Paramblynotus immaculatus, Liu, Zhiwei, Ronquist, Fredrik & Nordlander, Göran, 2007

Liu, Zhiwei, Ronquist, Fredrik & Nordlander, Göran, 2007, The Cynipoid Genus Paramblynotus: Revision, Phylogeny, And Historical Biogeography (Hymenoptera: Liopteridae), Bulletin of the American Museum of Natural History 2007 (304), pp. 1-151 : 66-67

Paramblynotus immaculatus , new species

FEMALE: Length 1.9 mm. Head and mesosoma black and other body parts dark brown. Wings slightly, evenly ferruginous.

Antenna 13-segmented; flagellum gradually expanded apically; median flagellomeres not constricted. Vertex foveate-reticulate. Eye prominent, distinctly extended laterally beyond outer margin of gena. Distance between posterior ocelli 3.3 times as wide as the distance between posterior ocellus and eye. Ocellar plate slightly raised and not distinctly defined laterally by a carina; the reduced lateral carina of ocellar plate meeting medially above antennal sockets and lined by a row of foveae along interior side; ocellar plate foveate; median frontal carina distinct in lower face, reaching to epistomal sulcus anteriorly and bifurcated posteriorly to delimit a glabrous triangular area beneath anterior ocellus. Upper face glabrate laterally and antennal scrobes glabrous except foveate medially (at the level of anterior ocellus); antennal scrobe defined by carina laterally. Lower face coarsely foveate-reticulate with pubescence. Anterior tentorial pits distinct. Clypeo-pleurostomal sulcus and epistomal sulcus almost form an arch. Clypeus longitudinally carinate. Gena coarsely foveaterugose. Lateral occipital carina extended dorsally to meet lateral carina of antennal scrobe. Occiput glabrous.

Anterior flange of pronotum finely longitudinally carinate. Submedian pronotal depression fused medially. Anterior plate of pronotum glabrous and punctate. Pronotum dorsomedially not distinctly raised; pronotal crest medially not raised into a process. Lateral pronotal carina distinct, meeting pronotal crest dorsomedially. Lateral surface of pronotum foveate-reticulate dorsolaterally and longitudinally rugose lateroventrally. Dorsal pronotal area glabrate-punctate and reaching to end of posterior margin of pronotum. Mesoscutum distinctly arched dorsally and foveate-reticulate. Scutellar sulcus divided by single median longitudinal carina; mesoscutellum foveate-reticulate; posterior margin broadly emarginate in dorsal view. Mesopleural triangle ventrally well defined by smoothly curved carina and with white pubescence. Upper mesopleuron glabrous, distinctly depressed; speculum longitudinally carinate; median longitudinal impression percurrent with three unevenly distributed transverse carinae. Lower mesopleuron glabrous; pubescence sparse above and dense ventrally. Metepisternum depressed and nude above, with an oblique, elevated glabrous area across the middle, and conspicuously pubescent ventrally. Propodeum, including median propodeal area, areolate-reticulate; lateral propodeal carina present only anteriorly and median longitudinal carina percurrent. Nucha glabrous. Rs+M of forewing nebulous, arising from posterior end of basal vein. Marginal cell 2.6 times as long as wide. Bulla on Sc+R 1 absent.

Abdominal petiole 0.36 times as long as wide in lateral view. Relative length of T3–8: 1.3:1.0:2.0:1.2:0:0.4; T3–5 glabrous; T6 finely punctate; T7 almost completely covered by T6 except posterolaterally slightly exposed; T8 punctate with a few foveae. Front and middle legs sparsely punctate with pubescence; hindlegs sparsely punctate with pubes- cence except metacoxa dorsally glabrous and metatibia and metatarsomeres densely punctate with pubescence. Apical teeth of metatibia long, thin, and pointed apically. 1mt/ 2–5mt 5 0.64.

MALE: Unknown.

Paramblynotus immaculatus , antistatus , scalptus , vannoorti , diminutus , angolensis , and minutus form a very distinct, monophyletic clade within the trisetosus group. This clade is characteristic in having (1) very conspicuous anterior tentorial pits; (2) distance between posterior ocelli at least three times as wide as distance between posterior ocellus and eye; (3) speculum longitudinally carinate; (4) T5 of female dorsally expanded and the largest metasomal tergum, T7 largely or entirely covered by T6, T8 exposed; and (5) wings with different color patterns; Rs+M vein of forewing arising from posterior end of basalis.

Paramblynotus immaculatus can be easily separated from the other species of this clade by having (1) wings evenly ferruginous and without darker bands, (2) mesoscutellum posteriorly with a broad emargination, and (3) median frontal carina almost reaching clypeus.

TYPE MATERIAL: HOLOTYPE: ♀, Zaire, Pidigala , 1952-iv-23, H. De Saeger coll. ( NHM).

DISTRIBUTION: Zaire.

ETYMOLOGY: From Latin, im- not, and macula, mark, spot. The name describes the evenly ferruginous wings of the species.