Dima evritaniensis Schimmel & Platia, 2008

Mertlik, Josef, Németh, Tamás & Kundrata, Robin, 2017, Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula, Zootaxa 4220 (1), pp. 1-63 : 14

Dima evritaniensis Schimmel & Platia, 2008

Figs 26–28 , 126–127 , 183 , 245 .

Dima evritaniensis Schimmel & Platia, 2008: 573 .

Type depositories. Holotype, ♂ (MRSN), 3 paratypes: ♀♀ (MCSB, PCGP, PCRS).

Type locality. Greece: Evritania province , Panetoliko Mts., 620 m, Proussos env.

Redetermined material. One of the paratypes of D. evritaniensis , a male from Mt. Ossa ( PCGP; Fig. 87 ), belongs under D. pecoudi Fleutiaux, 1944 .

New material. GREECE: distr. Evritania, Timfristos Mts. [Veluchi], 1 ♀, without further data ( PCVD) ; distr. Evritania , Timfristos Mts., Karpeníssi env., firry wood, 1446 m (38°55'25.22"N, 21°48'33.22"E), 13. VI.2012, 4 ♂♂, 14 ♀♀, J. Mertlik leg. ( PCJM) ; dtto, 1 ♂ (PCRK); dtto, 8 ♂♂, 28 ♀♀, V. Dušánek leg. (PCVD); dtto, 3 ♂♂, 11 ♀♀, P. Brůha leg. (PCPB); dtto, 4 ♂♂, 8 ♀♀, B. Zbuzek leg. (PCBZ); distr. Evritania , Timfristos Mts., Karpeníssi env., firry wood, 1446 m (38°55'25.22"N, 21°48'33.22"E), 8. VI.2015, 8 ♂♂, 9 ♀♀, J. Mertlik leg. ( PCJM) ; dtto, 4 ♂♂, 7 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 2 ♂♂, 14 ♀♀, P. Brůha leg. (PCPB); dtto, 6 ♂♂, 1 ♀, T. Németh leg. (PCRK); dtto, 1 ♂ (HNHM); dtto, 9. VI.2015, 1 ♀, J. Mertlik leg (PCJM).

Diagnosis. Dima evritaniensis is a medium-sized species (11.3–13.5 mm) with dense pronotal punctuation, semi-erect setae on the pronotal lateral margin, and elytra with short, dense, decumbent pubescence ( Figs 26–28 , 126–127 ). This species is morphologically very like D. fthiotidensis , based on the body shape, size, and matt pronotum with dense, fine punctation ( Figs 35–41 ). Dima fthiotidensis has slightly coarser punctation of pronotum ( Figs 130–134 ), shorter hairs on pronotum and elytra, decumbent to semi-erect pubescence on the pronotal sides (in most cases), and shorter apical lobe of paramera ( Figs 186–187 ). Dima evritaniensis differs from the remaining species in the central Greece by its matt body surface and pronotum with denser, finer punctation ( Figs 26–28 , 126–127 ). Additionally, D. pelionensis sp. nov. has the sides of pronotum with decumbent setae in the anterior two thirds (semi-erect setae along its whole length in D. evritaniensis ; Figs 126–127 , 152–153 ), D. etoliensis has the scutellar frontal margin flattened, declined gradually to the plane of elytra (convex, steeply declined in D. evritaniensis ; visible from the lateral view), D. hladilorum has more robust apical lobe of paramera ( Fig. 190 ), broader scutellum, which is less convex basally (in most cases; scutellum longer and more convex basally in D. evritaniensis ), and D. zbuzeki sp. nov. is a smaller species with much longer apical parameral lobe ( Fig. 221 ).

Intraspecific variability. In some cases, elytra or whole body is paler than in a typical form.

Distribution. Greece (Evritania: Panetoliko Mts., Timfristos Mts.; Fig. 245 ).

Remarks. We did not have an opportunity to study the holotype of D. evritaniensis , which is deposited in the Natural History Museum in Torino (MRSN). The collection is closed and not available for study (M. Garsena, pers. comm.). We studied the female paratype with the same locality data as the holotype (i.e. Panetoliko Mts., Proussos env.; from MCSB; Fig. 26 ) and this is conspecific with the specimens from Timfristos Mts. ( Figs 27–28 ). Based on the comparison of the high-resolution holotype male photograph (kindly provided us by R. Schimmel) and fresh male specimens collected in Timfristos Mts. in 2012 and 2015, these populations share similar body size and shape, pronotal punctation and pubescence, and length of antennae. According to the original description of D. evritaniensis ( Schimmel & Platia 2008) , the holotype has short antennomeres II and III, with antennomere III slightly longer than II, and both combined as long as antennomere IV. However, all specimens which we examined here (including the paratype) have antennomere II slightly elongate, subtriangular, narrower than antennomere III, antennomeres II and III of subequal length, both combined longer than antennomere IV. Additionally, the parameral apical lobes of D. evritaniensis figured in the original description (Schimmel & Platia 2 008) are shorter than in the specimens from Timfristos, however, it might be caused by the different angle and/or the low-quality photograph. Therefore, the real shape of paramera might differ from the one originally figured (compare e.g. the shape of paramera of D. assingi holotype in the original description and herein; Schimmel & Platia (2008); Fig. 175 ).

Figures

FIGURE 245. The distribution of Dima species in the Central Greece.

FIGURES 15 – 29. Habitus images of Dima species. Dima dalmatina Küster, 1844, 15 — syntype, male, 16 — non-type male from Vosac, Croatia, 17 — non-type female from Vosac, Croatia, 18 — non-type male from Rumija Mts., Montenegro, 19 — nontype female from Rumija Mts., Montenegro; D. dusaneki sp. nov., 20 — holotype, male, 21 — paratype, female; D. elateroides Charpentier, 1825, 22 — non-type male from Serbia; 23 — non-type male from Slovenia, 24 — non-type female from Croatia; 25 — D. etoliensis Platia, 2012, holotype, female; D. evritaniensis Schimmel & Platia, 2008, 26 — paratype, female, Panetoliko Mts., 27 — non-type male from Timfristos Mts., 28 — non-type female from Timfristos Mts.; 29 — D. fialai sp. nov., holotype, male. Not to scale.

FIGURES 119 – 130. Pronotal details of Dima species. 119 — D. arndti Platia, 2013, male; 120 — D. assingi Schimmel & Platia, 2008, female; 121 — D. bruhai sp. nov., holotype, male; 122 — D. dalmatina Küster, 1844, male; 123 — D. dusaneki sp. nov., holotype, male; 124 — D. elateroides Charpentier, 1825, male; 125 — D. etoliensis Platia, 2012, holotype, female; 126 — D. evritaniensis Schimmel & Platia, 2008, paratype, female; 127 — D. evritaniensis, male; 128 — D. fialai sp. nov., holotype, male; 129 — D. florinensis Platia, 2012, female; 130 — D. fthiotidensis Schimmel & Platia, 2008, male. Not to scale.

FIGURES 182 – 191. Aedeagus images of Dima species. 182 — D. elateroides Charpentier, 1825, non-type specimen from Serbia; 183 — D. evritaniensis Schimmel & Platia, 2008, non-type specimen from Timfristos Mts.; 184 — D. fialai sp. nov., holotype; 185 — D. florinensis Platia, 2012, non-type specimen; 186 — D. fthiotidensis Schimmel & Platia, 2008, non-type specimen from Kallidromo Mts.; 187 — D. fthiotidensis, non-type specimen from Vardoussia Mts.; 188 — D. giachinoi Schimmel & Platia 2008, non-type specimen; 189 — Dima sp. from Mani; 190 — D. hladilorum Schimmel, 1987, non-type specimen; 191 — D. kozufensis, sp. nov., holotype. Not to scale.

FIGURES 75 – 89. Habitus images of Dima species. Dima olympica Meschnigg, 1934, 75 — non-type specimen, male, 76 — non-type specimen, female; D. orientalis sp. nov., 77 — holotype, male, 78 — paratype, female, 79 — paratype, female; 80 — Dima sp. from Xanthi, Greece; D. parnonensis Schimmel & Platia 2008, 81 — holotype, female, 82 — non-type specimen, male, 83 — non-type specimen, female, 84 — non-type specimen, male; D. pecoudi Fleutiaux, 1944, 85 — non-type specimen, male, 86 — non-type specimen, female, 87 — paratype of D. evritaniensis, male, 88 — non-type specimen, teratological male; 89 — D. pelikani sp. nov., holotype, male. Not to scale.

FIGURES 30 – 44. Habitus images of Dima species. 30 — D. fialai sp. nov., paratype, female; D. florinensis Platia, 2012, 31 — holotype, female, 32 — non-type male from Varnous Mts., 33 — non-type female from Varnous Mts., 34 — non-type female from Galičica Mts.; D. fthiotidensis Schimmel & Platia, 2008, 35 — paratype, female, 36 — holotype of D. fokidensis Schimmel & Platia, 2008, female, 37 — holotype of D. scutellaris Platia, 2010, female, 38 — non-type male from Kallidromo Mts., 39 — nontype male from Vardoussia Mts., 40 — non-type female from Vardoussia Mts., 41 — non-type female from Vardoussia Mts.; D. giachinoi Schimmel & Platia 2008, 42 — paratype, female, 43 — non-type specimen, male, 44 — non-type specimen, female. Not to scale.

FIGURES 143 – 154. Pronotal details of Dima species. 143 — D. macedonica Schimmel, 1993, male; 144 — D. marvani Mertlik & Dusanek, 2006, holotype, male; 145 — D. moraveci Schimmel & Platia, 2008, holotype, female; 146 — D. neumanni Platia, 2013, female; 147 — D. olympica Meschnigg, 1934, male; 148 — D. orientalis sp. nov., holotype, male; 149 — D. parnonensis Schimmel & Platia, 2008, female; 150 — D. pecoudi Fleutiaux, 1944, male; 151 — D. pelikani sp. nov., holotype, male; 152 — D. pelionensis sp. nov., paratype, female, 153 — D. pelionensis sp. nov., paratype, female paratype of D. fokidensis; 154 — D. raineri Wurst, 1997, female. Not to scale.

FIGURES 212 – 221. Aedeagus images of Dima species. 212 — D. raineri Wurst, 1997, paratype of D. macedonica; 213 — D. raineri, holotype of D. pindosensis Schimmel & Platia, 2008; 214 — D. schimmeli Platia & Gudenzi, 2009, holotype; 215 — D. schimmeli, non-type specimen; 216 — D. schnitteri Platia, 2013, holotype; 217 — D. vailatii Schimmel & Platia, 2008, holotype; 218 — D. vailatii, non-type specimen; 219 — D. vailatii, holotype of D. riesei Platia, 2010; 220 — D. vonickai sp. nov., holotype; 221 — D. zbuzeki sp. nov., holotype. Not to scale.

FIGURES 172 – 181. Aedeagus images of Dima species. 172 — D. arndti Platia, 2013, holotype; 173 — D. arndti, non-type specimen; 174 — Dima sp. from Crete; 175 — D. assingi Schimmel & Platia, 2008, holotype; 176 — D. assingi, non-type specimen from Kilini Mts.; 177 — D. peloponnensis Schimmel & Platia 2008, holotype; 178 — D. bruhai sp. nov., holotype; 179 — D. dalmatina Küster, 1844, non-type specimen from Sutorman, Montenegro; 180 — D. dalmatina, non-type specimen from Vosec, Croatia; 181 — D. dusaneki sp. nov., holotype. Not to scale.