Coramus gedanensis, Skibinska, 2016

Skibinska, Kornelia, 2016, Nannotanyderinae: a new subfamily of Tanyderidae (Diptera), Palaeontologia Electronica 19 (3), pp. 1-16 : 11

Coramus gedanensis sp. nov.

Figures 13 -14

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Etymology. " Gedanensis " given after old name of Gdańsk — capital of amber in Poland.

Material. Holotype No MP/3376 ; Paratype No MP/ 3377 . Both from Upper Eocene Baltic amber. They are deposited in the Institute of Systematics and Evolution of Animals, PAS, Krakow, Poland .

Description. Antennae elongate, delicate, with 16 articles, slightly shorter than the whole body length ( Figures 13 ); scapus short and cylindrical, pedicel short and ovoid, flagellomeres one and two elongate, significantly longer than remaining flagellomeres; flagellomeres 3−14 progressively shorter; flagellomeres 10−14 short and wider than all preceding flagellomeres; mouthparts shorter than the width of head ( Figure 13 ); tibia with short and wide spine; palpus elongate, with apical segment longer than preceding segment; wing length 3.8 mm, width 1.7 mm; vein Sc ends almost opposite one-third length of R 2+3; Rs about 1/5 longer than R 2+3; R 2 twice as long as R 2+3; cross-vein r-m almost in a half of upper margin of d-cell length; vein m-cu situated about 1/10 length of vein M 4; d-cell almost equal in length to vein M 2; vein A 1 strongly angled at the end; anal lobe poorly developed (Figure 14.1); terminalia well developed, with slender and elongated basistylus (0.73 mm long) and gonostylus (0.64 mm long) aedeagus with three phallotrema ( Figures 13.1 , 14.2).

Remarks. Eocene Baltic amber is the youngest deposit in which Nannotanyderinae subfam. nov. is recorded. Wing venation and external built of hypopygium in Coramus gen. nov. is coincident for morphological scope of Nannotanyderinae subfam. nov. Coramus gen. nov. significantly differs from others genera in the wing venation (e.g., length of vein R 2; ending of vein Sc). Aedegal complex is bigger than in other genera, conspicuous, aedeadus with three phallotrema. Members of Nannotanyderinae subfam. nov. were in all probability very few because despite 150 years of research, Coramus gen. nov. is the first to represent this subfamily in Baltic amber. So far two fossil genera are known Macrochile Loew, 1850 and Podemacrochile Podenas, 1997 , which are classifies as subfamily Tanyderinae .

Figures

FIGURE 13. Photographs of Coramus gedanensis gen. et sp. nov. 1, whole specimen (paratype, MP/3377). 2, wing venation (holotype, MP/3376). 3, close-up of head (holotype, MP/3376). 4, close-up of legs (holotype, MP/3376) in lateral view. Scale equals 0.5 mm.