Smicridea (Rhyacophylax) sanctorum, Gibon & Sganga, 2019
publication ID |
https://doi.org/ 10.11646/zootaxa.4604.1.8 |
publication LSID |
lsid:zoobank.org:pub:23923191-4D90-4BBA-8669-2354791EB3F6 |
DOI |
https://doi.org/10.5281/zenodo.5684872 |
persistent identifier |
https://treatment.plazi.org/id/41D27427-C275-42CC-BA6D-9155058FD362 |
taxon LSID |
lsid:zoobank.org:act:41D27427-C275-42CC-BA6D-9155058FD362 |
treatment provided by |
Plazi |
scientific name |
Smicridea (Rhyacophylax) sanctorum |
status |
sp. nov. |
Smicridea (Rhyacophylax) sanctorum sp. n.
( Figures 1–10 View FIGURES 1–2 View FIGURE 3 View FIGURES 4–6 View FIGURES 7–10 )
Type material. Holotype. Male (disarticulated and mounted on slides), FRENCH GUIANA, Crique Toussaint, 5°20’13.50’’N, 52°59’4.70’’, 34 m a.s.l., 28.ix.2010, F.- M. Gibon leg. ( CBGP) . Paratypes. 6 males (3 on slides, 3 in alcohol), same label data as holotype ( CBGP) .
Diagnosis. Smicridea (R.) sanctorum sp. n. is closely similar to S. (R.) dentisserrata Albino, Pes & Hamada 2011 . In the two species, abdominal segments IX and X are fused in such a way that it is not easy to determine the origin of the observed structures. The tergum IX is positioned dorsally with respect to tergum X ( Fig. 1 View FIGURES 1–2 ), the whole is maintained by a sclerotized Y structure, composed of three branches, ventral, dorsal (Ydo, Fig. 1 View FIGURES 1–2 ) and distal (Ydi, Fig. 1 View FIGURES 1–2 ). The ventral branch arises from the ventral base of sternum IX; the distal branch is the lateroventral margin of tergum X; the dorsal branch may be an extension of one of the two previous. The apex of the distal branch generally corresponds to the distal end of tergum X but in S. dentisserrata and in S. sanctorum sp. n., they are clearly distinct, the distal branch is straight in the new species ( Fig. 2 View FIGURES 1–2 ) and curved inwards in S. dentisserrata (Fig. 22 in Albino et al. 2011). In the two species, the apex of tergum X is mesally divided, each hemitergite is truncated posteriorly in S. dentisserrata (Fig. 22 in Albino et al. 2011) and rounded in the new species ( Fig. 2 View FIGURES 1–2 ). The main diagnostic character is the size and distribution of the spines of the distal branch; they are larger and grouped apically and preapically in the new species, giving to the distal branch the appearance of a mace; they are distributed nearly all along the ventral edge in S. dentisserrata , giving it the appearance of a saw (which is the etymology of the species name). Small but clear morphological differences are also remarkable at the phallic apparatus. Albino et al. (2011) described «2 apico-ventral paramere spines», one anterior, ending in 7 points and one posterior, broad and truncate (Figs. 24, 25 in Albino et al. 2011). In the new species the apico-ventral margin of the phallotheca is elongated, curved dorsad and enlarged, two small spines are inserted latero-ventrally but are hardly visible in dorsal view. As this ventral projection of phallotheca is truncate, the dorsal view is quite similar to that of the posterior spine of S. dentisserrata (Fig. 24 in Albino et al. 2011). On the same dorsal view, the apex of the phallotheca is U-shaped whereas it is smoothly concave in the new species.
Description. Adult male. General coloration (in alcohol) brown. Interocular distance 0.14 mm. Maxillary palp with relative lengths of segments from base: 1, 1.5, 2.5, 2, 7.5 ( Fig. 2 View FIGURES 1–2 ). Wing color pattern of forewings brown, each with perpendicular clear strip at maximum width; hind wing light brown ( Fig. 9 View FIGURES 7–10 ). Length of each forewing 3.80 mm, hind wing 2.90 mm. Anterolateral glandular processes of sternum V long and slender composed of short base, rigid and slightly bent, and distal part, thinner, longer, with wrinkled appearance ( Fig. 6 View FIGURES 4–6 ).
Male genitalia. Segment IX, in lateral view, with Y-shaped internal sclerotization; anterolateral margins concave, anteromesal margins each produced into blunt lobe directed upwards ( Fig. 1 View FIGURES 1–2 ); in dorsal view, tergum subtriangular, posterior margin projected into blunt mesal lobe covered with small spines ( Fig. 2 View FIGURES 1–2 ). Tergum X laterally expanded; in dorsal view divided apicomesally; each hemitergite rounded posteriorly and slightly notched posteromesally, dorsal setal wart slightly oblique and bearing four long setae, and, apically, three tiny setae; ventrolateral margins (distal branch of the Y structure) sclerotized, posteriorly enlarged and covered of spines.
Inferior appendages each two-articulated, setose, basal article long with distal region slightly bent inward; apical article thinner and shorter (less than half as long as basal article) ( Fig. 3 View FIGURE 3 ). Phallic apparatus with tubular phallobase, its basal section slightly enlarged, forming angle of about 120° with distal section; dorsal margin with large, dorsal periphallic cap bearing small spines directed posterad ( Fig. 10 View FIGURES 7–10 ); apicoventral margin of phallotheca elongated, curved dorsad and enlarged with two small spines inserted lateroventrally on each side (paramere spines of Albino et al. 2011); apical portion of ejaculatory duct sclerotized, long and sinuous in lateral view, with two short and sclerotized dorsolateral processes ( Figs. 8, 9 View FIGURES 7–10 ).
Etymology. The name is a reference to the river where the new species was collected, the Crique Toussaint (French for all the saints).
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Smicrideinae |
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