Maculonola, László, Gyula M., Ronkay, Gábor & Ronkay, László, 2015

László, Gyula M., Ronkay, Gábor & Ronkay, László, 2015, Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species, Zootaxa 4052 (2), pp. 251-296 : 289-294

Maculonola gen. n.

(Plate 6, Figs 7, 8 ; gen. figs 41, 42)

Type species: Meganola apiensis Holloway, 2003 , The Moths of Borneo 18: 30, pl. 1, fig. 20. Type-locality: Borneo, Sarawak, Gunong Mulu NP, Api, 900 m. Holotype: male, in coll. BMNH.

Diagnosis. The habitus of the type-species of the new genus is unique within the whole tribe. The pale greyishbrown forewing lacks all but one crosslines except the oblique, straight, pale, but well visible brownish postmedial line. The other markings of the wing are conspicuous blackish patches on the basal and medial part of the costa, an oblique row of three dots in the terminal area and a larger patch at the middle of the ventral margin. The hindwing venation is typically „quadrifine”, with the M3+CuA1 long stalked.

PLATE VI. Adults of Fragilonola and Maculonola species.

The basic configuration of the male genitalia indicates the closer relationship of Maculonola rather with Nanola than Meganola according to the rather narrow-elongate valval shape and the configuration of harpe. The main autapomorphic character of the new genus is the presence of a rather curious, long, slim, strongly arcuate process of valva arising basally from the sclerotized costal margin, covered densely with short, fine, hair-like cornuti. This character is unknown in other species of the tribe, being eligible to delineate a distinct genus.

Description. Wingspan 19–20 mm, length of forewing 9–10 mm. Head and eyes relatively large, frons, vertex, collar and tegulae covered with pale greyish-brown hair-scales. Forewing elongate, apically rounded, ground colour pale greyish-brown, all but one transverse lines deleted, postmedial line present, as an oblique, straight, pale but well visible, narrow brownish stripe. Forewing costal margin with conspicuous blackish patches at the basal and medial sections and oblique row of three relatively large blackish dots in the terminal area; the mid-section of the ventral margin is marked by a larger patch. Hindwing shining greyish brown, somewhat paler than forewing, discal spot and transverse lines absent.

Male genitalia. Uncus short and broad, with relatively long, acute, fine medial process apically. Tegumen narrow, without medial dilatation; valvae long, rather narrow, apically barely dilated, rounded. Harpe arising at the ventral margin of valva, short ribbon-like with parallel margins, slightly arcuate, apically either with short, pointed process arising at the distal margin, or fully truncate. Valva is armed with long, slim, caudally curved process arising basally at the costal margin, covered densely with short, fine hair-like cornuti; vinculum relatively short, rounded. Aedeagus tubular, relatively short and narrow, without apical carinal process, coecum penis conspicuously short; vesica without cornuti.

Female unknown.

Figures

FIGURES 7 – 9. Genitalia of Meganola and Hampsonola species.