Sadala rufa ( Keyserling, 1880 ), 2022

Rheims, Cristina A. & Jäger, Peter, 2022, Revalidation of the genus Sadala Simon, 1880 with the description of a new genus of Neotropical huntsman spiders (Araneae, Sparassidae), Zootaxa 5135 (1), pp. 1-80 : 17-21

Sadala rufa ( Keyserling, 1880) comb. rest.

Figs 56–61 , 90

Sparassus rufus Keyserling, 1880: 263 , plate 7, fig. 145 (Female holotype from Bogota [4.6492, ‑74.0628], Bogota, Colombia, NHM 1890.7.1.3138, examined); Simon 1897: 35.

Sadala rufa: Simon 1880: 318 .

Olios rufus: Simon 1903a: 1020 ; Petrunkevitch 1911: 502; Jäger 2020: 94, figs 341–348 (misplaced in Olios ).

Olios corallinus Schmidt, 1971: 390 , fig. 2 (holotype female, 2 female paratypes assumingly from Ecuador, introduced to Germany: Hamburg, Lübeck, Mainz, depository uncertain, not examined). Jäger 2020: 81, figs 273–280 (misplaced in Olios ). syn. nov.

Note. The location of the types of O. corallinus is uncertain. Jäger (2020) illustrated an epigyne of what he believes might be a paratype, mounted on a glass slide, deposited in Senckenberg Research Institute (SMF 25776), which is in accordance to that depicted by Schmidt (1971). Based on these illustrations, this species is here considered a junior synonym of S. rufa (Keyserling) .

Additional material examined. PANAMA: Colon: 1♀, Frijoles [9.1733, ‑79.7967], 25 January 1958, A.M. Chickering leg. ( MCZ 69334) ; 1♀, same data as previous specimen ( MCZ 69348) ; 1♀; Panama Oeste : 1♀, Barro Colorado Island [9.1569, ‑79.8466], 3 July 1950, A.M. Chickering leg. ( MCZ 69336) ; 5♀, same locality as previous specimen, 27 June 1934, A.M. Chickering leg. ( MCZ 69333) ; 1♀, same locality as previous specimen, 30 June 1936, A.M. Chickering leg. ( MCZ 69346) ; 1♀, same locality as previous specimen, 18 February 1958, A.M. Chickering leg. ( MCZ 69347) ; 1♀, same locality as previous specimen, Smithsonian Tropical Research Institute , 15 September 1980, M.H. Purdy leg. ( MCZ 69750) ; 2♀, same locality as previous specimen, beach near Barbour Point [9.1605, ‑79.8194], 19 July 1950, A.M. Chickering leg. ( MCZ 69335) ; 1♂, same locality as previous specimen, 7–9 July 1936, A.M. Chickering leg. ( MCZ 69149) ; 1♂, same locality as previous specimen, 23–30 June 1939, A.M. Chickering leg. ( MCZ 69131) ; 1♂, same locality as previous specimen, 2 July 1954, A.M. Chickering leg. ( MCZ 69146) ; 1♂, same locality as previous specimen 1–31 August 1936, A.M. Chickering leg. ( MCZ 69142) ; 1♂, same locality as previous specimen, 20 May 1964, A.M. Chickering leg. ( MCZ 69126) ; 1♂, same locality as previous specimen, 16 June–15 July 1934, A.M. Chickering leg. ( MCZ 69150) ; 1♂, same locality as previous specimen, June 1950, A.M. Chickering leg. ( MCZ 69145) ; 1♂, same locality as previous specimen, 08 July 1939, A.M. Chickering leg. ( MCZ 69138) ; 1♂, same locality as previous specimen, 31 July 1954, A.M. Chickering leg. ( MCZ 69135) .

Diagnosis. Males of S. rufa resemble those of S. keyserlingi ( Figs 24–26 ) and S. velox ( Figs 72–74 ) by the palp with embolus, arising from tegulum at 6 o’clock position ( Fig. 57 ). They are distinguished from both species by the cymbium lacking hook-shaped dorsal proximal projection ( Fig. 58 ) and conductor base extending from central tegulum to embolus base ( Fig. 57 ) (cymbium with hook-shaped dorsal proximal projection and conductor base not extending to embolus base in S. keyserlingi and S. velox ). Females are distinguished from the remaining congeners by the epigyne with TP short, more than 7 times wider than long and arising anteriorly from MS (roughly as wide as wide as long or longer than wide and arising medially or posteriorly from MS in other species) ( Fig. 59 ).

Description. Male (MCZ 69149): Total length 11.5. Prosoma: 5.5 long, 5.0 wide. Opisthosoma: 6.1 long, 3.6 wide. Eyes: diameters: 0.40, 0.34, 0.30, 0.34; interdistances: 0.28, 0.10, 0.50, 0.42, 0.22, 0.20. Legs: I: 31.9 (8.6, 2.8, 8.8, 9.3, 2.4): II: 34.7 (9.3, 3.1, 9.7, 9.7, 2.4); III: 23.4 (7.0, 2.4, 6.1, 6.1, 1.8); IV: 26.7 (8.0, 2.2, 7.1, 7.4, 2.0). Spination follows the generic pattern except patella III: p0. Palp: RTA two times longer than wide, finger-like, pointing antero-ventrad in retrolateral view; PTA triangular, as long as wide; subtegulum visible prolaterally in ventral view; tegulum oval; conductor more than five times longer than wide, anteriorly fanned ( Figs 56–58 ).

Redescription. Female (MCZ 69333): total length 20.8. Prosoma: 8.8 long, 8.1 wide. Opisthosoma 11.7 long, 7.3 wide. Eyes: diameters: 0.60, 0.46, 0.34, 0.48; interdistances: 0.48, 0.30, 0.78, 0.70, 0.50, 0.36. Legs: I: 37.1 (10.5, 4.5, 9.9, 10.0, 2.2); II: 39.3 (11.3, 4.5, 10.5, 10.7, 2.3); III: 29.3 (9.1, 3.7, 7.4, 7.0, 2.1); IV: 32.0 (9.8, 3.5, 7.9, 8.6, 2.2). Spination follows the generic pattern. Epigyne: EF slightly wider than long with transversal anterior groove; MAB embedded in EF; MS rectangular, wider than long; LL touching posteriorly; ( Fig. 67 ). Vulva: FW postero-mediad; GP short, rounded, arising from ducts at second turn; SP elongate; FD laterad ( Figs 60–61 ).

Variation. Males (n = 9): total length 11.2–12.9; prosoma length 4.9–5.7; femur I length 8.2–9.9. Females (n = 9): total length 18.1–22.8; prosoma length 8.0–9.2; femur I length 8.3–10.7.

Distribution. Panama and northwestern South America ( Colombia and possibly Ecuador) ( Fig. 90 ).

Figures

FIGURES 56–61. Sadala rufa (Keyserling) comb. nov. 56–58 Male (MCZ 69149), left palp (56 prolateral, 57 ventral, 58 retrolateral); 59–61. Female (MCZ 69333) (59 epigyne, 60 vulva, 61 schematic course of internal duct system). ag—anterior groove; C—conductor; E—embolus; FD—fertilization duct; ft—first turn of duct system; GP—glandular projection; LL—lateral lobe; MAB—muscle attachment bands; MS—median septum; PTA—prolateral tibial projection; RTA—retrolateral tibial apophysis; SP—spermathecae; st—second turn of duct system; ST—subtegulum; TP—triangular projection. Scale lines: 1 mm.

FIGURES 89–90. Distribution maps for Sadala spp. 89 S. tabatinga spec. nov., S. velox Simon, S. nanay spec. nov., S. punicea Simon 90 S. rufa (Keyserling), S. kaiabi spec. nov., S. keyserlingi Simon, S. nigristernis Simon, S. yuyapichis spec. nov.

FIGURES 24–29. Sadala keyserlingi Simon 24–26 Male (INPA 4563), left palp (24 prolateral, 25 ventral, 26 retrolateral); 27–29 Female (MNHN 2459, syntype) (27 epigyne, 28 vulva, 29 schematic course of internal duct system). C—conductor; DpP—dorsal proximal projection; E—embolus; FD—fertilization duct; ft—first turn of duct system; GP—glandular projection; LL—lateral lobe; MS—median septum; TP—triangular projection; PTA—prolateral tibial projection; RTA—retrolateral tibial apophysis; SP—spermathecae; st—second turn of duct system; ST—subtegulum. Scale lines: 1 mm.

FIGURES 72–77. Sadala velox Simon 72–74 Male (MZSP 33349), left palp (72 prolateral, 73 ventral, 74 retrolateral); 75–77 Female (IBSP 9361) (75 epigyne, 76 vulva, 77 schematic course of internal duct system). C—conductor; DpP—dorsal proximal projection; E—embolus; FD—fertilization duct; ft—first turn of duct system; GP—glandular projection; LL—lateral lobe; MS—median septum; TP—triangular projection; PTA—prolateral tibial projection; RTA—retrolateral tibial apophysis; SP— spermathecae; st—second turn of duct system; ST—subtegulum. Scale lines: 1 mm.

FIGURES 62–67. Sadala tabatinga spec. nov., female 62–64 (ex MNHN 2459) (62 epigyne, 63 vulva, 64 schematic course of internal duct system); 65–67 (AMNH, paratype) (65 epigyne, 66 vulva, 67 schematic course of internal duct system). Scale lines: 1 mm. FD—fertilization duct; ft—first turn of duct system; GP—glandular projection; LL—lateral lobe; MS—median septum; st—second turn of duct system; TP—triangular projection.