Craspedochaeta chela, Lonsdale & Marshall, 2006

Lonsdale, Owen & Marshall, Stephen A., 2006, Revision of the New World species of Craspedochaeta (Diptera: Clusiidae), Zootaxa 1291 (1), pp. 1-101 : 36-38

Craspedochaeta chela spec. nov.

( Figs. 23 , 49, 50 , 62 , Map 5)

Description ( Fig. 23 )

Body length 3.1 mm. Bristles yellow (including bristles on basal section of costa). Acrostichal bristle present. Scutum yellow with well defined cruciate pattern. Scutellum and laterotergites brown. Pleuron light yellow with brown stripe from dorsal margin of anepisternum to anteromedial face of katepisternum. Coxae white. Legs yellow in part; fore femur brown excluding tip, or yellow with median dorsal spot; mid and hind femora brown excluding yellow apex and wide white base, or entirely yellow; tibiae brown excluding apex and base, or entirely yellow with mid and hind tibiae browned medially; tarsi white. Head yellow in part; frons brown centrally (more so anteriorly); first flagellomere black; dorsal half of face light brown; palpus, lower half of face, buccal cavity and gena black; ocellar spot brown. Abdomen dark brown with tergite 1 yellow and cerci and surstylus white. Wing dusky on distal 1/3, darkest around R 2+3; distance between crossveins distinctly wider than posterior cross­vein.

Female

As described for C. pollostos female except as follows: frons darkest along anterior margin, or evenly dark across entire surface excluding lateral margins (never yellow); palpus yellow; clypeus always brown; face black along dorsal and ventral margins; first flagellomere with faded infuscation on inner­ventral face of first flagellomere (usually) or around entire segment (occasionally); sometimes buccal cavity black and/or mid femur with faint median band.

Male terminalia ( Figs. 49, 50 , 62 )

Externally as described for C. pleuralis except emargination on cerci narrower and surstylus 2/3 as long and narrowest at tip. Hypandrial complex as described for C. pleuralis except posterodistal margin of pregonite more setose and anterior lobe more pronounced.

Etymology: The specific name refers the shape of the surstylus, which resembles a claw.

Distribution: Costa Rica, Ecuador, Guatemala, Peru (Map 5).

Holotype: COSTA RICA. Limón: Valle de la Estrella, R.R. Hitoy Cerere, 100–200m, 30.vii–26.viii.1993, G. Carballo, Malaise (1♂, INBC).

Paratypes: COSTA RICA. Estrella Valley , Pandora , 16.iii.1984, FIT, G.V. Manley & H.F. Howden (1♀, DEBU) , Alajuela: 20km S Upala , F.D. Parker , 1–10.vii.1991 (1♂, EMUS) , 22–30.vi.1991 (1♀, EMUS) , 24.vi.1991 (1♀, EMUS) , 1–15.vii.1991 (1♀, EMUS) , vi.1990 (1♀, EMUS) , 700m, Piedras Blancas , 18.viii.1986, rain forest, L. Masner (1♀, CNCI) , Upala, Dos Rios, Estacion San Gerrardo , 600m, D. Briceno , 10.ix–10.x.2000, Malaise (2♀♀, INBC) , 10.vii.2001, Trampa Malaise (1♀, INBC) , Guanacaste: Guanacaste Conservation Area, Sta. Rosa Sector, Administration Area , ix.1998, Malaise trap, J.J. Sullivan (1♀, USNM) , Finca Montezuma , 3km SE R. Naranjo , 17.vii.1992, F.D. Parker (1♂, EMUS) , 3km SE R. Naranjo, F.D. Parker, 1–5.vi.1992 (1♂, EMUS) , 21–31.viii.1992 (2♂♂, EMUS) , 10–14.viii.1992 (2♂♂, EMUS) , 11–20.vii.1992 (4♂♂, EMUS) , 1–10.viii.1992 (3♂♂ 3♀♀, EMUS) , 1.ii.1993 (1♂, EMUS) , 21–31.vii.1992 (2♂♂, EMUS) , 1–13.vii.1993 (1♂, EMUS) , 13.vii.1992 (2♂♂, EMUS) , 22–24.vii.1992 (1♀ 1♂, EMUS) , 29–31.vii.1993 (2♂♂, EMUS) , 14–15.vii.1993 (1♂, EMUS) , 15–25.i.1993 (1♂, EMUS) , 21–31.viii.1992 (1♂, EMUS) , 21–31.vi.1992 (1♂, EMUS) , 21–30.vi.1992 (1♂, EMUS) , 5–9.vi.1993 (1♂, EMUS) , 4–6.viii.1993 (1♀, EMUS) , 14–20.viii.1993 (2♀♀, EMUS) , 21–31.vii.1992 (10 ♀♀, EMUS; 1♀, DEBU) , 5–9.vii.1993 (1♀, EMUS) , 4.vi.1993 (1♀, EMUS) , 1–10.vii.1992 (1♀, EMUS) , 11–20.ix.1992 (2♀♀, EMUS; 1♀, DEBU), Heredia: La Selva Res. Sta ., 24–30.viii.1988, W.J. Hanson (1♂, EMUS) , 11–17.vi.1986, Hanson & Bohart (1♂, EMUS) , Limón: Amubrio (Talamanca) , 26.vii.1975, W.J. Hanson (1♂, EMUS) , Valle La Estrella , 100m, Malaise, G. Carballo , 8.iv–15.v.1994 (2♀♀ 3♂♂, INBC) , 30.vii–26.viii.1993 (2♀♀, INBC) , 21.v–22.vii.1993 (7♀♀ 6♂♂, INBC) , 100–200m, 14.ii–20.v.1993 (5♀♀ 5♂♂, INBC) , Valle de la Estrella , R.B. Hitoy Cerere, 100m , 8.viii–14.ix.1994, Malaise, G. Carballo (2♀♀, INBC) , P.N. Cahuita, Sector Puerto Vargas, Frente a casa administracion, 22.iv–12.v.2001, Malaise, F. Rojas (1♀, INBC) , R.B. Hitoy Cerere, 300m, 7.vi–8.viii.1994, Malaise, G. Carballo (5♀♀ 3♂♂, INBC) , Puntarenas: Pen. De Osa , Puerto Jiminez, 10m , xi.1991, P. Hanson (1♀, DEBU) , R.F. Golfo Dulce , 24km W Piedras Blancas, 200m , xi.1990, P. Hanson (1♀, DEBU) , Piedras Blancas (24km W, 200m, 8°47’N, 83°15’W) , xi.1990, Malaise trap, P. Hanson (1♂ 1♀, USNM) , Centro Juvenil Tropical , Sendero Mirador, 100m , 5–12.vii.1997, M. Lobo, Malaise trap (2♂♂, INBC) , Osa, Sierpe , Rancho Quemado, 200m , 23.x–29.xi.1993, Malaise, A.L. Marin & A.H. Gutierrez (1♀, INBC) , Osa, Sierpe , Rancho Quemado, 200m , 10.ix–10.x.1993, Manual, A.L. Martin & A.H. Guiterrez (1♀, INBC) , Jiminez, P.N. Corcovado , Estac. Los Patos, Send. Guaymi, 140m , 5.xii.2001 – 23.i.2002, Malaise, K. Caballero (1♀, INBC) . ECUADOR. Her Pto. , Viejo, 50m, rain forest , ii.1980, W. Mason (1♂, CNCI) , Jumboe Rov. , Zamora , 1–2.iv.1965, 1200m, L. Pena (1♂, CNCI) . GUATEMALA. Las Escobas, Izabal Dpto ., 15.vii.1986, L. LeSage (1♂, CNCI) . PERU. Dto. Madre de Dios: Manu Nat. Pk., Cocha Cashu Biol. Station , 30.ix.–5.x.1986, K. Petren (1♂, ROME) .

Additional material examined: COSTA RICA. 100–600m, vii [year unknown], Malaise (1♀, INBC), Puntarenas: Manuel Antonio N.P., 23–28.viii.1986, coastal rainforest, L. Masner (1♀, CNCI) .

Comments: Craspedochaeta chela can be separated from other species with a cruciate notal pattern by a small, hook­like surstylus, and a black first flagellomere in the male. It also differs from similar species in that the face and legs are at least partially brown.

Figures

FIGURES 10–23: Head and thorax, dorsal colouration of the New World Craspedochaeta without an entirely dark notum; 10: C. xanthonotum spec. nov.; 11: C. festiva (Sóos, 1962); 12: C. loreto spec. nov.; 13: C. trivittata (Sóos, 1962); 14: C. weemsi spec. nov.; 15: C. pacaraima spec. nov.; 16: C. candida spec. nov.; 17: C. apsilutea spec. nov.; 18: C. ornata (Hennig, 1938); 19: C. transversa Czerny, 1903; 20: C. synneura (Hennig, 1938); 21: C. pleuralis (Williston, 1896); 22: C. pollostos spec. nov; 23: C. chela spec. nov, male.

FIGURES 47–52: male terminalia, external: Figs. 47–48: Craspedochaeta pollostos spec. nov.; Fig. 47: left lateral; Fig. 48: posterior; Figs. 49–50: C. chela spec. nov.; Fig. 49: left lateral; Fig. 50: posterior; Figs. 51–52: C. weemsi spec. nov.; Fig. 51: left lateral; Fig. 52: posterior.

FIGURES 61–66: Hypandrial complex, left lateral; Fig. 61: Craspedochaeta transversa Czerny, 1903; Fig. 62: C. chela spec. nov.; Fig. 63: C. pollostos spec. nov.; Fig. 64: C. pleuralis (Williston, 1896); Fig. 65: C. weemsi spec. nov.; Fig. 66: C. candida spec. nov.