Dima fthiotidensis Schimmel & Platia, 2008

Mertlik, Josef, Németh, Tamás & Kundrata, Robin, 2017, Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula, Zootaxa 4220 (1), pp. 1-63 : 17-18

Dima fthiotidensis Schimmel & Platia, 2008

Figs 35–42 , 130–134 , 166 , 186–187 , 229 , 245 .

Dima fthiotidensis Schimmel & Platia, 2008: 576 .

Dima fokidensis Schimmel & Platia, 2008: 575 , syn. n.; Platia 2013: 99. Dima scutellaris Platia, 2010: 30 , syn. n.

Type depositories. Holotype of D. fthiotidensis , ♂ (MRSN); 7 paratypes of D. fthiotidensis , ♀♀ (MCSB, MRSN, PCGP, PCRS); holotype of D. fokidensis , ♀ (MCSB); holotype of D. scutellaris , ♀ (PCGP).

Type locality. Greece: Phthiotis , Kallidromo Mt., 990 m ( D. fthiotidensis ) ; Greece: Fokida province , O. Iti, Pira, 1150 m ( D. fokidensis ) ; Greece: Phocis, Giona Mts. , vers. N str. Kaloskopi Stromi, 1170 m ( D. scutellaris ).

New material. GREECE: distr. Fthiótida , Vardoússia Mts., road Plátanes–Gramméni Oxiá, beechwood, 1447 m (38°46'57.47"N, 21°58'06.86"E), 14.VI.2012, 12 ♂♂, 11 ♀♀ J. Mertlik leg. ( PCJM) ; dtto, 1 ♀ (PCRK); dtto, 32 ♂♂, 32 ♀♀, V. Dušánek leg. (PCVD); dtto, 9 ♂♂, 12 ♀♀, P. Brůha leg. (PCPB); dtto, 18 ♂♂, 26 ♀♀, B. Zbuzek leg. (PCBZ); dtto 11 ♂♂, 8 ♀♀, M. Samek leg. (PCMS); Fthiotis env., Mt. Kallidromo, 3 km SW Pavliani, oakpine forest, singled at night, 23.V.2011, 2 ♂♂, 2 ♀♀, leg. A. Kotán, A. Márkus & T. Németh ( PCRK) ; dtto, 1 ♀ (HNHM); Ftiotida , Iti Mts., Kastania, 1500 m, 4.VI.2006, 1 ♂, 1 ♀, Pavel Štěpánek leg. ( PCJM) .

Diagnosis. Dima fthiotidensis is a medium-sized species (body length: 10.5–14.5 mm) with matt, densely punctate pronotum ( Figs 130–134 ), more or less decumbent pubescence on the pronotal lateral margin, moderately densely pubescent pronotum and elytra, and the tongue-like, convex scutellum with steeply declined frontal margin ( Fig. 166 ). This species is similar in the above mentioned characters to D. evritaniensis , however the latter differs in the slightly finer punctation of pronotum, longer hairs on pronotum and elytra, semi-erect pubescence on the pronotal sides ( Figs 26–28 , 126–127 ), and longer apical lobe of paramera ( Fig. 183 ). The remaining species occurring in the surrounding mountains have a shiny pronotum with sparser punctation. Additionally, D. pelionensis sp. nov. and D. etoliensis have a flattened scutellum, and D. hladilorum and D. zbuzeki sp. nov. have a more robust and longer apical lobe of paramera, respectively ( Figs 190 , 221 ). The sclerotized spines of bursa copulatrix as in Fig. 229 .

Intraspecific variability. The specimens from Mt. Iti have a darker body. Several specimens (including the holotype of D. scutellaris ) have slightly wrinkled to rugose pronotum with denser punctation ( Fig. 132 ). The scutellum is more or less convex, in some cases with a longitudinal shiny keel running from the base towards apex. Most specimens have pubescence near anterior pronotal angles oriented latero-posteriorly, but a female from Mt. Iti has pubescence oriented medio-posteriorly.

Distribution. Greece (Mt. Kallidromo, Mt. Iti, Vardousia Mts., Giona Mts., Oxia Mts.; Fig. 245 ).

Remark. We were not able to study the holotype of D. fthiotidensis (deposited in Torino, Italy), however we examined four paratypes, recently collected specimens from the type locality (Mt. Kallidromo; Fig. 38 ), and the specimens from the surrounding mountain ranges to investigate the limits of this species ( Figs 39–41 ). Platia (2010) described D. scutellaris from a single female from Mt. Giona, with a raised scutellum and wrinkled pronotum with denser punctation. This specimen, however, fits the definiton of D. fthiotidensis in all aspects, and what is more, we found similarly wrinkled (teratological?) female specimens in the Vardousia Mts. (see the similar cases in D. assingi / D. peloponnensis and D. fialai ). Therefore, we herein synonymize Dima scutellaris Platia, 2010 under Dima fthiotidensis Schimmel & Platia, 2008 .

Schimmel & Platia (2008) described D. fokidensis from three female specimens. The holotype comes from Mt. Iti and paratypes from Mt. Pelion, which is located more than 80 km away from the holotype locality. This holotype was found to be conspecific with D. fthiotidensis (described in the same paper as D. fokidensis ; Fig. 36 ) and both paratypes represent a new species, described here as D. pelionensis sp. nov. ( Fig. 95 ). Therefore, we herein synonymize Dima fokidensis Schimmel & Platia , 2 0 0 8 under Dima fthiotidensis Schimmel & Platia, 2008 .

Figures

FIGURE 245. The distribution of Dima species in the Central Greece.

FIGURES 30 – 44. Habitus images of Dima species. 30 — D. fialai sp. nov., paratype, female; D. florinensis Platia, 2012, 31 — holotype, female, 32 — non-type male from Varnous Mts., 33 — non-type female from Varnous Mts., 34 — non-type female from Galičica Mts.; D. fthiotidensis Schimmel & Platia, 2008, 35 — paratype, female, 36 — holotype of D. fokidensis Schimmel & Platia, 2008, female, 37 — holotype of D. scutellaris Platia, 2010, female, 38 — non-type male from Kallidromo Mts., 39 — nontype male from Vardoussia Mts., 40 — non-type female from Vardoussia Mts., 41 — non-type female from Vardoussia Mts.; D. giachinoi Schimmel & Platia 2008, 42 — paratype, female, 43 — non-type specimen, male, 44 — non-type specimen, female. Not to scale.

FIGURES 119 – 130. Pronotal details of Dima species. 119 — D. arndti Platia, 2013, male; 120 — D. assingi Schimmel & Platia, 2008, female; 121 — D. bruhai sp. nov., holotype, male; 122 — D. dalmatina Küster, 1844, male; 123 — D. dusaneki sp. nov., holotype, male; 124 — D. elateroides Charpentier, 1825, male; 125 — D. etoliensis Platia, 2012, holotype, female; 126 — D. evritaniensis Schimmel & Platia, 2008, paratype, female; 127 — D. evritaniensis, male; 128 — D. fialai sp. nov., holotype, male; 129 — D. florinensis Platia, 2012, female; 130 — D. fthiotidensis Schimmel & Platia, 2008, male. Not to scale.

FIGURES 131 – 142. Pronotal details of Dima species. 131 — D. fthiotidensis Schimmel & Platia, 2008, paratype, female; 132 — D. scutellaris Platia, 2010, holotype, female; 133 — D. fthiotidensis, male; 134 — D. fthiotidensis, holotype of D. fokidensis Schimmel & Platia, 2008; 135 — D. giachinoi Schimmel & Platia 2008, female; 136 — D. hirtipennis Platia, 2011, holotype, female; 137 — D. hladilorum Schimmel, 1987, male; 138 — D. kozufensis sp. nov., paratype, female; 139 — Dima lebenbaueri Schimmel & Platia 2008, holotype, male; 140 — Dima lebenbaueri, non-type specimen, female; 141 — D. macedonica Schimmel, 1993, holotype, female, 142 — D. macedonica, female paratype of D. hladilorum from Galičica Mts. Not to scale.

FIGURES 163 – 171. Scutellum-shape of Dima species. 163 — D. bruhai sp. nov.; 164 — D. macedonica Schimmel, 1993; 165 — D. raineri Wurst, 1997. Elytral details of Dima species. 166 — D. fthiotidensis Schimmel & Platia, 2008; 167 — D. hirtipennis Platia, 2011; 168 — D. macedonica Schimmel, 1993; 169 — D. olympica Meischnigg, 1934; 170 — D. parnonensis Schimmel & Platia, 2008; 171 — D. pecoudi Fleutiaux, 1943. Not to scale.

FIGURES 182 – 191. Aedeagus images of Dima species. 182 — D. elateroides Charpentier, 1825, non-type specimen from Serbia; 183 — D. evritaniensis Schimmel & Platia, 2008, non-type specimen from Timfristos Mts.; 184 — D. fialai sp. nov., holotype; 185 — D. florinensis Platia, 2012, non-type specimen; 186 — D. fthiotidensis Schimmel & Platia, 2008, non-type specimen from Kallidromo Mts.; 187 — D. fthiotidensis, non-type specimen from Vardoussia Mts.; 188 — D. giachinoi Schimmel & Platia 2008, non-type specimen; 189 — Dima sp. from Mani; 190 — D. hladilorum Schimmel, 1987, non-type specimen; 191 — D. kozufensis, sp. nov., holotype. Not to scale.

FIGURES 222 – 233. The sclerotized spines of bursa copulatrix of Dima species. 222 — D. arndti Platia, 2013, paratype; 223 — D. bruhai sp. nov., paratype; 224 — D. dusaneki sp. nov., paratype; 225 — D. elateroides, Charpentier, 1825, non-type specimen; 226 — D. etoliensis Platia, 2012, holotype; 227 — D. fialai sp. nov., paratype; 228 — D. florinensis Platia, 2012, holotype; 229 — D. fthiotidensis Schimmel & Platia, 2008, holotype of D. scutellaris Platia, 2010; 230 — D. giachinoi Schimmel & Platia 2008, paratype; 231 — D. hirtipennis Platia, 2011, holotype; 232 — D. kozufensis sp. nov., paratype; 233 — D. neumanni Platia, 2013, non-type specimen. Not to scale.

FIGURES 15 – 29. Habitus images of Dima species. Dima dalmatina Küster, 1844, 15 — syntype, male, 16 — non-type male from Vosac, Croatia, 17 — non-type female from Vosac, Croatia, 18 — non-type male from Rumija Mts., Montenegro, 19 — nontype female from Rumija Mts., Montenegro; D. dusaneki sp. nov., 20 — holotype, male, 21 — paratype, female; D. elateroides Charpentier, 1825, 22 — non-type male from Serbia; 23 — non-type male from Slovenia, 24 — non-type female from Croatia; 25 — D. etoliensis Platia, 2012, holotype, female; D. evritaniensis Schimmel & Platia, 2008, 26 — paratype, female, Panetoliko Mts., 27 — non-type male from Timfristos Mts., 28 — non-type female from Timfristos Mts.; 29 — D. fialai sp. nov., holotype, male. Not to scale.

FIGURES 212 – 221. Aedeagus images of Dima species. 212 — D. raineri Wurst, 1997, paratype of D. macedonica; 213 — D. raineri, holotype of D. pindosensis Schimmel & Platia, 2008; 214 — D. schimmeli Platia & Gudenzi, 2009, holotype; 215 — D. schimmeli, non-type specimen; 216 — D. schnitteri Platia, 2013, holotype; 217 — D. vailatii Schimmel & Platia, 2008, holotype; 218 — D. vailatii, non-type specimen; 219 — D. vailatii, holotype of D. riesei Platia, 2010; 220 — D. vonickai sp. nov., holotype; 221 — D. zbuzeki sp. nov., holotype. Not to scale.

FIGURES 90 – 104. Habitus images of Dima species. Dima pelikani sp. nov., 90 — paratype, female, 91 — paratype, male; D. pelionensis sp. nov., 92 — holotype, male, 93 — paratype, female, 94 — paratype, female; 95 — paratype, female, paratype of D. fokidensis; D. raineri Wurst, 1997, 96 — holotype, male, 97 — non-type specimen, male, 98 — non-type specimen, female, 99 — holotype of D. pindosensis Schimmel & Platia, 2008, male, 100 — male paratype of D. macedonica from Mt. Tomorr, Albania; 101 — Dima sp. from Librazhd, Albania; D. schimmeli Platia & Gudenzi, 2009, 102 — holotype, male, 103 — non-type specimen, male, 104 — non-type specimen, female. Not to scale.