Creophilus, Leach, 1819

Clarke, Dave J., 2011, Testing the phylogenetic utility of morphological character systems, with a revision of Creophilus Leach (Coleoptera: Staphylinidae), Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 163 (3), pp. 723-812 : 755

THE CREOPHILUS COMPLEX

Adult diagnosis: Species of the Creophilus complex can be recognized by the combination of: disc of head and pronotum glabrous; ventroscapal macrosetae absent or reduced; superior marginal pronotal carina separate from inferior marginal carina along entire length, effaced apically ( Fig. 2E, il, sl, Fig. 9J, right and middle arrows); inferior marginal pronotal carina continuous with anterior marginal carina ( Fig. 9J, left and middle arrows); mesepimeron with single marginal ridge ( Fig. 9K, arrow); male sternite IX subtruncate to shallowly emarginate apically, produced into elongate process basally ( Fig. 3Q); median lobe of aedeagus with large apicolateral sclerites ( Fig. 3, as); internal sac without ventral basomedian sclerotized strip (as in Fig. 3K, bms).

Remarks: The following taxa are here included in the Creophilus complex: Creophilus (12 species), Hadrotes crassus , Ht. wakefieldi , Hadropinus fosso r, Liusus (two species), Thinopinus pictus .