Syntemna Winnertz, 1863

BLAGODEROV, VLADIMIR & GRIMALDI, DAVID, 2004, Fossil Sciaroidea (Diptera) in Cretaceous Ambers, Exclusive of Cecidomyiidae, Sciaridae, and Keroplatidae, American Museum Novitates 3433 (1), pp. 1-76 : 32

Syntemna Winnertz, 1863

Syntemna Winnertz, 1863: 767 .

DIAGNOSIS: Mediotergite bare; laterotergites setose; wing membrane with macrotrichia; Sc meets R; radial cell very small, subquadrate; base of M 3+4 and CuA fork proximal to base of M 1 and M 2 fork; segment 7 of abdomen reduced.

TYPE SPECIES: S. morosa Winnertz, 1863: 767 , by monotypy.

COMMENTS: About 20 extant species are known from the Holarctic ( Zaitzev, 1994), with an additional 12 species described from Baltic amber ( Meunier, 1904, 1917a, 1922) and 4 from the Lower and Upper Cretaceous of northeast Asia ( Blagoderov, 1995, 1998a, 2000).

Syntemna fissurata , new species

Figures 39, 40 , Plate 4C

DIAGNOSIS: Sc ends at R at the base of RS; R 4 present; crossvein r­m 0.5Χ length of M3 section; base of fork of M 3+4 and CuA proximal to the base of M 1 and M 2 fork.

DESCRIPTION: Body length = 3.06 mm, wing length = 2.28 mm. Head not fully visible. Flagellum 14­segmented, flagellomeres barrel­shaped, length of each 1.5Χ the width, setose, length of setae about equal to flagellomere width. Thorax: Scutum with strong, long setae in lateral, dorsocentral, and acrostichal rows, with short setulae. Antepronotum, proepisternum, laterotergites, and me­ diotergite setose. Wing membrane with microtrichia and short macrotrichia. Costa ends beyond the tip of R 5, one­fourth distance between tips of R 5 and M 1. Sc meets R just before base of RS. Sections of RS1 and RS2 equal and 1.5Χ length of r­m. Crossvein r­m 0.5Χ length of M3 section. Fork of M 1 and M 2 3.5Χ length of its stem (M3). Base of fork of M 3+4 and CuA proximal to the base of M2 section. Legs: Inner surface of mid tibia with fissurelike sensory pit in apical third. Tarsal claws with two short teeth at the base. Abdomen setose, with 7 visible segments, 7th one very small. Cerci 2­segment­ ed, basal segment roundish, with numerous long trichia; apical segment bacilliform, with length twice the width, bare. Lateral lobes of 8th sternite setulose.

MATERIAL: Holotype TMPD P83.15.3.8, female. Canada: Alberta, Grassy Lake , coll. T. Pike

ETYMOLOGY: The species epithet is the Latin word fissures, meaning ‘‘full of cracks’’, in reference to the state of preservation of the specimen.

COMMENTS: The new species differs from all other known Mesozoic species in having section M3 long, at least twice the length of crossvein r­m.

Figures

Figs. 39–43. Syntemna and Gregikia. 39. S. fissurata, n.sp., holotype TMPD P83.15.3.8, left wing. 40. Female genitalia of the holotype. 41. G. pallida, n.sp., holotype AMNH NJ 117j. 42. Male genitalia dorsally. 43. Male genitalia ventrally.