus Pau in Caballero, Trab. Mus. Nac. Ci. Nat., Ser. Bot. 11: 22 (1917)

=§ Dianthus sylvestris subsp. longibracteatus (Maire) Greuter & Burdet

Ind. loc.:—“Habitat in Monte Gurugú. Junio fl.”

Holotype:—Gurugú (Riff), June 1915, Caballero s.n. ( MA 33506 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00033506])

Notes:—The D. sylvestris -group is one of the most complex within the genus, and its treatment still remains unclear. Ibn Tattou (1999b) recognises three subspecies in Morocco, namely D. sylvestris subsp. longicaulis (Ten.) Greuter & Burdet, subsp. longibracteatus , and subsp. siculus (C.Presl) Tutin , which differ from each other by the length of the epicalyx bracts, whereas Romo (2002b) considers D. siculus C.Presl a synonym of the subsp. longicaulis . APD (2020) synonymized var. lanceolatus under D. sylvestris subsp. siculus , but according to Romo (2002b) it should be synonymized under subsp. longibracteatus . A study of the type specimen has demonstrated that the main diagnostic characteristics (epicalyx bracts narrow and long) do not match those of the Italian populations of D. siculus (epicalyx bracts elliptical to obovate) and those of D. longicaulis Ten. (epicalyx bracts transversally elliptical to rounded) studied by Bacchetta et al. (2010). They rather correspond to the Moroccan endemic D. sylvestris subsp. longibracteatus (epicalyx bracts lanceolate to ovato-lanceolate, cf. Romo 2002b).

1,2,3 Digitalis obscura var. riphaea Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 584 (1928), in sched.

≡ Digitalis obscura subsp. riphaea (Pau & Font Quer) Valdés & Mateos

Ind. loc.:—“Hab. in declivibus calc. montis Yebel Hammam (Bocoya), 100 m. alt.; 4 majii”

Lectotype (designated here):—In declivibus calc. montis Yebel Hammam (Bocoya), 100 m, 4 May 1927, Font Quer 584-1927 (lectotype BC 43877! [https://www.ibb.csic.es/herbari/ JPEG / BC 43877.jpg]; isolectotypes BCN 70871 [photo!], GDA 39252 [photo!], MA 112836!, MPU 006382 [photo!], MPU 006383 [photo!]) Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as D. laciniata subsp. riphaea (Pau & Font Quer) Romo by Romo in 1995. Although Romo (2002c) and Ibn Tattou (2007b) subordinate this taxon, as subspecies, to D. laciniata Lindl., APD (2020) considers it as subspecies of D. obscura L.

1,3 Diplotaxis catholica var. maroccana Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 278 (1924a), nom. nov. pro Diplotaxis siifolia var. bipinnatifida Coss. in Ill. Fl. Atlant. 1: 40 (1882)

≡§ Diplotaxis siifolia subsp. bipinnatifida (Coss.) Mart. -Laborde

Ind. loc.:—“ MAR. In provinciis austro-occidentalibus: Chtouka, Dar-ould-Delimi, Assaka, Oudjan, Dj. Tizelmi, Tazeroualt (Mard.)”

Lectotype (designated here):—Dar Ould Delimi, 1875, Mardoché s.n. (lectotype P 05355459 [photo!]* [https:// science.mnhn.fr/institution/mnhn/collection/p/item/p05355459];isolectotype MPU 100739 [photo!]*, P05355420 [photo!]*)

Notes:— Pau (1924a) proposed this name for Diplotaxis siifolia var. bipinnatifida as a variety under D. catholica ( L.) DC., since there existed a taxon described by Kunze with the same name. Martínez-Laborde (1992: 242) indicated two possible types for the variety described by Cosson (1882), at Herbs. MPU and P, but did not select a lectotype. Moreover, in Herb. P, which conserves the main collection of Cosson ( Stafleu & Cowan 1976), there are two specimens. Between these two specimens, the one with leaves best conserved is chosen here as the lectotype. In APD (2020) the variety of Pau (1924a) is included within D. catholica , but it is indeed the avowed substitute for Diplotaxis siifolia subsp. bipinnatifida (replaced synonym), which is the name accepted in that database and should be treated accordingly.

1,2,3 Dryopteris rigida var. rhiphaea Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 3 (1928), in sched.

= Athyrium filix-femina ( L.) Roth

Ind. loc.:—“Hab. pr. Zarkat (Atlante rhiphaeo), ad rivulos, 1400 m. alt.; 11 junii”

Lectotype (designated here):— Pr. Zarkat (Atlante rhiphaeo), ad rivulos, 1400 m, 11 June 1927, Font Quer 3-1927 (lectotype BC 72531 ! [https://www.ibb.csic.es/herbari/ JPEG / BC72531 .jpg]; isolectotypes BCN 15685 [photo!], MA 327 !, MPU 006362 [photo!])

Note:—The BC and MA specimens were identified as Athyrium filix-femina by Jury in 1996 and Rivas Martínez 1963, respectively.

1,2,3 Echinops fontqueri Pau in Font Quer, Iter Marocc. 1928 nº 409 (1929), in sched.

Ind. loc.:—“Hab. in incultis, pr. Xauen, 450 m. alt.; 10 junii”

Lectotype (designated by Valdés 1996a: 306):—In incultis, pr. Xauen, 450 m, 10 June 1928, Font Quer 409-1928 (lectotype BC 32791 ! [https://www.ibb.csic.es/herbari/ JPEG / BC32791 .jpg]; isolectotypes BCN 19406 [photo!], EMMA 17728 [photo!], GDA 4411 [photo!], MA 131169 !, MPU 006598 [photo!], MPU 006597 [photo!], G 00023520 [photo!], Z 000003303 [photo!])

Notes:—The GDA specimen was labelled as isolectotype by Vizoso in 2015. Determination of the BC and GDA specimens was confirmed by Ortega in 2005 and 2006, respectively.

1,2,3 Echium melillense Pau in Bol. Soc. Esp. Hist. Nat. 24: 100 (1924b)

= Echium horridum Batt.

Ind. loc.:—“Rif Oriental ”

Holotype:— Riff Oriental, September 1920, Vicioso s.n. ( MA 97400 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00097400])

Note:—The specimen was identified as E. horridum by Valdés in 2007 and with the indication of “Tipo” in the original label.

1,2,3 Echium pomponium var. tangerinum Pau in Bol. Soc. Esp. Hist. Nat. 24: 100 (1924b), nom. nud.

= Echium boissieri Steud.

Note:—In the protologue there is only an indication by Lacaita in litt. as “ forma extrema ”.

1,2,3 Echium pomponium var. velutinum Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 364 (1924a)

= Echium boissieri Steud.

Ind. loc.:—“Laucien”

Holotype:—Laucien, 3 May 1921, Pau s.n. ( MA 96889 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00096889])

Note:—In the original label Pau indicates “ var. tangerinum = velutinum ”. Specimen identified as E. boissieri by Valdés in 2006.

1,2,3 Echium rifeum Pau in Caballero, Trab. Mus. Nac. Ci. Nat., Ser. Bot. 11: 27 (1917)

≡ Echium sabulicola subsp. rifeum (Pau) Valdés

Ind. loc.:—“en la cima del Monte Gurugú, bastante copiosa, alrededor de Hardú”

Holotype:—Gurugú (Riff) circa Hardú, 7 June 1917 [sphalm. for 1915], Caballero s.n. ( MA 97395 ! [http:// colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00097395])

Notes:—The specimen was identified as E. sabulicola subsp. rifeum by Valdés in 2007. The name of Pau published in Caballero (1917) was previously typified by Klotz (1962: 1091) on a MA specimen collected by Font Quer in mount Gurugú in 1929 (some years after its publication in 1917) and distributed in the Iter Maroccanum 1929 nº 361; nevertheless, this typification has to be superseded according to Art. 9.19 ( Turland et al. 2018), because original material exists. It should be noted that there is a possible confusion in the date of the type sheet, which bears two original labels, the first by Caballero indicating “7 junii” and the other by Pau specifying “7 junii 1917”. However, Caballero visited Morocco only in 1912, 1915, 1923, and 1934 (see González-Bueno & Gomis 2007), and the publication of the taxon states that all the specimens were collected in 1915.

1,2,3 Echium sennenii Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 363 (1924a)

= Echium plantagineum L.

Ind. loc.:—“ Tánger ”

Holotype:— Tanger, April 1921, Pau s.n. ( MA 97390 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00097390])

Note:—The specimen was identified as E. plantagineum by Valdés in 2006.

1,2,3 Elaeoselinum foetidum var. brachylobum Pau in Font Quer, Iter Marocc. 1930 nº 492 (1932), in sched.

= Elaeoselinum foetidum ( L.) Boiss.

Ind. loc.:—“Hab. in quercetis suberis inter El Araix et El Ksar-el-Quebir”

Lectotype (designated by García Martín & Silvestre 1985: 222):— In quercetis suberis inter El Araix et El Ksar-el-Quebir, 24 June 1930, Font Quer 492-1930 (lectotype MA 88815 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00088815]; isolectotypes BC 98158 !, BCN 73360 [photo!], GDA 40469 [photo!], MA 88814 !, MAF 53374 , MPU 006722 [photo!], MPU 006723 [photo!])

Note:— The GDA specimen was labelled as isolectotype by Vizoso in 2015, and MA 88815 as lectotype .

1 Elaeoselinum foetidum f. hydrofila Pau in Vidal, Asoc. Esp. Progr. Ci. Congr. Cádiz 6: 147 (1928), nom. nud.

= Elaeoselinum foetidum (L.) Boiss.

Note:—In the protologue, there is only an indication that this form could be a new or different species.

1,3 Epilobium caballeroi Pau in Vidal, Bol. Soc. Esp. Hist. Nat. 22: 59 (1922), pro hybr.

= Epilobium parviflorum Schreb.

Ind. loc.:—“Xauen”

Type:—not seen

Notes:—This name was proposed by Pau in Vidal (1922) for a hybrid between Epilobium hirsutum L. and E. tournefortii Michal. Only one specimen from Xauen, identified as E. tournefortii ( MA 84193!), was found, but no original material of this hybrid. Raven (1967) cite a photograph of the type material kept at Herb. DS (now transferred to Herb. CAS), but it has not been located (comm. pers. D. Trock).

1,2,3 Erinus alpinus var. macranthus Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 372 (1929), in sched.

= Erinus alpinus L.

Ind. loc.:—“Hab. in rupibus calc. montis Magot (Gomara), 1300 m. alt.”

Lectotype (designated here):—In rupibus calc. montis Magot (Gomara), 1300 m, 21 May 1928, Font Quer 372- 1928 (lectotype BC 44950 ! [https://www.ibb.csic.es/herbari/ JPEG / BC44950 .jpg]; isolectotypes BCN 18144 [photo!], MPU 006605 [photo!])

Note:—Identification of the BC specimen was confirmed by Benedí in 1996.

1 Erodium baeticum Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 302 (1924a), pro hybr.

= Erodium cicutarium ( L.) L’Hér.

Ind. loc.:—“ Tánger, un solo pie”

Lectotype (designated here):— Tanger, April 1921, Pau s.n. (lectotype MA 72366 ! [http://colecciones.rjb.csic. es/#cardAdv.php?CatalogNumber=MA-01-00072366]; isolectotype MA 72274 !)

Notes:— Pau (1924a) proposed this new nothotaxon name as a hybrid between Erodium moschatum ( L.) L’Hér. and E. praecox (Cav.) Willd. The sheet MA 72366!, originally labelled by Pau as E. baeticum , is here selected as the lectotype. It should be noted that the sheet MA 72274 has a revision label by Aldasoro in 1995 as E. cicutarium , with an indication of “ lectotypus E. baeticum Pau ”. However, on the original label, Pau only indicated “ Erodium cicutarium L’Hér (auct.?)”, and the typification of Aldasoro was apparently not published.

3 Erodium chaerophyllum f. latisectum Pau in Font Quer, Iter Marocc. 1930 nº 399 (1932), in sched., nom. nud.

=§ Erodium salzmannii Delile

Note:—In APD (2020) this form appears under E. cicutarium ( L.) L’Hér., but plants distributed in Iter Maroccanum 1930 nº 399 have whitish pits, a character of E. salzmannii .

1 Erodium chium f. macrophyllum Pau in Cavanillesia 1: 140 (1929a)

= Erodium chium ( L.) Willd.

Ind. loc.:—“Larache ( MAS)”

Holotype:—Larache, 27–29 April 1928, Mas Guindal ( MA 71891 [photo!] [http://colecciones.rjb.csic.es/ #cardAdv.php?CatalogNumber=MA-01-00071891])

Notes:—The name was previously published by Pau (1924a: 301), but, according to Navarro (2015), it should be considered invalid, since the plant described by Pau (1924a) was obtained by cultivation from seeds of the typical form of E. chium . Some years later, Pau (1929a) provided a valid diagnosis. The holotype is identified as Geranium malacoides L. following Cavanilles (1787: tab. XCI), as indicated in Pau (1924a). The accepted name follows Navarro (2015).

1,2,3 Erodium macradenum var. heteradenum Pau & Font Quer in Font Quer, Cavanillesia 4: 91 (1931c)

≡ Erodium heteradenum (Pau & Font Quer) Guitt.

Ind. loc.:—“ Hab. in rupibus calc. umbrosisque montis Bu-Zeitun (Beni Hosmar), ad 1000 m. alt.; 14 junii” Lectotype (designated here):—in rupibus calc. umbrosisque montis Bu-Zeitun (Beni Hosmar), 1000 m, 14 June 1930, Font Quer 401-1930 (lectotype BC 98271 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98271 .jpg]; isolectotypes BCN20123 !, G 00439862 [photo!], MA 72065 !, MPU 007147 [photo!])

Notes:—On the original label of the Iter Maroccanum exsiccata there is the indication of “Cotypus” (= syntype). The MA specimen was labelled as lectotype by Aldasoro in 1999, but this typification was not published. The BC specimen bears two revision labels: one as E. heteradenum by Rutherford in 1996, and another as E. antariense Rouy by Aldasoro in 2010. According to Jury & Rutherford (2002a) and Ibn Tattou (2007c), these two taxa differ by the size of mericarps and beaks, and the density of the cilia on the outer edge of the pits. Based on the present study of these characters the identification as E. heteradenum is confirmed.

1,2,3 Erodium masguindalii Pau in Cavanillesia 2: 88 (1929c)

Ind. loc.:—“Frecuente en el Dersa, según el Sr. MAS ”

Holotype:—Dersa, 13 April 1929, Mas Guindal s.n. ( MA 72368 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00072368])

Notes:—The specimen was identified by Aldasoro, Aedo, and Navarro in 2001 as E. acaule ( L.) Bech. & Thell. This species of Pau is accepted as endemic to Morocco by Jury & Rutherford (2002a), Ibn Tattou (2007c), and APD (2020), although Aldasoro et al. (2009) consider it to be a synonym of E. acaule .

1,2,3 Erucastrum minutiflorum Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 230 (1928), in sched.

= Brassica tournefortii Gouan

Ind. loc.:—“Hab. in Littore rhiphaeo ad agrorum margines pr. Marsa Saguira (Bocoya)”

Lectotype (designated here):—In Littore rhiphaeo ad agrorum margines pr. Marsa Saguira (Bocoya), 15 May 1927, Font Quer 230-1927 (lectotype BC 4018 ![https://www.ibb.csic.es/herbari/ JPEG / BC4018 .jpg]; isolectotypes BCN 16379 [photo!], GDA 28226 [photo!], MA 46477 !, MPU 006333 [photo!])

Note:—The BC and MA specimens were identified as Brassica tournefortii by Gómez Campo in 1996 and 1979, respectively.

1,2,3 Eryngium caespitiferum Font Quer & Pau in Font Quer, Cavanillesia 4: 30 (1931a)

Ind. loc.:—“Hab. in cacumine montis Lexhab (Gomara), ad 2100 m.”

Lectotype (designated by Wörz 2011: 189, second-step designated here):—In cacumine montis Lexhab (Gomara), 2100 m, 21 July 1930, Font Quer 465-1930 (lectotype BC 98522 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98522 .jpg]; isolectotypes BC 809318 !, BCN 20126 [photo!], GDA 40484 [photo!], MA 84839 !*, MA 84840 !*, MPU 007143 [photo!]*, Z 000028671 [photo!])

Notes:— Wörz (2011) lectotypified this taxon on a BC specimen, but without mention to any particular sheet ( MA, MPU, and Z specimens are cited as isolectotypes). In BC there are two sheets of this gathering with different barcode numbers so a second-step lectotype is here designated under Art. 9.17 ( Turland et al. 2018); BC 98522 is selected, because it conserves more samples. The MA specimen was labelled as isotype by Jury in 1995, and the GDA specimen as syntype by Vizoso in 2013. The BC specimens were identified as E. huteri Porta by Jury in 1996, since Jury (1996) considers E. caespitiferum to be conspecific with E. huteri . According to Nieto Feliner (2001) E. huteri is an Iberian endemic plant while E. caespitiferum a Moroccan endemic; the characters that distinguish both species are leaf and bract venation, leaf division and colour, and bract length and margin.

1,2,3 Eryngium × mohamedanii Font Quer & Pau in Font Quer, Cavanillesia 4: 31 (1931a) [ Eryngium bourgatii Gouan × E. caespitiferum Font Quer & Pau ]

Ind. loc.:—“ Hab. in cacumine montis Lexhab (Gomara-Imp. Maroc), inter parentes, ad 2000–2100 m. alt., ubi, d. 22 julii 1930 legi ”

Holotype:— s.l., s.d., s.c. (holotype BC 98523 [center specimen!] [https://www.ibb.csic.es/herbari/ JPEG / BC98523 .jpg]; isotypes BC 98523 [two upper-left specimens!])

Notes:—New hybrid originally published as E. bourgatii var. hispanicum Lange × E. caespitiferum . Wörz (2011) selected the specimen BC 98523 as lectotype, but this typification is not needed, taking into account that there is a photograph accompanying the description that is an unequivocal indication of the type specimen according to the authors. One of the samples of this BC specimen matches this photograph, therefore it should be considered as holotype. As this voucher bears more samples, other samples should be considered as isotypes. The BC specimen was labelled as type by Jury in 1995. Other specimens were distributed in Iter Maroccanum 1930 nº 466, but they are not considered isotypes, because the date does not match the one indicated in the protologue (21 July instead of 22 July).

1,2,3 Eryngium triquetrum var. xauense Pau in Vidal, Bol. Soc. Esp. Hist. Nat. 21: 277 (1921)

≡ Eryngium triquetrum subsp. xauense (Pau) Jovet & Sauvage

Ind. loc.:—“Xauen”

Lectotype (designated by Breton 1962: 56):— s.l., 15 May 1921, s.c. ( MA 84914 ! [http://colecciones.rjb.csic. es/#cardAdv.php?CatalogNumber=MA-01-00084914])

1,3 Erysimum incanum var. longipes Pau in Broteria Ser. Ci. Nat. 2(29): 45 (1933)

= Erysimum incanum Kunze subsp. incanum

Ind. loc.:—“muestra repartida el año 1925 por Jahandiez (Plantas marocc, n.° 190)”

Lectotype (designated here):— Moyen Atlas : Itzer, Vallée de l’Oued Bou Haffs, 1900 m, 6 May 1925, Jahandiez 190 (lectotype MA 48758 ! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00048758]; isolectotypes MPU 102628 [photo!])

Notes:—The specimen was distributed by Jahandiez and originally labelled as Erysimum kunzeanum Boiss. & Reut.

1,2,3 Erysimum kuntzeanum var. maroccanum Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 141 (1929), in sched.

=§ Erysimum incanum Kunze subsp. incanum

Ind. loc.:—“Hab. in saxosis calc. montis Kalaa, 1100 m. alt.”

Lectotype (designated here):—In saxosis calc. montis Kalaa, 1100 m, 25 May 1928, Font Quer 141-1928 (lectotype BC 3505 ! [https://www.ibb.csic.es/herbari/ JPEG / BC3505 .jpg]; isolectotypes BCN 17864 [photo!], G 00414632 [photo!], GDA 28252 [photo!], MA 48762 !, MPU 006420 [photo!])

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013 and the MA specimen as type by Nieto Feliner in 1991, who identified it as E. incanum subsp. incanum . APD (2020) includes this taxon in E. incanum although different subspecies are accepted there; therefore, the name accepted here follows Montserrat (2002e).

1,2,3 Euphorbia bivonae var. tangerina Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 384 (1924a)

= Euphorbia squamigera Loisel.

Ind. loc.:—“Peñascos antes de llegar al Cabo Espartel. Rarísima”

Holotype:—Cabo Espartel, peñascos, 24April 1921, Pau s.n. ( MA 75011 ![http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00075011])

Note:—The specimen was identified as E. squamigera by Vicens in 1993.

1,3 Euphorbia clementei f. hozmariensis Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 383 (1924a)

= Euphorbia clementei Boiss. subsp. clementei

Ind. loc.:—“Beni Hozmar”

Lectotype (designated by Simon & Vicens 1999: 588):—Beni Hosmar, May 1921, Pau s.n. ( MA 74972 ! [http:// colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00074972])

Notes:—The specimen was labelled as lectotype by Vicens in 1993, who also identified it as E. clementei . According to Simon & Vicens (1999) this form should be attributed to subsp. clementei .

1,2,3 Euphorbia dracunculoides subsp. melillensis Pau ex Maire in Bull. Soc. Hist. Nat. Afrique N. 26(7): 228 (1935) = Euphorbia medicaginea Boiss.

Ind. loc.:—“Hab. in lapidosis aridis ditionis Rifanae prope Zaio! ( SENNEN et MAURICIO)”

Holotype:— El Zaio, coteaux, 23 April 1931, Mauricio & Sennen ( MPU 188136 [photo!] [https://explore.recolnat. org/occurrence/ 6828C95DFA634FE1B2479B0A54B4897E])

Note:— Sennen & Mauricio (1934) also published the name “ Euphorbia melillensis Nob. ” from Hidum without description. Maire (1935) reported “ subsp. melillensis (Pau ined. pro specie)”, but no reference of this species of Pau was found.

1,2,3 Euphorbia rugosissima Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 255 (1929), in sched.

= Euphorbia phymatosperma subsp. cernua (Coss. & Durieu ex Boiss.) Vindt

Ind. loc.:—“Hab. in saxosis calc. montis Kalaa, 950 m. alt.”

Lectotype (designated here):—In saxosis calc. montis Kalaa, 950 m, 11 May 1928, Font Quer 255-1928 (lectotype BC 57611 ! [https://www.ibb.csic.es/herbari/ JPEG / BC57611 .jpg]; isolectotypes BCN 17875 [photo!], BM 001050424 [photo!], GDA 29313 [photo!], MA 75186 !, MA 75188 !, MPU 008247 [photo!], Z 000017118 [photo!])

Notes:—The BC and BM specimens were identified as E. phymatosperma subsp. cernua by Molero in 1988, and by Berry and Peirson in 2012, respectively. The MPU specimen was labelled as syntype by Mathez in 1992, the BM specimen as type by Berry and Peirson in 2012, and the GDA specimen as syntype by Vizoso in 2013.

1,2,3 Euphorbia tetuanensis Pau in Monde Pl. 30(66–181): 1 (1929b)

= Euphorbia clementei Boiss. subsp. clementei

Ind. loc.:—“Los Dersas in Tetuan ( Mas Guindal )”

Lectotype (designated by Simon & Vicens 1999: 624):—Al pie del Dersa (Tetuan), 12 April 1929, Mas Guindal s.n. ( MA 205851! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00205851]) Note:—The specimen was labelled as lectotype by Vicens in 1993, who also identified it as E. clementei without indicating subspecies name; later, Simon & Vicens (1999) synonymized this taxon under subsp. clementei .

1,2,3 Evax pygmaea var. gaditana Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 340 (1924a)

≡§ Filago gaditana (Pau) Andrés-Sánchez & Galbany

Ind. loc.:—“Yebel Quebir … Esta misma forma la poseía en mi herbario de Casas Viejas (Cádiz)”

Lectotype (designated by Andrés-Sánchez et al. 2011: 573):—Djebel Kebir, April 1921, Pau s.n. (lectotype MA 123865 ; isolectotype BC 32432 !)

Notes:— Andrés-Sánchez et al. (2011) mention a specimen from “Casas Viejas” as syntype ( MA 123956). The BC specimen was labelled also as syntype by Andrés-Sánchez in 2010, who identified it as Filago gaditana . APD (2020) accepts this taxon at the subspecies rank [ Filago pygmaea subsp. gaditana (Pau) Dobignard ], but according to Andrés-Sánchez et al. (2011) it should be considered a segregate species.

1,3 Evax pygmaea var. virescens Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 648 (1928), in sched.

= Filago pygmaea L.

Ind. loc.:—“Hab. in aridis, supra Targuist, 1100 m. alt., solo arenaceo”

Lectotype (designated here):—In aridis, supra Targuist, 1100 m, 23 May 1927, Font Quer 648-1927 (lectotype BC 32431 ! [https://www.ibb.csic.es/herbari/ JPEG / BC32431 .jpg]; isolectotypes BCN 18027 [photo!], GDA 4478 [photo!], MPU 006374 [photo!])

Notes:—The GDA specimen was labelled as isotype by Andrés-Sánchez in 2010, and as syntype by Vizoso in 2015. The BC specimen was identified as Evax pygmaea ( L.) Brot. by Devesa in 1996, and the GDA one as Filago pygmaea by Andrés-Sánchez in 2010.

1,2,3 Ferula bolivari Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 332 (1924a)

= Ferula tingitana L.

Ind. loc.:—“Beni Hozmar, en los peñascos; altura, 1.000 metros”

Lectotype (designated here):—Beni Hozmar, 1000 m, May 1921, Pau s.n. (lectotype MA 88424! [http:// colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00088424]; isolectotype MA 88426!) Notes:—There are two sheets at MA from Beni Hozmar; the lectotype is labelled as F. bolivari and the isolectotype as F. communis L., both by Pau. But according to the Pau (1924a) himself, plants from Beni Hosmar should be referred to F. bolivari , not to F. communis .

1,3 Ficaria verna var. africana Pau in Font Quer, Iter Marocc. 1929 nº 159 (1930a), in sched.

= Ficaria verna subsp. grandiflora (Robert) Hayek

Ind. loc.:—“Hab. in humidis montis Djebel Tersetx (Beni Seddat), 1750 m. alt.; 23 mart”

Lectotype (designated here):—In humidis montis Djebel Tersetx (Beni Seddat), 1750 m, 23 March 1929, Font Quer 159-1929 (lectotype BC 98210 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98210 .jpg]; isolectotypes BCN 72349 [photo!], GDA 35282 [photo!], MA 40041 !, MPU 006692 [photo!])

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013, and the MA as type by Aedo in 2006. Veldkamp (2015) cites the MA specimen as holotype, but since different specimens considered original material are extant, the designation of a lectotype is necessary.

1 Ficaria verna f. disjunta Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 272 (1924a)

= Ficaria verna subsp. grandiflora (Robert) Hayek

Ind. loc.:—“Crece al pie de Beni Hozmar”

Holotype:—Beni Hozmar, May 1921, Pau s.n. ( MA 40045 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00040045])

Note:—The specimen was originally labelled by Pau as “ F. verna var. grandiflora f. divergens Pau ”. It should be noted that the main diagnostic feature of the variety is the divergent lobes of the leaf base.

1,2,3 Frankenia composita Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 392 (1928), in sched., pro hyb.

≡ Frankenia laevis subsp. composita (Pau & Font Quer) Nègre

Ind. loc.:—“Hab. in salsuginosis littoris rhiphaei, pr. Marsa Saguira (Bocoya), 10 m. alt.; 13 majii”

Lectotype (designated by Santos Guerra 1992: 136):—In salsuginosis littoris rhiphaei, pr. Marsa Saguira (Bocoya), 10 m, 13 May 1927, Font Quer 392-1927 (lectotype MA 78717 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00078717]; isolectotypes BC 8140 !, BCN 17312 [photo!], GDA 29450 [photo!], MA 78716 !, MPU 005218 [photo!], MPU 006298 [photo!])

Notes:—Pau & Font Quer in Font Quer (1928) published this nothotaxon as a hybrid between Frankenia laevis L. and F. pulverulenta L., but according to Santos Guerra (1992) there is not any evidence of the hybrid condition of the plant. The specimen MA 78717 was labelled as type by Santos Guerra in 1991, and the GDA voucher as isolectotype by Vizoso in 2013.

1 Fraxinus angustifolia f. perangusta Pau in Font Quer, Iter Marocc. 1930 nº 519 (1932), in sched., nom. nud.

= Fraxinus angustifolia Vahl subsp. angustifolia

Notes:—The form was published as var. rostrata Guss. The lack of pubescence in plants distributed in the exsiccata Iter Maroccanum 1930 nº 519 is a diagnostic character of subsp. angustifolia .

1,2,3 Fumana bracteifera Pau in Font Quer, Iter Marocc. 1930 nº 450 (1932), in sched.

=§ Fumana juniperina (Lag. ex Dunal) Pau

Ind. loc.:—“Hab. in quercetis suberis juxta ‘Cuesta Colorada’, 50 m. alt.; 1 apr.”

Lectotype (designated here):—In quercetis suberis juxta ‘Cuesta Colorada’, 50 m, 1 April 1930, Font Quer 450-1930 (lectotype BC 98415 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98415 .jpg]; isolectotypes GDA 3195 [photo!], MA 81850 !, MPU 006727 [photo!], S 07-8627 [photo!], S-G 2729 [photo!])

Notes:—On the original label the name Fumana thymifolia var. bracteifera (Pau) Font Quer is also mentioned, as synonym. The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as F. juniperina by G ̧emes in 2013. APD (2020) refer this name to F. thymifolia ( L.) Webb, but according to G ̧emes & Molero (2002) it should be synonymized under F. juniperina . Leaves of the types are lanceolate, acute, with thickened, scarcely revolute margins, features that match with F. juniperina .

1,2,3 Fumaria ajmasiana Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 120 (1929), in sched.

Ind. loc.:—“Hab. in sepibus, c. Xauen, 600 m. alt.; 25 aprilis”

Lectotype (designated here):—In sepibus, c. Xauen, 600 m, 25 April 1928, Font Quer 120-1928 (lectotype BC 39965 ! [https://www.ibb.csic.es/herbari/ JPEG / BC39965 .jpg]; isolectotypes B 10 0159918 [photo!]*, BCN 18023 [photo!], GDA 36430 [photo!], MA 43524 !, MPU 006431 [photo!]*)

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. Dobignard (2009) considers the MPU specimen as isotype, and El Oualidi et al. (2012) the B specimen as type. The BC specimen was identified as Fumaria ouezzanensis subsp. ramosa Lidén by Valdés, the MA specimen as F. ouezzanensis Pugsley by Lidén in 1983, and the B specimen as “ F. ouezzanensis subsp. ajmasiana (Pau & Font Quer) Lidén ” (unpublished name) by Lidén in 1981. It should be noted that Lidén (1986: 75) considers this name as a nomen nudum and includes it in the synonymy of his new name F. ouezzanensis subsp. ramosa . Nevertheless, according to Art. 30.8 ( Turland et al. 2018) F. ajmasiana is validly published, since Pau & Font Quer included the description in the label, and printed matter accompanying specimens; this being before 1953 makes the concerned name as effectively published.

1,2,3 Fumaria capreolata var. oscilans Pau & Font Quer in Font Quer, Iter Marocc. 1929 nº 171 (1930a), in sched.

= Fumaria macrosepala subsp. obscura (Pugsley) Lidén

Ind. loc.:—“Hab. in umbrosis c. Sidi Brahim (Mtigua), 1000 m. alt.; 12 majii”

Lectotype (designated here):—In umbrosis c. Sidi Brahim (Mtigua), 1000 m, 12 May 1929, Font Quer 171-1929 (lectotype BC 98222 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98222 .jpg]; isolectotypes BCN 72347 [photo!], BM 000552398 [photo!]*, G 00014157 [photo!], GDA 36439 [photo!], MA 43386 !*, MPU 006690 [photo!]*, MPU 006691 [photo!]*)

Notes:—The BC specimen was identified as F. macrosepala subsp. obscura by Valdés, the G specimen was labelled as syntype and identified as F. macrosepala Boiss. by Sell in 1963, and the MA specimen also as F. macrosepala by Lidén in 1983. The GDA specimen was labelled as syntype by Vizoso in 2013. Lidén (1986: 64) records different specimens ( BM, MPU, MA, etc.) without selecting any lectotype. As in the case of F. ajmasiana , this author considers this variety a nomen nudum, although the description is included in the label and, according to Art. 30.8 ( Turland et al. 2018), it is validly published.

1,2,3 Fumaria megasepala Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 275 (1924a)

≡ Fumaria macrosepala subsp. megasepala (Pau) Lidén

Ind. loc.:—“Beni Hozmar, en las grietas de los peñascos de Gorgues, a unos 1.000 metros de altura”

Lectotype (designated here):—Beni Hozmar, en las rendijas de los peñascos, May 1921, Pau s.n. (lectotype MA 43457 [photo!] [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00043457]; isolectotype BC 39677 !)

Notes:—The BC specimen was identified as F. macrosepala by Valdés. Lidén (1986: 64) typifies this taxon on a K specimen collected by Ball in Tetouan, surely because Pau (1924a) cites as synonym the F. macrosepala included in the catalogue of plants from Morocco by Ball (1878: 315). But Pau did not examine the specimens in the herbarium of Ball, only those of his own collection. So the one cited by Lidén could not be used by Pau to describe this taxon. Actually, he used one collected by himself in 1921 in Beni Hozmar, as reported in the protologue.

1 Fumaria muralis var. curta Pau in Bol. Soc. Aragonesa Ci. Nat. 7: 69 (1908)

= Fumaria sepium Boiss. & Reut. subsp. sepium

Ind. loc.:—“D. Benito Vicioso, en su visita hecha á Ceuta por los días 15–17 de Mayo”

Holotype:— Inmediaciones de Ceuta, 16 May 1907, Vicioso s.n. ( MA 43421 ! [http://colecciones.rjb.csic.es/ #cardAdv.php?CatalogNumber=MA-01-00043421])

Notes:—The specimen was identified as F. sepium subsp. sepium by Lidén in 1983. It should be noted that Lidén did not mention this specimen as a type in his monograph—he merely indicated “ type not seen” ( Lidén 1986: 75).

1,3 Galium brunneum var. rhiphaeum Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 607 (1928), in sched.

= Galium brunneum Munby

Ind. loc.:—“Hab. in rupestribus calcareis montis Yebel Hammam (Littore rhiphaeo-Bocoya), 150 m. alt.; 4 majii”

Lectotype (designated here):—In rupestribus calcareis montis Yebel Hammam (Littore rhiphaeo-Bocoya), 150 m, 4 May 1927, Font Quer 607-1927 (lectotype BC 27440 ! [https://www.ibb.csic.es/herbari/ JPEG / BC27440 . jpg]; isolectotypes BCN 74133 [photo!], MA 118427 !, MPU 006380 [photo!])

1,3 Galium constrictum var. rhiphaeum Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 611 (1928), in sched.

= Galium debile Desv.

Ind. loc.:—“ Hab. in paludosis altiplanitiei Isaguen d. (Atlante rhiphaeo), 1600 m. alt., solo siliceo; 7 julii” Lectotype (designated here):—In paludosis altiplanitiei Isaguen d. (Atlante rhiphaeo), 1600 m, 7 July 1927, Font Quer 611-1927 (lectotype BC 27074 ! [https://www.ibb.csic.es/herbari/ JPEG / BC27074 .jpg]; isolectotypes BCSennen 826346!, BCN 74132 [photo!], GDA 38508 , MA 117763 !, MPU 005224 [photo!])

Notes:— The MA and GDA specimens were identified as G. debile by Ortega in 2000 and 2004, respectively. The GDA specimen was labelled as syntype by Vizoso in 2013 .

1,3 Galium gerardii var. parviflorum Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 335 (1924a)

=§ Galium lucidum All.

Ind. loc.:—“Yebel Dersa ( M. Martínez)”

Holotype:—Djebel Dersa, 1921, Martínez s.n. (MA 117327!; [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00117327])

Notes:—The specimen was identified as G. lucidum by Ortega in 2000. According to APD (2020) the accepted name for this variety is Galium mollugo subsp. erectum Syme , probably because Maire combined the variety of Pau into G. mollugo in Jahandiez & Maire (1934: 713). However, it should be noted that Maire considered that this species is included G. lucidum . Both species differ by the shape and size of the leaves: G. lucidum has lanceolate to linear leaves (as the holotype), and G. mollugo elliptical to oblanceolate ( Jury & Rutherford 2002b).

1,3 Galium lucidum var. guilhelmi Pau & Sennen in Sennen, Diagn. Nouv.: 192 (1936b)

=§ Galium lucidum All.

Ind. loc.:—“ Maroc: Metalza massif de Tendri, vers 1700 m.”

Lectotype (designated here):— Maroc: Metalza massif de Tendri, vers 1700 m, 27 May 1933, Sennen & Mauricio 8809 (lectotype MA 117369 ! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00117369]; isolectotypes BC-Sennen 826399!, MPU 009337 [photo!], VAL 142736 [photo!])

Notes:—In Carrasco (1975) this name appears as “ G. mollugo var. guilhelmi ”. The gathering was distributed in the Sennen’s exsiccata Plantes d’Espagne 1933 nº 8809. The MA specimen was identified as G. lucidum by Ortega in 2000. This should be the accepted name, instead of Galium mollugo subsp. erectum Syme as indicated in APD (2020).

1,3 Galium mollugo var. hosmariense Pau ex Maire in Bull. Soc. Hist. Nat. Afrique N. 22(8): 294 (1931)

=§ Galium lucidum All.

Ind. loc.:—“Rochers calcaires du Mont Fahies près de Ceuta (FONT-QUER et MAIRE, 1930)”

Lectotype (designated here):—In montis Djebel Fahies (Andjera), 28 June 1930, Font Quer 622-1930 (lectotype MPU 335206 [photo!] [https://explore.recolnat.org/occurrence/ EB9FBDBCB9F14ED78403814C8F24B72E]; isolectotypes BC 98458 !, BCN 73773 , MA 117317 )

Notes:— Pau (1924a) published this variety under Galium gerardii Vill. , but without adding any description, only noting that it is a form growing on fresh and shady places. Later Maire (1931) added a description and subordinated it to G. mollugo subsp. erectum Syme. The MA specimen was identified as G. lucidum by Krendl in 1971. This should be the accepted name, not Galium mollugo subsp. erectum as indicated in APD (2020).

1,2,3 Genista acutiflora Pau in Font Quer, Iter Marocc. 1930 nº 292 (1932), in sched.

= Genista ancistrocarpa Spach

Ind. loc.:—“Hab. in paludibus, pr. emporium Sok-el-jemis (Sahel), c. El Araix, 100 m. alt.; 25 mart. fl., 4 junii fr.”

Lectotype (designated here):— In paludibus, pr. emporium Sok-el-jemis (Sahel), c. El Araix, 100 m, fl. 25 March 1930, fruct. 4 June 1930, Font Quer 292-1930 (lectotype BC 802820 ! [https://www.ibb.csic.es/herbari/ JPEG / BC802820 .jpg]; isolectotypes BCN 73038 [photo!], BM 000843206 [photo!], GDA 33754 [photo!], MA 58846 !, MA 58847 !, MPU 005278 [photo!], Z 000022793 [photo!])

Note:—The GDA specimen was labelled as syntype by Vizoso in 2013, and the BC specimen was identified as G. ancistrocarpa by Talavera in 1995.

1,2,3 Genista caballeroi Pau in Caballero, Trab. Mus. Nac. Ci. Nat., Ser. Bot. 11: 19 (1917)

= Genista tricuspidata Desf.

Ind. loc.:—“ copiosissima infra Hardú in latere septentrionale Montis Gurugú. Fl. Junio ” Lectotype (designated by Nualart et al. 2018: 617):—Gurugú, 12 June 1915, Caballero s.n. (lectotype MA 58809 ! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00058809]; isolectotype MA 58812 !)

1,2,3 Genista mogadorensis Pau in Actas Real Soc. Esp. Hist. Nat. 26: 232 (1897)

= Genista tricuspidata Desf.

Ind. loc.:—“-”

Type:—not seen

Notes:— Pau (1897) referred to his new species the western and southern Moroccan citations of Genista tridens Cav. by Ball (1878), since Ball had misapplied this name. Although Pau (1897) published this taxon name without a description, it can be considered validly published according to Art. 41 Note 3 ( Turland et al. 2018), because the publication by Ball (1878) includes a description for these citations.

1,2,3 Genista nociva Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 276 (1928), in sched.

≡ Genista carpetana subsp. nociva (Pau & Font Quer) C.Vicioso & M.Laínz

Ind. loc.:—“ Hab. in pratis humidis altiplanitiei Isaguen dictae (Atlante rhiphaeo), 1600 m. alt.; 7 julii” Lectotype (designated here):—In pratis humidis altiplanitiei Isaguen dictae (Atlante rhiphaeo), 1600 m, 7 July 1927, Font Quer 276-1927 (lectotype BC 13941 ! [https://www.ibb.csic.es/herbari/ JPEG / BC13941 .jpg]; isolectotypes BCN 16524 [photo!], BM 000843208 [photo!], GDA 33843 , MA 58947 !, MPU 006316 [photo!]) Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as Genista carpetana Lange by Talavera in 1995 without indicating any infraspecific taxon. El Oualidi et al. (2012) cite the specimen RAB 077738 [photo!] as type, although it corresponds to a later collection in the type locality (collected in 1929 by Font Quer).

1,3 Genista retamoides var. mauritiana Pau & Sennen ex Maire in Bull. Soc. Hist. Nat. Afrique N. 27(6): 219 (1936) = Genista spartioides Spach

Ind. loc.:—“Rif: Mouley-Rechid (Oulad-Settout), collines calcaires (SENNEN & MAURICIO)”

Lectotype (designated here):— Maroc: Muley-Rechid , coteaux calcaires (Ubast-Settut), 6 May 1934, Sennen & Mauricio 9316 (lectotype BC-Sennen 825368! [https://www.ibb.csic.es/herbari/ JPEG / BC825368 .jpg]; isolectotypes BC 80739 !, MA 58578 !, MA 159033 !, MPU 009473 [photo!], MPU 009474 [photo!], VAL 170534 [photo!])

Notes:—This taxon name was proposed by Sennen & Mauricio (1934: 145) at species level (“ Genista mauritiana ”) without a description or diagnosis, and distributed in Sennen’s exsiccata Plantes d’Espagne 1934 nº 9316. Two publications in 1936 included a description for this taxon as a variety under G. retamoides Spach : Maire (1936) published in June, and Sennen (1936b: 229) in July. As the first published description was made by Maire the authorship of this taxon should be “Pau & Sennen ex Maire”. The BC and MA specimens were identified as G. spartioides by Talavera in 1995 and 1997, respectively.

1,2,3 Genista tournefortii var. transfretana Pau & Font Quer in Font Quer, Iter Marocc. 1929 nº 230 (1930a), in sched. =§ Genista tournefortii subsp. jahandiezii (Batt.) Talavera & P.E.Gibbs

Ind. loc.:—“ Hab. in quercetis suberis juxta Terxed Tamstat, supra Tala Mzalla (Mtigua), 1400 m. alt.; 12 majii” Lectotype (designated here):—In quercetis suberis juxta Terxed Tamstat, supra Tala Mzalla (Mtigua), 1400 m, 12 May 1929, Font Quer 230-1929 (lectotype BC 98068 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98068 .jpg]; isolectotypes BCN 72409 [photo!], GDA 33875 [photo!], MA 58789 !, MPU 002625 [photo!]*)

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The MPU specimen was cited as type by El Oualidi et al. (2012). The BC specimen was identified as G. tournefortii subsp. jahandiezii by Talavera in 1995. APD (2020) synonymizes this variety under G. tournefortii Spach ; in addition, according to Talavera (1999) all the North African plants of this species should be attributed to subsp. jahandiezii .

1,2,3 Genista triacanthos var. intermedia Pau in Font Quer, Iter Marocc. 1930 nº 296 (1932), in sched.

= Genista tridens subsp. juniperina (Spach) Talavera & P.E.Gibbs

Ind. loc.:—“Hab. in declivibus arenaceis c. Iaarguiz (Beni Hosmar), ad 200 m. alt.; 14 junii”

Lectotype (designated here):—In declivibus arenaceis c. Iaarguiz (Beni Hosmar), 200 m, 14 June 1930, Font Quer 296-1930 (lectotype BC 868773 ! [https://www.ibb.csic.es/herbari/ JPEG / BC868773 .jpg]; isolectotypes BCN 73042 [photo!], GDA 33885 [photo!], MA 58606 !, MPU 006751 [photo!])

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as G. tridens subsp. juniperina by Talavera in 1995.

1,2,3 Genista ulicina var. densiflora Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 272 (1928), in sched.

=§ Genista tournefortii subsp. jahandiezii (Batt.) Talavera & P.E.Gibbs

Ind. loc.:—“Hab. in dumosis, pr. Zarkat (Atlante rhiphaeo), 1400 m. alt.; 11 junii”

Lectotype (designated here):—In dumosis, pr. Zarkat (Atlante rhiphaeo), 1400 m, 11 June 1927, Font Quer 272-1927 (lectotype BC 13692 ! [https://www.ibb.csic.es/herbari/ JPEG / BC13692 .jpg]; isolectotypes BCN 17299 [photo!], GDA 33901 [photo!], MA 58791 !, MPU 006317 [photo!])

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as G. tournefortii subsp. jahandiezii by Talavera in 1995. APD (2020) synonymizes this variety under G. tournefortii Batt. , but according to Talavera (1999) all the North African plants of G. tournefortii should be attributed to the cited subespecies.

1,2,3 Genistella riphaea Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 277 (1928), in sched.

=§ Pterospartum tridentatum subsp. rhiphaeum (Pau & Font Quer) Talavera & P.E.Gibbs

Ind. loc.:—“Hab. in collibus arenaceis juxta Tizzi Iffri, (Spec. fl.) nec non c. Badú (spec fr.) (Atlante rhiphaeo), 1800 et 1600 m.; fl 7 junii, fr. 6 julii”

Lectotype (designated by Fernández Casas 1974: 329):—In collibus arenaceis juxta Tizzi Iffri (Sp. fl.) nec non c. Badú (sp. fr.) (Atlante rhiphaeo), 1600–1800 m, fl. 7 June 1927, fruct. 6 July 1927, Font Quer 277-1927 (lectotype BC 138958 ! [https://www.ibb.csic.es/herbari/ JPEG / BC138958 .jpg]; isolectotypes BCN 17311 [photo!], BM 000843205 [photo!], GDA 33910 [photo!], MA 105511 !, MA 105512 !, MPU 006312 [photo!], MPU 006313 [photo!])

Notes:—The BC specimen was labelled as type by Fernández Casas in 1974, and the GDA as syntype by Vizoso in 2013. APD (2020) does not recognize any infraspecific taxa for Pterospartum tridentatum (L.) Willk., but according to Talavera (2002a) and El Oualidi et al. (2012) subsp. rhiphaeum should be accepted as endemic to Morocco.

1,2,3 Geranium maurum Pau in Font Quer, Iter Marocc. 1930 nº 395 (1932), in sched.

= Geranium atlanticum Boiss.

Ind. loc.:—“Hab. in declivibus umbrosis c. Sidi Ali (Beni Aros), ad 350 m. alt.; 6 majii”

Lectotype (designated here):—In declivibus umbrosis c. Sidi Ali (Beni Aros), 350 m, 6 May 1930, Font Quer 395-1930 (lectotype BC 98264 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98264 .jpg]; isolectotypes BCN 73186 [photo!], BM 000796487 [photo!], GDA 29736 , JE 00010352 [photo!], JE 00010351 [photo!], MA 71451 !, Z 000017699 [photo!])

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The BC and MA specimens were identified as G. atlanticum by Rutherford in 1996 and Aedo in 1997, respectively.

1 Geranium robertianum var. villosolum Pau in Bol. Soc. Ibér. Ci. Nat. 33: 98 (1935)

= Geranium robertianum L.

Ind. loc.:—“Abdelkader”

Holotype:—( Marruecos) Sidi-Abdelkader (Beni-Sicar), 7 April 1933, Pardo s.n. ( MA 71374 ! [http://colecciones. rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00071374/])

Note:—The specimen was identified as G. robertianum by Aedo in 1997.

1,2,3 Hedysarum capitellatum Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 339 (1928), in sched.

=§ Hedysarum aculeolatum Munby ex Boiss. subsp. aculeolatum

Ind. loc.:—“ Hab. in herbosis, ad pedem montis Yebel Malmusi dictis (Littore rhiphaeo), 50 m. alt.; 3 majii” Lectotype (designated by Valdés 1996b: 264):—In herbosis, ad pedem montis Yebel Malmusi dictis (Littore rhiphaeo), 50 m, 3 May 1927, Font Quer 339-1927 (lectotype BC 17938 ! [https://www.ibb.csic.es/herbari/ JPEG / BC17938 .jpg]; isolectotypes BC 809644 !, BCN 17317 [photo!], BM 000842794 [photo!], G 00424834 [photo!], GDA 33921 [photo!], MA 68785 !, MPU 006293 [photo!], S 09-22205 )

Notes:— The specimen BC 17938 was labelled as lectotype by Valdés in 1996 and the GDA voucher as syntype by Vizoso in 2013. The BC and MA specimens were identified as H. aculeolatum subsp. aculeolatum by Valdés in 1996 and 1998 respectively .

1,2,3 Hedysarum zeluanum Pau in Ann. Sci. Acad. Polytecn. Porto 6: 99 (1911)

=§ Hedysarum aculeolatum Munby ex Boiss. subsp. aculeolatum

Ind. loc.:—“-”

Holotype:— Zeluan: La Puntilla, 2 May 1910, Pau s.n. ( MA 68840 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00068840])

Notes:—Although there is no mention of a concrete locality in the protologue, it should be noted that the publication by Pau (1911) reports a trip to Melilla and Zeluan in 1910. This specimen was identified as H. aculeolatum subsp. aculeolatum by Valdés in 1999. Pau’s taxon is synonymized under H. aculeolatum s.l. in APD (2020), but according to Valdés (2002b), it should be attributed to subsp. aculeolatum .

1 Helianthemum appeninum var. riffeum Pau in Ann. Sci. Acad. Polytecn. Porto 6: 97 (1911)

= Helianthemum virgatum (Desf.) Pers.

Ind. loc.:—“ Melilla … En sitios incultos”

Holotype:— Frequens ad Tetuan et Melilla, April May 1910, Pau s.n. ( MA 80650 ! [http://colecciones.rjb.csic. es/#cardAdv.php?CatalogNumber=MA-01-00080650])

Notes:— Pau (1911) describes this taxon by indicating only the pinkish colour of the petals, which is sufficient to characterize it. This specimen was identified as H. apenninum ( L.) Mill. s.l. by López González in 1992, but Moroccan plants with pinkish petals of this group are usually attributed to H. virgatum ( Raynaud 1999) .

3 Helianthemum atriplicifolium var. macrocalycinum Pau in Font Quer, Iter Marocc. 1928 nº 279 (1929), in sched.

= Cistus atriplicifolius var. macrocalycinus (Pau) Dobignard

Ind. loc.:—“Hab. in quercetis suberis, c. Ankod (Gomara), 750 m. alt.; 27 junii”

Lectotype (designated by Soriano 2008: 31):—In quercetis suberis, c. Ankod (Gomara), 750 m, 27 June 1928, Font Quer 279-1928 (lectotype BC 6509 ! [https://www.ibb.csic.es/herbari/ JPEG / BC6509 .jpg]; isolectotypes AMD 60638 [photo!], BC 809653 !, BCN 42609 !, G 00396559 [photo!]*, GDA 3327 [photo!], MA 79999 !, MPU 006625 [photo!], MPU 006626 [photo!], GDA 3327 )

Notes:—The GDA specimen was labelled as isolectotype by Vizoso in 2015. The G specimen was mentioned by Dobignard (2009) as isotype. The MA and BC specimens were identified as Halimium atriplicifolium subsp. macrocalycinum (Pau) Greuter & Burdet by Nogueira in 1991 ( MA 79999), Gamarra in 1995 ( BC 809653 ), and Soriano in 1997 ( BC 6059).

1,2,3 Helianthemum grosii Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 414 (1928), in sched.

Ind. loc.:—“Hab. in declivibus calc. Yebel Malmusi (Bocoya); 100 m. alt.; 6 majii”

Lectotype (designated by Soriano 2008: 34):—In declivibus calc. Yebel Malmusi (Bocoya), 100 m, 6 May 1927, Font Quer 414-1927 (lectotype BC 7226 ! [https://www.ibb.csic.es/herbari/ JPEG / BC7226 .jpg]; isolectotypes BCN 20158 !, G 00424837 [photo!], GDA 3299 [photo!], MA 80898 !, MA 80899 !)

Notes:—The GDA specimen was labelled as isolectotype by Vizoso in 2015. The BC and BCN specimens were revised by Soriano in 1996, who accepts the species rank.

1 Helianthemum halimifolium var. minoriifolium Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 284 (1924a)

= Cistus halimifolius L.

Ind. loc.:—“Rincón de Medik”

Lectotype (designated here):— Rincon de Medik , 7 May 1921, Pau s.n. (lectotype MA 80049 ! [http://colecciones. rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00080049]; isolectotypes MA 80047 !, MA 80050 !) Note:—The selected lectotype is the specimen bearing the biggest sample.

1 Helianthemum lasiocalycinum var. angustifolium Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 278 (1929), in sched.

=§ Cistus halimifolius var. lasiocalycinus (Boiss. & Reut.) Demoly

Ind. loc.:—“Hab. in quercetis suberis montis Sugna (Yebala), 700 m. alt.; 1 junii”

Lectotype (designated here):—In quercetis suberis montis Sugna (Yebala), 700 m, 1 June 1928, Font Quer 278- 1928 (lectotype BC 7227 ! [https://www.ibb.csic.es/herbari/ JPEG / BC7227 .jpg]; isolectotypes BCN 42470 !, GDA 3306 [photo!], MA 80064 !, MPU 006627 [photo!])

Notes:—In Carrasco (1975) this taxon is named “ Halimium lasiocalycinum var. angustifolium Pau & Font Quer ”.

The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as Halimium lasiocalycinum (Boiss. & Reut.) Engl. & Pax by Soriano in 1997, and the MA voucher as Halimium halimifolium subsp. multiflorum (Dunal) Maire by Nogueira in 1991. According to Demoly (2006) both names fall in synonymy of Cistus halimifolius var. lasiocalycinus .

1 Helianthemum ledifolium var. hirtulum Pau in Sennen, Pl. Espagne 1931 nº 8077 (1931–1932), in sched., nom. nud.

= Helianthemum ledifolium ( L.) Mill.

Note:—This variety name was also mentioned by Sennen (1936a), but without description nor diagnosis.

1 Helianthemum mauritianum Pau & Sennen , in sched. ( MA 81145! [http://colecciones.rjb.csic.es/#cardAdv.

php?CatalogNumber=MA-01-00081145]), nom. nud.

= Helianthemum papillare Boiss.

Notes:—The name was handwritten by Pau on a revision label of a specimen distributed in Sennen’s exsiccata Plantes d’Espagne 1934 nº 9266, originally identified as Helianthemum pediforme Pomel. Based on examination of this specimen it should be attributed to H. papillare .

1,3 Helianthemum origanifolium f. leiocladum Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 418 (1928), in sched.

= Helianthemum origanifolium subsp. africanum M.B.Crespo, M.Á.Alonso, A.Vicente & J.L.Villar

Ind. loc.:—“Hab. in declivibus calc. Yebel Malmusi (Littore rhiphaeo), 100 m. alt.; 3 majii”

Lectotype (designated here):—In declivibus calc. Yebel Malmusi (Littore rhiphaeo), 100 m, 3 May 1927, Font Quer 418-1927 (lectotype BC 7045 ! [https://www.ibb.csic.es/herbari/ JPEG / BC7045 .jpg]; isolectotypes BCN 42614 [photo!], MA 81449 !, MPU 006286 [photo!], MPU 006285 [photo!])

Notes:—The MA and BC specimens identified as H. marifolium subsp. origanifolium (Lam.) G.López by G. López himself (=López González) in 1991 and Soriano in 1996, respectively. According to Crespo et al. (2016) this form proposed by Pau & Font Quer in Font Quer (1928) does not differ from the new subspecies described by the same authors for the North African populations of the species.

1,2,3 Helianthemum rhiphaeum Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 405 (1929), in sched.

=§ Cistus halimifolius var. rhiphaeus (Pau & Font Quer) Demoly

Ind. loc.:—“Hab. in collibus saxosis, juxta Tizzi Iffri, 1800 m. alt. (Atlante rhiphaeo), 8 junii”

Lectotype (designated by Soriano 2008: 31):—In collibus saxosis, juxta Tizzi Iffri, 1800 m, 8 June 1927, Font Quer 405-1927 (lectotype BC 6547 ! [https://www.ibb.csic.es/herbari/ JPEG / BC6547 .jpg]; isolectotypes BCN 42455 !, GDA 3381 [photo!], MA 79987 !, MPU 006507 [photo!], S-G 3025 [photo!])

Notes:—The MPU specimen was labelled as isotype by Durand and Loup in 2006, and the GDA voucher as isolectotype by Vizoso in 2015. The BC specimen was identified as Halimium lasiocalycinum subsp. riphaeum (Pau & Font Quer) Maire by Soriano in 2006. Vizoso (2018) indicates as type the specimen BC 6546! that corresponds to var. elatum Font Quer. The accepted name follows Demoly (2006).

1 Helminthia comosa var. spinulosa Pau in Vidal, Asoc. Esp. Progr. Ci. Congr. Cádiz 6: 150 (1928)

= Helminthotheca comosa (Boiss.) Holub

Ind. loc.:—“Dar Riffien”

Type:—not seen

Notes:—The name was cited as H. echioides var. spinulosa Pau by Talavera et al. (2016).

1,2,3 Hieracium praecox var. ajmasianum Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 437 (1929), in sched.

≡ Hieracium ajmasianum (Pau & Font Quer) Dobignard

Ind. loc.:—“ Hab. in silvis cedrorum montis Yebel Afestal dictis (Gomara), solo siliceo, 1900 m. alt.; 25 junii” Lectotype (designated by Ferrer-Gallego et al. 2019):—In silvis cedrorum montis Yebel Afestal dictis (Gomara), 1900 m, 25 June 1928, Font Quer 437-1928 (lectotype BC 36453 ! [https://www.ibb.csic.es/herbari/ JPEG / BC36453 .jpg]; isolectotypes BCN 19394 [photo!], GDA 4742 [photo!], MA 141575 !, MA 141576 !, MPU 006593 [photo!], MPU 006594 [photo!])

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as Hieracium murorum L. by Mateo in 1999. This author includes this variety in the H. murorum -complex ( Mateo 2002), but Dobignard (2009) and APD (2020) give the plant specific status, as an endemic to the Riff mountains.

1,3 Hieracium riofrioi Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 438 (1929), in sched.

Ind. loc.:—“ Hab. in silvis pinsaporum inter montes Kalaa et Tisuka, solo calc., 1500 m. alt.; 15 junii” Lectotype (designated by Ferrer-Gallego et al. 2019):—In silvis pinsaporum inter montes Kalaa et Tisuka, 1500 m, 15 June 1928, Font Quer 438-1928 (lectotype BC 37369 ! [https://www.ibb.csic.es/herbari/ JPEG / BC37369 .jpg]; isolectotypes BC 37367 !, BC 37368 !, GDA 4748 [photo!], MA 141688 !, MA 141690 !, MPU 006592 [photo!]*, MPU 006638 [photo!]*, MPU 006694 [photo!]*)

Notes:— The GDA specimen was labelled as syntype by Vizoso in 2013. The selected lectotype is the BC specimen with more mature samples. Dobignard (2009) mentions the MPU specimens as isotypes .

1,2,3 Hippocrepis scabra var. maroccana Pau & Font Quer in Font Quer, Iter Marocc. 1929 nº 276 (1930a), in sched.

= Hippocrepis monticola Lassen

Ind. loc.:—“Hab. in collibus Tizi Usli dictis (Guesnaia), 1500 m. alt.”

Lectotype (designated here):—In collibus Tizi Usli dictis (Guesnaia), 1500 m, 10 June 1929, Font Quer 276- 1929 (lectotype BC 806562 ! [https://www.ibb.csic.es/herbari/ JPEG / BC806562 .jpg]; isolectotypes BCN 72149 [photo!], BM 000842793 [photo!], G 00022225 [photo!], GDA 33237 [photo!], MA 68546 !, MA 68550 !, MPU 006667 [photo!], MPU 006668 [photo!], S-G 8866 [photo!])

Notes:— The G and S specimens were labelled as isotypes by Lassen in 1982 and the GDA voucher as syntype by Vizoso in 2013. The BM, G, and S specimens were identified as Hippocrepis monticola by Lassen in 1982, the BC, GDA, and MA specimens were similarly identified by Talavera, in 1999 .

1,2,3 Hirschfeldia littorea Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 229 (1928), in sched.

≡ Erucastrum littoreum (Pau & Font Quer) Maire

Ind. loc.:—“ Hab. in dumosis c. Torres de Alcalá, cab. Beni Bu-Frah (Littore rhiphaeo), 50 m. alt.; 4 junii” Lectotype (designated here):—In dumosis c. Torres de Alcalá , cab. Beni Bu-Frah (Littore rhiphaeo), 50 m, 4 June 1928, Font Quer 229-1927 (lectotype BC 4019 ! [https://www.ibb.csic.es/herbari/ JPEG / BC4019 .jpg]; isolectotypes BCN 17304 [photo!], BM 010752158 [photo!], GDA 28269 [photo!], MA 46479 !, MA 46480 !, MPU 006334 [photo!], MPU 006335 [photo!])

Notes:—The MPU specimens were labelled as isotypes by Sennedot and Scouppe in 2006, and the GDA specimen as syntype by Vizoso in 2013. The BC specimen was identified as Erucastrum littoreum by Gómez Campo in 1996.

3 Hordeum murinum var. pedicellatum Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 96 (1928), in sched.

= Hordeum murinum subsp. glaucum (Steud.) Tzvelev

Ind. loc.:—“Hab. in locis salsuginosis juxta Marsa Saguira (Littore rhiphaeo), 50 m. alt.; 13 majii”

Lectotype (designated here):—In locis salsuginosis juxta Marsa Saguira (Littore rhiphaeo), 50 m, 13 May 1927, Font Quer 96-1927 (lectotype BCN 73997 ! [http://www.ub.edu/cedocbiv/herbarivirt/122236]; isolectotypes BM 000060862 [photo!], MPU 006356 [photo!])

Notes:—The BM specimen was identified as Hordeum murinum subsp. glaucum by Jacobsen, without date.

1,3 Hypericum aegyptiacum var. maroccana Pau in Cavanillesia 4: 157 (1931)

≡ Hypericum aegyptiacum subsp. maroccanum (Pau) N.Robson

Ind. loc.:—“-”

Lectotype (designated here):—Sud-Ouest: Cap Ghir, versant Sud, 27 March 1931, Jahandiez 57 (lectotype MA 77865 ! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00077865]; isolectotypes BC 138333 !, BM 000617919 [photo!], MARS 64215 [photo!])

Notes:— Pau (1931) described this new variety from a specimen distributed by Jahandiez (nº 57) originally labelled as H. aegyptiacum L. The BM specimen was labelled as type and identified as H. aegyptiacum subsp. maroccanum by Robson in 1991.

1,2,3 Hypericum andjerinum Font Quer & Pau in Font Quer, Iter Marocc. 1930 nº 423 (1932), in sched.

Ind. loc.:—“Hab. in declivibus argilloso-calcareis montis Hafa-el-Gorbeb (Andjera), ad 480 m. alt.”

Lectotype (designated here):—In declivibus argilloso-calcareis montis Hafa-el-Gorbeb (Andjera), 480 m, 11

June 1930, Font Quer 423-1930 (lectotype BC 98364 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98364 .jpg]; isolectotypes BM 001204683 [photo!], BCN 20125 [photo!], GDA 31291 [photo!], MA 78024 !, MPU 006729 [photo!], S-G 3303 [photo!], Z 000018582 [photo!])

Notes:— The GDA specimen was labelled as syntype by Vizoso in 2013. Determination of the BC specimen was confirmed by Robson in 1996 .

1,3 Hypericum tomentosum var. viridulum Pau in Vidal, Bol. Soc. Esp. Hist. Nat. 22: 57 (1922)

=§ Hypericum pubescens Boiss.

Ind. loc.:—“Xauen”

Lectotype (designated by Robson 1996: 182):— Xauen ( Marruecos), 10 June 1921, Vidal s.n. ( MA 78357 ! [http:// colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00078357])

Notes:—The specimen was identified as H. pubescens by Ramos in 1989, although APD (2020) includes this variety in H. tomentosum L. Both species differ by the sepals, and those of the lectotype are long-aristate, a diagnostic feature of H. pubescens .

1 Iberis gibraltarica f. hosmariensis Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 280 (1924a)

= Iberis gibraltarica L.

Ind. loc.:—“Rendijas de los peñascos elevados de Gorgues, en Beni Hozmar”

Lectotype (designated here):—Gorgues, en los peñascos, 1000 m, 8 May 1921, Pau s.n. (lectotype MA 44631 ! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00044631]; isolectotype MA 44632 !) Notes:—The selected lectotype is the only specimen that bears a plant with silicules. The shape of the base of these silicules (attenuate-cuneate) is one of the diagnostic characters of this form. The isolectotype was labelled originally as “ Iberis gibraltarica forma tetuanensis ”, but in the original publication, Pau (1924a) proposed the name “ hosmariensis ”. Specimens were identified as I. gibraltarica by Moreno in 1987.

1,2,3 Iberis grosmiquelii Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 220 (1928), in sched.

≡ Iberis carnosa subsp. grosmiquelii (Pau & Font Quer) Dobignard

Ind. loc.:—“Hab. in cedretis montis Yebel Lerz d. (Atlante rhiphaeo); fl. ad 2100 m. alt., 12 junii, fr. ad 1900 m. alt., 14 julii, lectis”

Lectotype (designated by Montserrat 1996: 247):—in cedretis montis Yebel Lerz d. (Atlante rhiphaeo), 1900– 2100 m, fl. 12 June 1927, fruct. 14 July 1927, Font Quer 220-1927 (lectotype BC 5238 ! [https://www.ibb.csic.es/ herbari/ JPEG / BC5238 .jpg]; isolectotypes BC 809753 !, BCN 17289 [photo!], GDA 28298 [photo!], MA 44766 !, MPU 006336 [photo!]*, S 07-10823 [photo!])

Notes:—The GDA specimen was labelled as isolectotype by Vizoso in 2013. Dobignard (1997, 2009) cites as isotype the MPU specimen, which is labelled also as isotype by Sennedot and Scouppe in 2006. The identification of the isolectotype from Herb. BC was validated by Bergés and Guàrdia in 1993.

1,2,3 Iberis grosmiquelii var. senneniana Pau in Bol. Soc. Iber. Ci. Nat. 31: 99 (1934)

≡ Iberis carnosa subsp. senneniana (Pau) Dobignard

Ind. loc.:—“Tidiguin, VI, 1933 (Sennen y Mauricio)”

Lectotype (designated here):—Tidiguin, 19–22 June 1933, Sennen & Mauricio s.n. (lectotype MA 44767 ! [http:// colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00044767]; isolectotype BM 010779450 [photo!] MA 44769 !, MPU 301555 [photo!], VAL 159008 [photo!])

Notes:—The selected lectotype is part of the materials of Sennen’s exsiccata Plantes d’Espagne 1933 nº 8659. It does not have the printed label of the exsiccata, but two handwritten labels, one by Sennen with the indication of the genus, locality and collection data, and the other by Pau identifying it as var. senneniana .

1,2,3 Iberis linifolia var. maroccana Pau in Font Quer, Iter Marocc. 1928 nº 126 (1929), in sched.

= Iberis contracta subsp. rifana (Emb. & Maire) Valdés

Ind. loc.:—“ Hab. in silvis pinsaporum collis Bab de Ruida (Beni Darkul), solo calc., 1500 m. alt.; 23 junii” Lectotype (designated here):—In silvis pinsaporum collis Bab de Ruida (Beni Darkul), 1500 m, 23 June 1928, Font Quer 126-1928 (lectotype BC 5142 ! [https://www.ibb.csic.es/herbari/ JPEG / BC5142 .jpg]; isolectotypes BCN 74219 [photo!], GDA 28304 [photo!], MA 44796 !, MPU 006428 [photo!], S 12-16225 [photo!], Z 000170719 [photo!])

Note:— The GDA specimen was labelled as syntype by Vizoso in 2013 .

1,2,3 Inula montana var. lanata Font Quer & Pau in Font Quer, Iter Marocc. 1930 nº 657 (1932), in sched.

= Pentanema montanum (L.) D.Gut.Larr., Santos-Vicente, Anderb., E.Rico & M.M.Mart.Ort.

Ind. loc.:—“Hab. in collibus calc. montis Djebel Dersa, supra Tetauen, ad 400 m. alt.; 20 junii”

Lectotype (designated here):—In collibus calc. montis Djebel Dersa, supra Tetauen, 400 m, 20 June 1930, Font Quer 657-1930 (lectotype BC 137328 ! [https://www.ibb.csic.es/herbari/ JPEG / BC137328 .jpg]; isolectotypes BCN 73553 [photo!], MPU 005246 [photo!])

Note:—The BC specimen was identified as I. montana L. by Vogt and Oberprieler in 1995.

Iris riphienensis Pau , in sched. (MA 23216! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01- 00023216]), nom. nud.

= Iris tingitana Boiss. & Reut.

Note:—The specimen was identified as I. tingitana by Carvalho in 1994.

1,2,3 Jasione perennis var. cedretorum Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 643 (1928), in sched.

≡ Jasione crispa subsp. cedretorum (Pau & Font Quer) Dobignard

Ind. loc.:—“ Hab. in cedretis montis Yebel Lerz (Atlante rhiphaeo), 1900 m. alt., solo arenaceo; 14 julii” Lectotype (designated here):—In cedretis montis Yebel Lerz (Atlante rhiphaeo), 1900 m, 14 July 1927, Font Quer 643-1927 (lectotype BC 40267 ! [https://www.ibb.csic.es/herbari/ JPEG / BC40267 .jpg]; isolectotypes BCN 74809 [photo!], G 00237595 [photo!], GDA 1205 [photo!], MPU 006552 [photo!], MPU 006553 [photo!]) Notes:—Mentioned in Carrasco (1975) as “ Jasione humilis var. cedrotorum Pau & Font Quer ”. The MPU and G specimens were labelled as types by Romo in 2003 and 2005 respectively. The GDA specimen was labelled as syntype by Vizoso in 2013. Romo identified the G specimen as Jasione crispa subsp. cedretorum in 2005.

3 Juncus obtusiflorus var. latifolius Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 112 (1928), in sched.

=§ Juncus punctorius L.f.

Ind. loc.:—“Hab. in paludosis c. Badú (Atlante rhiphaeo), 1500 m. alt.; 6 julii”

Lectotype (designated here):—In paludosis c. Badú (Atlante rhiphaeo), 1500 m, 6 July 1927, Font Quer 112-1927 (lectotype BC 62805! [https://www.ibb.csic.es/herbari/JPEG/BC62805.jpg]; isolectotypes BCN 25698 [photo!], BCN 25710 [photo!], G 00080409 [photo!], GDA 31647 [photo!], MA 19219!, MPU 006354 [photo!]) Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013, and the MPU as isotype by Sennedot and Scouppe in 2006. The BC specimen was identified as J. subnodulosus Schrank by Rutherford in 1996, and the G voucher as J. punctorius by Romero in 2008. According to APD (2020) the accepted name for this variety is J. subnodulosus . However, based on examination of the original material, these plants should be attributed to J. punctorius due to the subacute perianth-segments and the large heads, with more than ten flowers.

1,3 Juncus rhiphaenus Pau & Font Quer in Font Quer, Iter Marocc. 1929 nº 64 (1930a), in sched.

= Juncus foliosus Desf.

Ind. loc.:—“Hab. in silviculis betularum pr. Asib de Ktama, solo uliginoso, ad 1450 m. alt.; 1 julii”

Lectotype (designated here):—In silviculis betularum pr. Asib de Ktama , solo uliginoso, 1450 m, 1 July 1929, Font Quer 64-1929 (lectotype BC 98562 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98562 .jpg]; isolectotypes BCN 71567 [photo!], BM 000803696 [photo!]*, G 00190451 [photo!], GDA 31655 [photo!], MA 18876 !, MPU 006710 [photo!], RAB 026578 *, RAB 026578 bis [photo!]*)

Notes:—The BM specimen was labelled as type by Snogerup in 1972, and the GDA voucher as syntype by Vizoso in 2013. The BC and BM specimens were identified as J. foliosus by Rutherford in 1996 and by Kirschner in 2004, respectively. The BM and RAB specimens were cited as types by Cope & Stace (1978) and El Oualidi et al. (2012), respectively.

1,3 Knautia arvensis var. glandulifera Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 629 (1928), in sched.

= Knautia mauritanica Pomel

Ind. loc.:—“Hab. in cedretis montis Yebel Iguermalez (sp. fl.), nec non pr. Imasinen (sp. fruct.), 1700–1800 m. alt. solo siliceo; fl. 26, fruct. 30 junii”

Lectotype (designated here):—In cedretis montis Yebel Iguermalez (sp. fl.), nec non pr. Imasinen (sp. fruct.),

1700–1800 m, fl. 26 June 1927, fruct. 30 June 1927, Font Quer 629-1927 (lectotype BC 28644 ! [https://www. ibb.csic.es/herbari/ JPEG / BC28644 .jpg]; isolectotypes B 10 0158910 [photo!], BCN 74121 [photo!], GDA 28750 [photo!], MA 120144 !, MPU 006377 [photo!], MPU 006378 [photo!], S 12-7858 [photo!])

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as K. mauritanica by Romo in 1996, as also the GDA and MA specimens by Devesa, in 2000. The specimen at Herb. B was cited as type by El Oualidi et al. (2012).

1,2,3 Lactuca cornigera Pau & Font Quer in Font Quer, Iter Marocc. 1929 nº 476 (1930a), in sched.

≡ Lactuca virosa subsp. cornigera (Pau & Font Quer) Emb. & Maire

Ind. loc.:—“Hab. in quercetis suberis juxta Asib de Ktama, solo saxoso, arenaceo, ad 1650 m. alt.; fl. 12 julii, fr. 10 oct.”

Lectotype (designated here):—In quercetis suberis juxta Asib de Ktama , 1650 m, fl. 12 July 1929, fruct. 10 October 1929, Font Quer 476-1929 (lectotype BC 136595 ! [https://www.ibb.csic.es/herbari/ JPEG / BC136595 . jpg]; isolectotypes BCN 72569 [photo!], GDA 4868 [photo!], MA 140196 !, MA 140197 !, MPU 006640 [photo!], MPU 006641 [photo!], RAB 028348 [photo!]*, S-G 3517 [photo!], Z 000003603 [photo!])

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. El Oualidi et al. (2012) cited the RAB specimen as type. The BC specimen was identified as “ L. virosa f. cornigera ” (unpublished name) by Mejías 1996. Mejías (2002) and Fennane (2014f) do not accept this taxon, although El Oualidi et al. (2012) and Rankou et al. (2013) include it in their catalogues of Moroccan endemisms.

1,2,3 Lamium berengueri Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 378 (1924a)

≡ Lamium flexuosum subsp. berengueri (Pau) Dobignard

Ind. loc.:—“Tetuán a Beni Hozmar”

Lectotype (designated by Mennema 1989: 32):—Tetuan, margen derecha del rio Martín, 5 May 1921, Pau s.n. (lectotype FI 000833 [photo!] [http://parlatore.msn.unifi.it/img72/FI000833.jpg]; isolectotype BC 48550!) Notes:— Dobignard (2009) cites as isotype a G specimen from Xauen of the exsiccata Iter Maroccanum 1928 nº 339, but this gathering cannot be considered original material, since it was collected some years after publication of the taxon by Pau (1924a), and in a different locality from the one in the protologue. On the other hand, the specimen MA 102652! collected by Pau in Beni Hozmar should be considered as original material.

1 Lathyrus setifolius f. majoriflorus Pau in Cavanillesia 2: 89 (1929c)

= Lathyrus setifolius L.

Ind. loc.:—“Xauen (Vidal y López, Font Quer)”

Lectotype (designated here):—Xauen, 15 May 1921, Vidal s.n. ( MA 70764 ! [http://colecciones.rjb.csic.es/ #cardAdv.php?CatalogNumber=MA-01-00070764])

Notes:—The original material includes two gatherings collected in Xauen: one by Vidal and another by Font Quer, the latter distributed in the exsiccata Iter Maroccanum 1928 nº 236 (BC 18224!, BC 18223!, MA 70763!). The selected lectotype is the one collected by Vidal and labelled as belonging to the personal collection of Pau. This specimen was identified as L. setifolius by Gallego in 1995.

1 Lavandula multifida f. glabrifolia Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 372 (1924a)

= Lavandula multifida L.

Ind. loc.:—“Tetuán”

Holotype:—Tetuan, 10 May 1921, Pau s.n. ( MA 99851 !)

Note:—The specimen was identified as L. multifida by Upson in 1997.

1 Lavandula stoechas f. gracilis Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 372 (1924a)

= Lavandula stoechas L.

Ind. loc.:—“ Tetuán … se encuentra, además de en Yebel Dersa en el Rincón de Medik ( M. Martínez)” Holotype:—Tetuan, May 1921, Pau s.n. ( MA 99495 !)

1,2,3 Lavatera vidalii Pau in Vidal, Bol. Soc. Esp. Hist. Nat. 22: 58 (1922)

≡ Malva vidalii (Pau) Molero & J.M.Monts.

Ind. loc.:—“Xauen”

Holotype:—Xauen, 10 May 1921, Vidal s.n. ( MA 77180 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00077180])

1,2,3 Legousia speculum-veneris var. maroccana Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 639 (1928), in sched.

=§ Legousia falcata (Ten.) Janch.

Ind. loc.:—“Hab. in saxosis montis Iguermalez, versus collem Tizzi Iffri d. (Atlante rhiphaeo), 1700 m. alt., solo schistose; 8 junii”

Lectotype (designated here):—In saxosis montis Iguermalez, versus collem Tizzi Iffri d. (Atlante rhiphaeo), 1700 m, 8 June 1927, Font Quer 639-1927 (lectotype BC 137429 ! [https://www.ibb.csic.es/herbari/ JPEG / BC137429 . jpg]; isolectotypes BCN 63553 [photo!], GDA 1220 [photo!], MA 121890 !, MA 121918 !)

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The MA specimen was identified as L. speculum-veneris ( L.) Chaix by Guinea in 1968. Peris et al. (2002) synonimized the variety of Pau to L. scabra (Lowe) Gamisans. Later, Fennane & Mathez (2014) considered that all the Moroccan L. scabra should be included in L. falcata , a similar approach to that of Wahlsteen & Tyler (2019), who sustain that differences among these two species are not clear and there are not enough features to distinguish them, so that they suggest uniting them in L. falcata . However, in APD (2020) var. maroccana is synonymized under L. falcata subsp. castellana (Lange) Jauzein.

1 Leontodon taraxacoides f. leiolepis Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 353 (1924a)

= Leontodon saxatilis Lam.

Ind. loc.:—“Yebel Dersa”

Type:—not seen

1,2,3 Lepidium petrophilum var. afrum Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 127 (1929), in sched.

≡§ Lepidium hirtum subsp. afrum (Pau & Font Quer) J.M.Monts.

Ind. loc.:—“Hab. in silvis pinsaporum montis Yebel Tisuka (Gomara), 1800 m. alt.; 13 majii”

Lectotype (designated by Montserrat 1996: 243):— In silvis pinsaporum montis Yebel Tisuka (Gomara), 1800 m, 13 May 1928, Font Quer 127-1928 (lectotype BC 5578 ! [https://www.ibb.csic.es/herbari/ JPEG / BC5578 .jpg]; isolectotypes BCN 18021 [photo!], BM 000593726 [photo!], GDA 45283 [photo!], MA 43874 !, MA 43875 !, MPU 006426 [photo!], MPU 006427 [photo!])

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as L. hirtum subsp. afrum by Montserrat in 2006. APD (2020) includes this variety under L. hirtum subsp. dhayense (Munby) Thell. ; but Montserrat (1996) and Valdés (2013: 45) consider that subsp. afrum is clearly different from subsp. dhayense by its bigger silicules with a rather laxer indument.

1 Leucanthemum glabrum f. pinnatisectum Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 344 (1924a)

= Mauranthemum decipiens (Pomel) Vogt & Oberpr.

Ind. loc.:—“Tetuán, por encima de la Alcazaba”

Holotype:—Tetuan, 13 May 1921, Pau s.n. ( MA 128516 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00128516])

Note:—The specimen was labelled as holotype by Vogt in 1996, who also identified it as Mauranthemum decipiens .

1,2,3 Linaria arenicola Pau & Font Quer in Font Quer, Iter Marocc. 1930 nº 590 (1932), in sched.

Ind. loc.:—“Hab. in arenaceis maritimis, pr. El Araix; 7 febr.”

Lectotype (designated here):— In arenaceis maritimis, pr. El Araix, 7 February 1930, Font Quer 590-1930 (lectotype BC 98322 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98322 .jpg]; isolectotypes B 10 0165092 [photo!], BCN 20132 [photo!], G 00424862 [photo!], GDA 39357 [photo!], MA 110064 !, MPU 005267 [photo!], S 10- 19130 [photo!], Z 000028440 [photo!])

Notes:— The GDA specimen was labelled as syntype by Vizoso in 2013. Determination of the BC specimen was confirmed by Valdés in 1996 .

1,2,3 Linaria pectinata Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 360 (1929), in sched.

≡ Linaria tristis subsp. pectinata (Pau & Font Quer) Maire

Ind. loc.:—“Hab. in rupestribus saxosisque calc. montis Kalaa, supra Xauen, 900 m. alt.; 22 aprilis”

Lectotype (designated here):—In rupestribus saxosisque calc. montis Kalaa, supra Xauen, 900 m, 22 April 1928, Font Quer 360-1928 (lectotype BC 44898 ! [https://www.ibb.csic.es/herbari/ JPEG / BC44898 .jpg]; isolectotypes BCN 18134 [photo!], G 00424868 [photo!], GDA 39457 [photo!], MA 109878 !, MA 109879 !, Z 000028443 [photo!])

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as L. tristis subsp. pectinata by Valdés in 1996.

1,2,3 Linaria riffea Pau in Ann. Sci. Acad. Polytecn. Porto 6: 99 (1911)

Ind. loc.:—“Zeluan, Melilla ”

Holotype:— Melilla, Zeluan , April May 1910, Pau s.n. ( MA 110310 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00110310])

Notes:— There are two sheets labelled as Linaria riffea in Herb. MA from Zeluan (the Spanish name for Selouane in Nador province), both collected in 1910 by Pau. MA 101310 has two labels, the original one handwritten by Pau and another typewritten with a locality mistake: Tetuan instead of Zeluan. The sheet MA 101311 ! comes from the personal herbarium of Vicioso and has the indication “cult. sem. ex.”. Both plants match the description by Pau (1911) for this species. However, only the first can be considered the holotype, under the pretext that the one of the Vicioso herbarium may be a cultivated specimen from seeds collected by Pau in Selouane and, therefore not original material .

1,3 Linaria sennenii Pau in Sennen & Mauricio, Cat. Fl. Rif Orient.: 86 (1934), nom. nud.

= Linaria micrantha (Cav.) Hoffmanns. & Link

Notes:—This name was cited by Sennen & Mauricio (1934: 86) with the only indication of the locality “Targuist; Tizi-ifri”. Two years later, Sennen (1936a: 78; 1936b: 160) remarks that this species was cultivated by Pau and distributed in the exsiccata Plantes d’Espagne nº 8464 as L. munbyana Boiss & Reut. Specimens of this gathering (BC 137788!, BC-Sennen 825377!) were identified as L. arvensis by Valdés in 1996. In the personal collection of Maire there is a specimen (MPU 074226 [photo!]) labelled as L. sennenii cultivated from seeds of the type and with the indication of “corolla vivide caerulea” (corolla vivid blue), a character of the L. arvensis -group. According to APD (2020) Pau’s taxon should be synonymized under L. micrantha , a species of L. arvensis -group.

1 Linaria spuria var. minoripes Pau in Bol. Real Soc. Esp. Hist. Nat. 21: 202 (1921)

= Kickxia spuria (L.) Dumort.

Ind. loc.:—“Benisicar”

Holotype:— Riff Oriental Cala Charrana (Beniscar), 13 September 1920, Vicioso s.n. ( MA 109083 ! [http:// colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00109083])

Note:— There are two specimens by Vicioso labelled as f. minoripes in MA, but only one of them is a good match for the locality in the protologue; the other ( MA 109082 !) comes from Melilla .

1,2,3 Linaria supina var. ajmasiana Pau in Vidal, Bol. Soc. Esp. Hist. Nat. 22: 59 (1922)

= Linaria tristis subsp. pectinata (Pau & Font Quer) Maire

Ind. loc.:—“Todas las plantas son de Xauen”

Type:—not seen

1,2 Linaria verticillata var. tetuanensis Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 367 (1924a)

= Linaria tristis subsp. pectinata (Pau & Font Quer) Maire

Ind. loc.:—“Frecuente en las rendijas de los peñascos de las cercanías de Tetuán, y subiendo hasta Beni Hozmar”

Lectotype (designated here):—Tetuan, 10 May 1921, Pau s.n. (lectotype MA 110248 ! [http://colecciones.rjb.csic. es/#cardAdv.php?CatalogNumber=MA-01-00110248]; isolectotype BC 44549 !)

Note:—The BC specimen was identified as L. tristis subsp. pectinata by Valdés in 1996.

1,2,3 Linum moroderorum Pau in Ann. Sci. Acad. Polytecn. Porto 6: 98 (1911)

= Linum usitatissimum L.

Ind. loc.:—“ Melilla … En sitios incultos”

Holotype:— Melilla (Riff) in locis incultis et in aggeribus, April 1910, Pau s.n. ( MA 72779 ! [http://colecciones. rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00072779])

Note:— Pau also labelled as “ type ” a specimen from the same locality ( MA 72777 !), collected in 1932, some years after the publication .

1,3 Linum munbyanum var. curtifolium Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 353 (1928), in sched.

= Linum tenue subsp. munbyanum (Boiss. & Reut.) Batt.

Ind. loc.:—“Hab. in quercetis superbis pr. Badú (Atlante rhiphaeo), 1600 m. alt.; 6 julii”

Lectotype (designated here):—In quercetis superbis pr. Badú (Atlante rhiphaeo), 1600 m, 6 July 1927, Font Quer 353-1927 (lectotype BC 11229 ! [https://www.ibb.csic.es/herbari/ JPEG / BC11229 .jpg]; isolectotypes BCN 17315 [photo!], GDA 37273 [photo!], MA 73075 !, MPU 006295 [photo!])

Note:— The GDA specimen was labelled as syntype by Vizoso in 2013 .

1,3 Linum munbyanum var. tenuifolium Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 352 (1928), in sched.

= Linum tenue subsp. munbyanum (Boiss. & Reut.) Batt.

Ind. loc.:—“Hab. in quercetis supra Targuist, 1300 m. alt., solo arenaceo; 20 junii”

Lectotype (designated here):—In quercetis supra Targuist, 1300 m, 20 June 1927, Font Quer 352-1927 (lectotype BC 11230 ! [https://www.ibb.csic.es/herbari/ JPEG / BC11230 .jpg]; isolectotypes BCN 17314 [photo!], GDA 37272 [photo!], MA 73074 !, MPU 006294 [photo!], S 10-1057 [photo!])

Note:— The GDA specimen was labelled as syntype by Vizoso in 2013 .

1,3 Linum narbonense var. afrum Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 245 (1929), in sched.

= Linum narbonense L.

Ind. loc.:—“Hab. in collibus calc. montis Kalaa, 900 m. alt.; 26 majii”

Lectotype (designated here):—In collibus calc. montis Kalaa, 900 m, 26 May 1928, Font Quer 245-1928 (lectotype BC 11272 ! [https://www.ibb.csic.es/herbari/ JPEG / BC11272 .jpg]; isolectotypes BCN 17956 [photo!], GDA 37278 [photo!], MA 72719 !, MPU 006633 [photo!]*, S 10-1219 [photo!])

Notes:— The BC specimen was labelled as type by Montserrat in 2006, and the GDA specimen as syntype by Vizoso in 2013. Dobignard (2009) cites the MPU specimen as isotype .

1,3 Linum squarrosum var. ericoides Pau in Font Quer, Iter Marocc. 1927 nº 355 (1928), in sched., nom. nud.

= Linum suffruticosum L.

Note:— Carrasco (1975) records this name at the rank of form.

Linum strictum var. genuinum Pau , in sched. ( MA 72803 [photo!] [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00072803]), nom. inval.

= Linum strictum L.

Note:—Invalid name according to Art. 24.3 ( Turland et al. 2018).

1,3 Linum suffruticosum var. maroccanum Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 354 (1928), in sched. = Linum squarrosum subsp. maroccanum (Pau & Font Quer) Mart.Labarga & P.P.Ferrer

Ind. loc.:—“Hab. in declivibus arenaceis c. emporium Sok-et-Tnin (Beni Hadifa)”

Lectotype (designated by Martínez Labarga & Ferrer-Gallego 2020):—In declivibus arenaceis c. emporium Sok-et-Tnin (Beni Hadifa), 1927, Font Quer 354-1927 (lectotype BC 11357 ! [https://www.ibb.csic.es/herbari/ JPEG / BC11357 .jpg]; isolectotypes BCN 17316 [photo!]*, G 00412323 *, GDA 37305 [photo!]*, MA 73166 !*, MPU 006297 [photo!]*)

Notes:—The GDA specimen was labelled as syntype by Vizoso.The MA specimen was identified as L. tenuifolium subsp. suffruticosum ( L.) Litard. by Nicholls in 1983. All isolectotypes were cited by Martínez Labarga & Ferrer-Gallego (2020).