Alaudes californicus Aalbu, Caterino, and Smith

Aalbu, Rolf L., Caterino, Michael S. & Smith, Aaron D., 2018, Studies in the Cnemeplatiini I: A New Subtribe and Revision of the Genus Alaudes Horn (Coleoptera: Tenebrionidae: Pimeliinae: Cnemeplatiini), with Descriptions of New Species from the Southwestern USA and Mexico, Including Notes on Distribution and Biology, The Coleopterists Bulletin 72 (2), pp. 249-268: 253-254

publication ID 10.1649/0010-065X-72.2.249

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Alaudes californicus Aalbu, Caterino, and Smith

new species

Alaudes californicus Aalbu, Caterino, and Smith   , new species ( Figs. 1A, C, E View Fig , 2C View Fig , 4 View Fig , 10 View Fig , 11D View Fig )

Diagnosis. Elytral vestiture always much longer than twice width, composed of thick, strongly recumbent posterior setae ( Fig. 4 View Fig ).

Description. Length 1.47 mm, width 0.63 mm. Color dark brown, luster opaque. Head: Transverse, broader behind, anterior margin broadly rounded, with central aspect straight. Laterally with indentation at eye level. Upper surface transversely, shallowly rounded, upper and lower surfaces completely covered with short, appressed scales. Mentum short, transverse, ligula visible. Maxillary palpi short, apical segment elongate oval. Pronotum: About as broad as elytra, transverse. Surface covered with short, appressed scales except central aspect of base, which is bare and forms a broad hexagon, apex of bare area lacking a small pore. Anterior margin slightly concave, lateral aspect with anterior angles broadly rounded, posteriorly narrowing, very slightly concave before base. Posterior margin bearing 2 pairs of longer, flattened setal clumps, 1 positioned around central bare area with very long, appressed scales and 1 on posterior lateral margin with moderately long, appressed scales. Lateral aspect of pronotum with short, appressed scales. Elytra: Slightly inflated, surface strongly punctate, punctures set in uneven striae on disc, laterally becoming more confused; punctures large, impressed near lateral margins, deeply perforate centrally. intervals mostly bare, with 7 series of moderately long, appressed scales. Anterior margin concave with 2 bare, depressed areas mid-laterally and 1 centrally and 4 alternately elevated areas of concentrated scales corresponding to areas on posterior margin of thorax. Venter: Prosternal process vertically convex between procoxae, mesosternum not excavate; Prosterum bare except central aspect with rounded scales, extending laterally on prosternal process; metasternum with few scattered punctures, extending to first 3 visible abdominal sterna; first visible sternite with 2 shallow groves extending laterally from sternal base; sternal ratios (anterior to posterior midline) 1:0.8: 0.6:0.4:0.7. Legs: Covered with short appressed scales, leg ratios (femur:tibia) pro– 1:0.6; meso– 0.8:0.6; meta– 0.8:0.6. Tarsal length ratios as follows (base to apex): protarsus 1:1:1:1:2; mesotarsus 1:1:1:1:1.7; metatarsus 1.7:1:1.3:3. Genitalia: Male ( Fig. 2C View Fig ) with basal piece longer and broader than parameres, gradually curved basally. Parameres strongly curved basally at midpoint.

Type Material. HOLOTYPE: Male. CALIFORNIA: Kern Co.: Ft. Tejon SP, 34.8756°N,  

118.8953°W, II-9-2008, Caterino & Chatzimanolis. Deposited in SBMN. PARATYPES: (39 specimens from 13 localities) – CALIFORNIA: Fresno Co.: 1 mi. N Cedar Brook , VI-4-1961, manzanita duff, D. J. Burdick (1) CASC   ; 10 mi. NE Auberry , I/31/ 1966, berlese Arctostaphylos   duff, J. Prince (2) FSCA   ; 15 mi. E Squaw Valley , II-26-1977, A. J. Gilbert (3) RLAC   ; same data except (13) CDFA   ; Italian Creek , 6 mi. NE Auberry, V-29 to XI-13-1982, A. J. Gilbert & N. Smith (1) CASC   ; Kern Co.: 5 mi. SE Tehachapi , XI-8-1984, berlese Ceanothus   duff, A. J. Gilbert (1) CDFA   ; Sequoia NF Breckenridge Mt. Rd   ., 35.4644°N, 118.6475°W, XI-8-2007, Caterino & Short (1) SBMN GoogleMaps   ; Sequoia NF, Water Cyn. Rd., 35.0899°N, 118.4941°W, XII- 6-2005, Quercus   litter, Caterino & Chatzimanolis (1) SBMN GoogleMaps   ; Los Angeles Co.: Angeles NF, Hideaway Cyn   ., 34.6993°N, 118.5465°W, V-14- 2007 to V-28-2007, cantharidin pitfall, Caterino & Chatzimanolis (1) SBMN GoogleMaps   ; same locality except VI- 13-2007 to VI-23-2014, unbaited pitfall, Caterino & Chatzimanolis (1) SBMN GoogleMaps   ; same locality except V- 14-2007 to VI-28-2007, cantharidin pitfall, Caterino & Chatzimanolis (4) SBMN GoogleMaps   ; same locality except unbaited pitfall, Caterino & Chatzimanolis (3) SBMN GoogleMaps   ; same locality except VI-13-2007 to VI-23- 2007, unbaited pitfall, Caterino & Chatzimanolis (1) SBMN GoogleMaps   ; Mariposa Co.: 1977, A. J. Gilbert (1) CDFA   ; San Benito Co.: 1.8 mi. SW Idria , III-25 to VI-8- 1981, A. J. Gilbert & N. Smith (1) CDFA   ; San Luis Obispo Co.: Carrizo Plain NM, Selby Campg   ., 35.1227°N, 119.8288°W, IV-2-2004 to IV-25-2004, FIT, Caterino (1) SBMN GoogleMaps   ; Tulare Co.: 5 mi. SW Badger , II-26-1977, Berlese deciduous oak litter, A. J. Gilbert (1) CDFA   .

Other Material Examined. CALIFORNIA: Los Angeles Co.: Angeles NF, Hideaway Cyn., 34.6993°N, 118.5465°W, VI-23-2007 to VII-2-2007, cantharidin pitfall, Caterino & Chatzimanolis (1) SBMN GoogleMaps   .

Distribution. Restricted to the hills along the mid- and lower San Joaquin Valley in California.

Collection Data. Mainly Berlesed oak litter and from Arctostaphylos   (manzanita) and Ceanothus   duff or in pitfall traps. Elevation range 450–1,524 m.


Florida State Collection of Arthropods, The Museum of Entomology