Slaterocoris atratus (Uhler)Slaterocoris croceipes (Uhler)

Schwartz, Michael D., 2011, Revision And Phylogenetic Analysis Of The North American Genus Slaterocoris Wagner With New Synonymy, The Description Of Five New Species And A New Genus From Mexico, And A Review Of The Genus Scalponotatus Kelton (Heteroptera: Miridae: Orthotylinae), Bulletin of the American Museum of Natural History 2011 (354), pp. 1-290: 100-107

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http://doi.org/ 10.1206/354.1

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http://treatment.plazi.org/id/0395E50B-FFEE-FF84-DC69-FC48FD85A004

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Tatiana

scientific name

Slaterocoris atratus (Uhler) Slaterocoris croceipes (Uhler)
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Slaterocoris atratus (Uhler)  

Figures 12 View Fig , 19 View Fig ; plate 1 View Plate 1 ; map 4

Stiphrosoma atrata, Uhler, 1894: 268   (orig. desc.).

Strongylocoris atratus: Van Duzee, 1921: 135   (disc., in part); Knight, 1938: 7, fig. 9 (male genit.).

Slaterocoris atratus: Kelton, 1968: 1135   , fig. 23 (disc.); Knight, 1970: 246, fig. 42 (disc.); Henry and Wheeler, 1988: 448 (catalog); Schuh, 1995: 197 (catalog).

DIAGNOSIS: This is the only species of the apache   group with dense, relatively long, brown vestiture and male genitalia with a bifurcate tergal process, fanlike apical region of the right paramere, and endosomal spicule with basal portion of the dorsal lobe narrow. Distinguished from non– apache   -group species with dense brown vestiture by the completely black coloration; in S. maculatus   the head is partially pale. The tergal process is small and bifurcate, the endosomal spicule occupies the entire base of dorsal lobe, and the basal portion of right paramere is prominent and short. The species of the hirtus   group, containing the only other species with dense brown vestiture, differ from S. atratus   by the board tergal process with a solitary point, the usually bifurcate base of the dorsal lobe of the endosomal spicule, and the long basal portion of the right paramere without prominent basal lobe.

REDESCRIPTION: Male: Moderately large, obovate; length 4.50–4.90, width 2.13–2.40 (pl. 1). COLORATION: Pale on antennal segment I distally, femur at apex, tibia distally; hind tibia missing (fig. 12). VESTI- TURE: Dense, moderately long, suberect, brown to black simple setae. STRUCTURE: Frons slightly striate; antennal segment II just longer than foretibia. GENITALIA: As in species group, but differing as follows. Pygophore: Aperture with short, bifurcate tergal process (fig. 19E). Endosomal spicule: Distal portion of ventral lobe gradually expanded with some marginal serration on lateral surface; dorsal lobe narrow basally, distally flat and relatively narrow with weak marginal serration (fig. 19A). Right paramere: Apical region flat, fanlike with three sharp points; basal lobe bifurcate (fig. 19D).

Female: Unknown.

HOST: Unknown.

DISTRIBUTION: Known from the two specimens of the type series found in San Julio, which is apparently located in the middle of Baja California, Mexico (map 4).

DISCUSSION: Uhler (1894) described S. atratus   based on ‘‘several specimens’’ acquired from the CAS. The introduction of his paper indicated that the ‘‘types’’ were returned to the CAS. Van Duzee (1921) provided notes on Slaterocoris species   (as Strongylocoris   ) retained in the CAS based presumably on specimens determined by Uhler. He mentioned that an ‘‘examination of Uhler’s types [of atratus   ] show that they were males.’’ Knight, during a trip to the CAS in 1931, illustrated the parameres of S. atratus   based on observations of ‘‘ two type specimens conserved there’’ ( Knight, 1938: 7, fig. 9, 1970: 266, fig. 46). Kelton (1968: 1133, fig. 23B, C) illustrated the endosomal spicule and right paramere of S. atratus   (identical to that documented by Knight) and designated the lectotype. For this current study four male syntypes of Stiphrosoma atrata   , were located, two in the CAS and two in the CNC; no specimens of the type series were found in the USNM. The lectotype designation by Kelton (1968) was ambiguous—apparently no specimen was labeled. I have labeled the male intended by Kelton as the lectotype and returned it to the USNM. It is dissected with parts of the genitalia in a vial affixed to the pin and bears the labels: ‘‘San Julio, Lower Cal., Mex.,’’ ‘‘PR Uhler Collection,’’ ‘‘atratus,’’ ‘‘USNM_ENT AMNH_PBI 00122271,’’ ‘‘ LECTOTYPE Stiphrosoma atrata Uhler, 1894   , desig. L.A. Kelton, 1968, labeled by M.D. Schwartz, 2010 [red label].’’ The remaining three syntype males are now recognized as paralectotypes.

All California specimens in the CAS determined as S. atratus   by Van Duzee (1921) (denoted by attached identification labels in Van Duzee’s handwriting) are S. solidaginis   (see Other Specimens Examined under S. solidaginis   ). I am in agreement with the assessment of Knight (1970) that none of the specimens (except the two former syntypes) retained in the CAS are S. atratus   . Non-California specimens of S. atritibialis   , S. breviatus   , S. pallidicornis   , and S. stygicus   incorrectly identified as S. atratus   by Van Duzee (1921) are housed in the CAS and several other institutions (see Other Specimens Examined under those species). Two fifth-instar nymphs (CAS) have the same locality label as the lectotype; however, I have refrained from assigning these immatures to S. atratus   because they lack the relatively long, suberect dorsal vestiture of the adult. These glabrous nymphs are perhaps S. solidaginis   , and would extend the range of that species 360 km further south in Baja California.

Both Kelton (1968) and Knight (1970) suggested that S. atratus   has a relationship with S. alpinus   , one of three species placed in the hirtus   -species group by the phylogenetic analysis. Based strictly on external characters, where the vestiture is dense, relatively long, and brown, the frons is striate, and the tibiae are black, this would be a reasonable conclusion. However, the structure of the tergal process (small with two apical points), right paramere (with a short basal region and prominent basal lobe), and endosomal spicule (entire dorsal lobe and relatively short distal portion of the ventral lobe) are clearly of the form diagnostic for the apache   -group species. Species of the hirtus   group have the

tergal process with a broad base and one point, the right paramere with long, parallel apical spines and long basal region, and the dorsal lobe of the endosomal spicule bifurcate.

LECTOTYPE: MEXICO: Baja California: San Julio [29.656 N 114.766666 W], Apr 1889, Charles D. Haines, 18 ( AMNH _ PBI 00122271 View Materials ) ( USNM). GoogleMaps  

ADDITIONAL SPECIMENS: Same label data as lectotype GoogleMaps   ; paralectotypes, 18 (00122272) ( USNM), 28 (00242726, 00242727) ( CAS)   .

Slaterocoris croceipes (Uhler)  

Figures 12 View Fig , 20A, 20E View Fig , 21–23 View Fig View Fig View Fig ; plates 1 View Plate 1 , 4F; map 3, cover

Stiphrosoma croceipes Uhler, 1893: 373   (orig. desc.).

Strongylocoris croceipes: Van Duzee, 1921: 136   (disc., in part).

Slaterocoris croceipes: Kelton, 1968: 1128   , fig. 13 (disc.); Knight, 1970: 253, fig. 14, (disc.); Henry and Wheeler, 1988: 449 (catalog); Schuh, 1995: 197 (catalog).

DIAGNOSIS: Distinguished from other apache   -group species by the variable bright yellowish to orange-brown distal portions of the femora, the yellowish orange tibiae (fig. 12); the sparse, recumbent, moderately long, golden brown vestiture (pl. 4F); and the male genitalia with the apex of the basal lobe in the right paramere usually serrate (fig. 21A, D–I) and the endosomal spicule with distal portion of the ventral lobe somewhat expanded and moderately serrate (fig. 22A, B, D, E, G). Slaterocoris croceipes   differs from S. sheridani   in antennal segment II longer than the foretibia, mostly yellowish to orange-brown femora, and untwisted dorsal lobe of the endosomal spicule. The more sparse dorsal vestiture, mostly pale antennal segment I, pale yellowish orange femur, shorter hemelytron, and the twisted dorsal lobe of the endosomal spicule (fig. 60D, E) of S. sparsus   will distinguish it from S. croceipes   ; both species can have the femora mostly pale, although the femora in S. sparsus   are usually more extensively pale yellowish orange than in S. croceipes   , whose palest specimens have a more orange or reddish-brown tint. When antennal segment I is pale in the female, S. croceipes   , S. sheridani   , and S. sparsus   can be difficult to separate. Antennal segment II is on average longer and the costal margin is somewhat curved, and the pale portions of the femora are pale orange-brown in S. croceipes   . In the latter two species antennal segment II is generally shorter, the costal margin is arcuate, and the pale portions of the femora have a more yellow tint.

REDESCRIPTION: Male: Moderately large, elongate; length 3.15–5.28, width 1.58–1.90 (pl. 1). COLORATION: Antennal segments I and II and base of femur black, variable distal portion of femur pale orange-brown, tibia yellow (fig. 12). VESTITURE: Consisting of sparse, recumbent to suberect, moderate length, golden brown setae (pl. 4F). STRUCTURE: Costal margin of hemelytron curved to subparallel; frons smooth (fig. 20A); antennal segment II longer than foretibia; mesothoracic spiracle and metathoracic scent-efferent system as in figure 20E. GENITALIA: Tergal process: Short, with one sharp point. Endosomal spicule: Distal portion of ventral lobe variable, somewhat narrow (fig. 22F) or slightly expanded (fig. 22B), usually with moderately dense marginal serration, length short to moderately long; dorsal lobe parallel sided, predominantly wide distally (fig. 22E), sometimes narrow (fig. 22F), terminating with apical serration of variable number and size; lateral edge usually without conspicuous medial and basal spine (fig. 22A–H). Right paramere: Number of apical spines variable, basal lobe usually with apex serrate (fig. 21A–I).

Female: Moderately large, obovate; length 3.45–4.20, width 1.60–2.15 (pl. 1). COLOR- ATION: Sometimes antennal segment II dusky to pale. STRUCTURE: Costal margin curved. GENITALIA: As in species group except: Ventral labiate plate: Strongly produced, base triangular, moderately wide, apex entire or cleft (fig. 23A, B). Interramal lobe: Dorsomedial margin with short process (fig. 23D, E).

HOSTS: Many specimens, from a large number of collection events, indicate that Ericameria nauseosa   ( Asteraceae   ) is a host plant. Other probable host plants are E. parryi var. vulcanica   and Chrysothamnus viscidiflorus   . A number of specimens were also collected from species of Artemisia   ( A. filifolia   and A. tridentata   ) and Chrysothamnus   . The latter generic identification could pertain to either Chrysothamnus   of Erica- meria considering the current state of botanical taxonomy. The asteraceous genus, Aster   , and three non-asteraceous plants are considered to be sitting records.

DISTRIBUTION: From the mountain ranges of southern California that border the western portion of the Mojave Desert, north along the eastern foothills of the Sierra   Nevada, isolated ranges of the Great Basin in Nevada and Utah, lower elevations of northwestern Colorado, and scattered populations in the Great Plains of Colorado, South Dakota, and Texas (map 3).

COLLECTION SUMMARY: 719 specimens from 112 collection events with the earliest record from April in California and the majority of specimens from June to early August.

DISCUSSION: The original description of croceipes ( Uhler, 1893)   stated that it was ‘‘based upon two males specimens from Los Angeles, Cal., which were sent to me by Mr. D.W. Coquillett.’’ Kelton received these two males, deposited in the USNM, and designated one the lectotype ( Kelton, 1968), stating that it bore the No. 69539. At the beginning of the current project both males were found at the CNC without labels indicating the lectotype designation or the USNM type number. To correct the ambiguous lectotype designation, I have chosen the male in the best condition, and applied two labels to it in order to stabilize the concept of croceipes: AMNH   _PBI 00122110 and LECTOTYPE 8 Stiphrosoma croceipes Uhler, 1893   , desig. by M.D. Schwartz, 2010 [red label]. I also labeled the other syntype [right hemelytron missing] to recognize it as the paralectotype (AMNH_PBI 00070120).

In the original description, ‘‘several specimens were collected at American Fork, [UT], June 22 [91]’’ and included in croceipes   , by Uhler (1893). During the course of this current project seven specimens (38, 4♀) fitting the original description have been located—six in the USNM and one female in the CAS. The CAS specimen has an additional printed label, ‘‘ Heideman [sic] Collector’   ’; I assume Heidemann collected all seven specimens. Van Duzee (1921) probably included the latter specimen in his treatment of the genus (see Other Specimens Examined). The seven Utah specimens are not considered to be syntypes and along with the lectotypes and paralectotype have been returned to the museum of provenance   .

Easternmost specimens are somewhat smaller than western specimens. The size difference is easily apprehended by comparing the length of antennal segment II in both sexes; those from Colorado, South Dakota, and Texas are shorter (0.80–0.938, 0.78– 0.81 ♀) than those from California, Nevada, and Utah (1.00–1.318, 1.06–1.23 ♀). The hemelytron length in males is conspicuously longer in California specimens (total body length 4.13–5.28) than in those from the remainder of the range (3.15–4.10).

As noted for S. apache   , variation also occurs in the number and conformation of the apical spines and apical serration on the basal lobe of the right paramere in S. croceipes   (cf. fig. 21A–I). Variable structure was observed during this study in specimens from California, Nevada, Texas, and Utah. The dorsal and ventral lobes of the endosomal spicule (fig. 22A–H) are also variable across the distribution of the species. The variation of the width of the dorsal lobe was deemed great enough to code S. croceipes   as polymorphic for this character (37-0, 1; table 2).

Specimens of S. croceipes   from Oelrich, South Dakota, and near Lefors, Texas, in the eastern reaches of the distribution, have femoral coloration as observed in the specimens from the western portion of the range in California, Nevada, and Utah. The proportions of basal black, medial orangebrown, and apical yellow coloration regions on the femora are variable across the distribution and within populations. The extent of black varies from a basal onefourth to one-half, with the reciprocal extent of orange-brown varying from a medial three-quarters to one-half. The orange-brown color also grades from a darkest tone to apical yellow in a variable fashion.

LECTOTYPE: USA: California: Los Angeles Co.: Los Angeles [34.052226 N 118.242786 W], May , Coquillett Collection , 18 ( AMNH _ PBI 00122110 View Materials ) ( USNM). GoogleMaps  

OTHER SPECIMENS EXAMINED: USA: Arizona: Coconino Co.: Grand Canyon, south Rim, 36.054446 N 112.138616 W, 03 Jun 1930, R.L. Usinger, Artemisia sp.   , 18 (00119454), 2♀ (00119455–00119456) (UCB). Gila Co.   : nr Roosevelt Lake, 33.673236 N 111.149136 W, 686 m, 27 Jun 1947, H. and M. Townes, 1♀ (00134619) (USNM). California: Inyo Co.   : 2 mi E of Big Pine, 37.1656 N 118.252326 W,

03 Jun 1975, J.D. Pinto, 1♀ (00119871) (UCR). 5 mi E of Big Pine, 37.17096 N 118.21146 W, 1341 m, 11 Jun 1969, W.F. Barr, Aster sp.   , 1♀ (00122347) (UID). 9 mi NE of Big Pine, 37.2576 N 118.173016 W, 1920 m, 09 Jun 1966, W. Gagne, 18 (00119435), 1♀ (00276763) (UCB). 10 mi NE of Big Pine, 37.26116 N 118.15086 W, 2103 m, 09 Jun 1966, C.W.O’Brien, 98 (00119436, 00276754–00276761), 1♀ (00276762) (UCB); 09 Jun 1966, W. Gagne, Chrysothamnus sp.   , 18 (00119437), Chrysothamnus sp.   , 48 (00276773–00276776), 3♀ (00276777–00276779) (UCB). 30 mi N of Saline Valley Lake, 36.677136 N 117.768166 W, 335 m, 04 May 1974, J.D. Pinto, 28 (00119865, 00120016), 1♀ (00120017) (UCR). China Lake Naval Weapons Center, Coso Mountains, Silver Peak Springs on west side, 35.736 N 117.6346 W, 1996 m, 27 Jul 1999, M. Gates, 1♀ (00119866) (UCR). Deep [Spring] Valley, Antelope Spring[s], 37.330246 N 118.090436 W, 1698 m, 02 Jun 1987, W.F. Barr, Salix sp.   ( Salicaceae   ), 1♀ (00122425) (UID). Independence, 36.802786 N 118.199176 W, 17 Jun 1937, Norman W. Frasier, 1♀ (00119445) (UCB). Olancha, Owens Valley, 36.28066 N 118.01696 W, 1122 m, 20 May 1937, J.W. Johnson   , 18 (00119439) (UCB). Panamint Mountains, 36.40036 N 117.27976 W, 1512 m, 24 May 1937, Norman W. Frasier, 1♀ (00119444) (UCB). Saline Valley Road, 19.3 mi N of Hwy 190, 36.59316 N 117.62946 W, 919 m, 04 Jun 1974, J.D. Pinto, 38 (00119864, 00120013– 00120014), 1♀ (00120015) (UCR). Kern Co.: 11 mi NW of Tehachapi on Rt 58, 35.244666 N 118.585746 W, 25 Apr 1983, G.M. Stonedahl, Ericameria nauseosa   , 78 (00108331, 00170227–00170232), 3♀ (00170233–00170235), 2 immatures (00170236–00170237) (AMNH). 20 km W of Wofford Heights on Rt 155, 35.706946 N 118.67646 W, 1500 m, 26 Jul 1999, M.D. Schwartz, Chrysothamnus viscidiflorus   , 18 (00110996), 2♀ (00112105– 00112106) (CNC). E of Caliente on Caliente Creek Rd, Twin Oaks, 35.307026 N 118.42336 W, 836 m, 22 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Ericameria nauseosa   , det. A. Sanders UCR140644 [PBI_CAL04-H126], 108 (00088461–00088470), 14♀ (00088471– 00088484) (AM), 18 (00108333), 1♀ (00322727), 3 immatures (00322728–00322730) (AMNH). Kernville, 35.754726 N 118.424446 W, 06 Jun 1952, G.L. Smith, 1♀ (00119443) (UCB); 09 Jun 1952, unknown, 1♀ (00276781) (UCB); 09 Jun 1959, G.L. Smith, 1♀ (00276782) (UCB). Mount Pinos, 34.82346 N 119.13316 W, 2438 m, 01 Jul 1965, unknown, 1♀ (00119446) (UCB). Red Rock Canyon, 35.3256 N 117.949726 W, 18 May 1937, E.P. Van Duzee, Asteraceae   , 18 (00242481) (CAS). S of Bodfish on Caliente- Bodfish Road, 35.560896 N 118.50536 W, 1094 m, 22 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Fremontodendron californicum (Torr.) Corville   ( Sterculiaceae   ), det. Field ID, 1♀ (00322745) (AMNH). S of Tehachapi on Tehachapi Willow Springs Rd, 35.055026 N 118.36966 W, 1329 m, 21 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Ericameria nauseosa   , det. A. Saunders UCR 140653 [PBI_CAL04-H92], 88 (00091708, 00088439– 00088445), 7♀ (00088446–00088452) (AM). S of Tehachapi on Tehachapi Willow Springs Rd, 35.068896 N 118.39596 W, 1393 m, 21 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Ericameria nauseosa   , det. Field ID, 98 (00322736–00322744), 5♀ (00322746–00322750) (AMNH). W of Mojave on Oak Creek Road, 35.038486 N 118.28076 W, 1055 m, 21 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Ericameria nauseosa   , det. A. Sanders UCR140653, [PBI CAL04-H92], 68 (00137262, 00322731–00322735), 10♀ (00322751–00322760) (AMNH). Los Angeles Co.: 1 mi S of Gorman, 34.781656 N 118.851676 W, 13 Jun 1964, A.G. Raske, 38 (00119410, 00276769–00276770), 2♀ (00276771–00276772) (UCB). Big Rock Creek Wash, Mohave Desert, 34.66646 N 117.87266 W, 770 m, 12 May 1944, Timberlake, Chrysothamnus sp.   , 18 (00119862), 2♀ (00120018–00120019) (UCR). Largo Vista Rd 3.1 mi S of Rt 18, SE of Llano, 34.452516 N 117.76516 W, 1275 m, 17 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Ericameria nauseosa   , det. A. Saunders UCR 140595 [PBI_CAL04-H2], 178 (00088414–00088429, 00091710), 9♀ (00088430–00088438) (AM), 228 (00107881, 00108136, 00108853, 00137236–00137238, 00322993–00323008), 17♀ (00108135, 00108970, 00322987–00322992, 00323009– 00323017) (AMNH). Los Angeles, 34.052226 N 118.242786 W, Coquillett Collection, 1♀ (00122111) (USNM); May, Coquillett Collection, 18 (00070119) (USNM), 18 paralectotype [abdomen missing] (00070120) (USNM). Rt 18 at W 263 St, 34.498336 N 117.6696 W, 1033 m, 17 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Ericameria nauseosa UCR   140595 [PBI CAL04-H2], 98 (00137263, 00322687–00322694), 23♀ (00322695– 00322717) (AMNH). SE of jct Pearblossom Hwy (Rt 18) and 263rd Street 3300N, 34.50006 N 117.668666 W, 1035 m, 19 May 2000, M.D. Schwartz, Ericameria nauseosa   , 118 (00108334, 00170213–00170222), 4♀ (00170223–00170226) (AMNH). Mono Co.: Mono Lake, 1 mi S of Lee Vining, 37.944466 N 119.102146 W, 2064 m, 20 Aug 1987, D.A. Rider, 1♀ (00242954) (DAR). Oasis, 37.487716 N 117.915656 W, 1538 m, 27 May 1954, M. Cazier, 28 (00108332, 00170238), 1♀ (00108895) (AMNH). Riverside Co.: 2.5 mi NW of Desert Springs, 33.98866 N 116.52776 W, 408 m, 25 May 1955, Timberlake, Ericameria nauseosa   , 78 (00119860 [cat UCRC ENT 32032 View Materials ], 00120004 [cat UCRC ENT 32025 View Materials ]–00120009 [cat UCRC ENT 32035 View Materials ]), 3♀ (00120010 [cat UCRC ENT 32026 View Materials ]–00120012 [cat UCRC ENT 32031 View Materials ]) (UCR). Round Valley, 33.805026 N 116.662796 W, 2754 m, 27 Jul 1947, R.H. Beamer, 2♀ (00117852–00117853) (KU). Upper Palm Canyon, 14 mi SE of Keen Camp Summit, 33.63776 N 116.64526 W, 1341 m, 08 Aug 1964, E.I. Schlinger, 18 (00119868) (UCR). Vandevanter Flat, San Jacinto Mountains, 33.783896 N 116.957786 W, 04 May 1940, unknown, Asteraceae   , 18 (00242480), 1♀ (00242479) (CAS); 04 Jun 1940, unknown, Chrysothamnus sp.   , 18 (00119438), Chrysothamnus sp.   , 3♀ (00276764–00276766) (UCB); 04 Jun 1940, H.T. Reynolds, Chrysothamnus sp.   , 1♀ (00276767) (UCB); 04 Jun 1940, C.D. Michener, 1♀ (00276768) (UCB); top of Berdoo Canyon, 33.83386 N 116.07336 W, 1128 m, 14 Jun 1963, E.I. Schlinger, 1♀ (00119872) (UCR). San Bernardino Co.: 1 mi N of X of FS Rd 2N93 and Hwy 38, 34.21846 N 116.81626 W, 2350 m, 24 Jun 1997, J. Heraty, 18 (00119870) (UCR). 5.4 mi W of Lucerne Valley, 34.443896 N 117.06166 W, 935 m, 13 May 1978, J.D. Pinto and R.T. Schuh, Ericameria nauseosa   , 668 (00108329, 00170290–00170313, 00322857–00322897), 37♀ (00169981, 00322898–00322933) (AMNH); 13 May 1978, John D. Pinto, Chrysothamnus sp.   , 158 (00119983–00119997), 7♀ (001198756, 00119998–00120003) (UCR). Apple Valley, 34.531396 N 117.282786 W, 15 May 1955, W.R.M. Mason, 18 (00112099) (CNC). Barton Flats, 34.172516 N 116.863086 W, 1997 m, 22 Jul 1981, G. Gordh, 18 (00119863) (UCR). Cajon Pass Summit on I-15, 34.298896 N 117.455566 W, 02 Jul 1949, Flock, 18 (00119869) (UCR). Camp Baldy, 34.295286 N 116.913926 W, 11 Jul 1950, R. DeNoble, 28 (00119440, 00276780) (UCB). Ironwood Campground, 34.304166 N 117.012226 W, 2130 m, 22 Aug 1999, M. Gates, 18 (00119867) (UCR). NW of Pioneertown, T1N R4E S12, 34.18446 N 116.52696 W, 1280 m, 03 May 1994, M. Cooperband, 18 (00119861) (UCR). Phelan, Rt 138 at Phelan Road, 34.425316 N 117.61746 W, 1310 m, 16 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Ericameria nauseosa   , det. A. Saunders UCR 140595 [PBI_CAL04-H2], 38 (00088453–00088455), 5♀ (00088456– 00088460) (AM), 238 (00107875, 00322718– 00322721, 00322934–00322951), 41♀ (00169985, 00322722–00322726, 00322952– 00322986) (AMNH). Tulare Co.: 21 mi W of Rt 395 toward Kennedy Meadows, 36.009446 N 118.503086 W, 1875 m, 01 Jul 1980, R.T. Schuh, Ericameria parryi var. vulcanica   , 88 (00108335, 00108843– 00108849), 3♀ (00108850–00108852) (AMNH). Colorado: Delta Co.   : Delta   , 38.742226 N 108.068336 W, 27 Jun 1938, R. Bauer, 2♀ (00134594–00134595) (USNM); 27 Jun 1938, U. Lanham, 1♀ (00134596) (USNM). Mesa Co.   : Collbran, 39.240166 N 107.959736 W, 1829 m, 08 Aug 1922, E.R. Kalmbach, 1♀ (00164886) (USNM). Glade Park, 38.994746 N 108.741176 W, 2105 m, 09 Aug 1931, F.C. Hottes, 18 (00070130), 1♀ (00134615) (USNM). Moffat Co.: Massadona, 40.25286 N 108.64986 W, 1768 m, 01 Jul 1931, R.H. Beamer, 1♀ (00117855) (KU). Maybell, 40.517756 N 108.080286 W, 1805 m, 30 Jun 1931, R.H. Beamer, 1♀ (00117854) (KU). Morgan Co.: 5 mi N of Wiggins, 40.303036 N 104.072226 W, 12 Jun 1979, D.A. Polhemus, 28 (00116819, 00116953) (JTP). Weld Co.: 5 mi NW of Roggen, 40.218616 N 104.43866 W, 1461 m, 16 Jul 1983, D.A. and J.T. Polhemus, Artemisia filifolia Torr.   , 7♀ (00116958–00116964) (JTP). 5 mi W of Roggen, 40.16756 N 104.466266 W, 1585 m, 16 Jul 1983, R.T. Schuh, D.A. and J.T. Polhemus, Artemisia filifolia Torr.   , 18 (00108396), 1♀ (00108396) (AMNH). 10 mi E of Roggen along I-25, 40.20186 N 104.25496 W, 1426 m, 11 Jul 1979, D.A. Polhemus, 18 (00116727), 1♀ (00116965) (JTP). Fort Lupton, 40.084726 N 104.81256 W, 10 Jun, unknown, 18 (00134368) (USNM); 10 Jun 1910, unknown, 1♀ (00134614) (USNM). Keenesburg, 40.108336 N 104.519446 W, 11 Jun 1961, B.H. Poole, 3♀ (00112102–00112104) (CNC). Unknown co.: Colo 2304, unknown, 1♀ (00134357) (USNM). Nevada: Carson City Co.: 4 mi S of Carson City, 39.11626 N 119.78156 W, 1463 m, 30 May 1930, W.F. Chamberlain, Artemisia sp.   , 28 (00119044–00119045), 1♀ (00119046) (TAMU). Clark Co.: Charleston Peak, 36.272226 N 115.694176 W, 2286 m, 22 Jul 1982, J.T. Polhemus, 18 (00116728), 3♀ (00116954–00116956) (JTP). Kyle Canyon, 36.262746 N 115.606686 W, 2090 m, 26 Jun 1966, W.F. Barr, 18 (00122345), 1♀ (00122346) (UID). Lee Canyon, Charleston Peak, 36.354516 N 115.64056 W, 2286 m, 20 Jul 1982, J.T. Polhemus, 1♀ (00116957) (JTP). Esmeralda Co.   : 2 mi W of Lida, 37.45136 N 117.514916 W, 1935 m, 08 Jun 1966, W. Gagne, Light Trap, 18 (00119441), 38 (00276790–00276792) (UCB). 3 mi W of Lida, 37.458336 N 117.551856 W, 2073 m, 07 Jul 1966, C.W. O’Brien, 18 (00276783), 2♀ (00276784–00276785) (UCB); 07 Jul 1966, L. and C.W. O’Brien, 18 (00119411) (UCB). 6 mi N of Goldfield, 37.795356 N 117.234726 W, 1615 m, 08 Jun 1966, W. Gagne, 18 (00119039) (TAMU), 18 (00276793) (UCB). Lincoln Co.: Beaver Dam State Park, 37.518136 N 114.076366 W, 1555 m, 18 Jun 1986, J.B. Knight and K.R. Helms, 1♀ (00108896) (AMNH). Pioche, 37.929696 N 114.452216 W, 1852 m, 09 Jul 1965, W.F. Barr, Penstemon sp.   ( Scrophulariaceae   ), 28 (00122344, 00122439), 1♀ (00122440) (UID). Lyon Co.: 7 mi N of Sweetwater Summit on Rt 338, Toiyabe National Forest, 38.613626 N 119.240546 W, 1865 m, 11 Jul 1980, R.T. Schuh and G.M. Stonedahl, Ericameria nauseosa   , 28 (00108336, 00170314), 9♀ (00170315–00170323) (AMNH). 8 mi N of Sweetwater summit on highway 22, Toiyabe National Forest, 38.630466 N 119.260976 W, 1811 m, 11 Jul 1980, G.M. Stonedahl, Ericameria nauseosa   , 4♀ (00108337, 00170324–00170326) (AMNH). Nye Co.: 2 mi E of Tonopah, 38.067226 N 117.192446 W, 1890 m, 07 Jun 1966, W. Gagne, 28 (00119442, 00276786), 3♀ (00276787– 00276789) (UCB). 5.5 mi S of Belmont on Hwy 82, 38.514016 N 116.932316 W, 1981 m, 13 Jul 1980, G.M. Stonedahl, Artemisia tridentata   , 1♀ (00108897) (AMNH). Mercury, 18 M, 36.660566 N 115.993616 W, 20 Jun 1965, H.H. Knight and J. Merino, Ericameria nauseosa   , 1♀ (00119067) (TAMU). Mercury, 401 M, 36.660566 N 115.993616 W, 18 Jun 1965, H.H. Knight and J. Merino, Ericameria nauseosa   , 18 (00112100) (CNC), 28 (00119063–00119064), 2♀ (00119065– 00119066) (TAMU). White Pine Co.   : Lehman Caves, Great Basin National Park, 39.005556 N 114.219446 W, 24 Jun 1966, W. Gagne, J. Haddock, 18 (00119429) (UCB). South Dakota: Fall River Co.: Angostura Dam, Hot Springs, 43.344636 N 103.439396 W, 967 m, 06 Jul 1968, A.T. Howden, 1♀ (00115908) (CNC). Oelrichs, 43.178896 N 103.230286 W, 1022 m, 28 Jun 1973, L.A. Kelton, Artemisia sp.   , 28 (00110993, 00112135), Artemisia sp.   , 178 (00115880– 00115896), 11♀ (00115897–00115907) (CNC). Texas: Gray Co.: 3 mi E of Lefors, 35.44146 N 100.79696 W, 878 m, 29 May 2002, J.C. Schaffner, 58 (00118222, 00119058– 00119060, 00245354), 2♀ (00119061, 00245355) (TAMU). Hemphill Co.: 7.5 mi N of Canadian River on Hwy 83, 35.983616 N 100.465276 W, 18 May 2002, E.G. Riley, 1♀ (00119062) (TAMU). Utah: Beaver Co.: Milford, 39.396916 N 113.017896 W, 1513 m, 07 Jun 1969, W.F. Chamberlain, 18 (00119047) (TAMU). Garfield Co.: Capitol Reef National Park, Grand Wash, Cobab Canyon Trail, 38.156 N 111.166676 W, 1827 m, 21 Jun 1983, R.T. Schuh, M.D. Schwartz, Ericameria nauseosa   , 28 (00108390–00108391), 2♀ (00108392– 00108393) (AMNH). Jct. Rts 95 & 276, S of Hanksville, T32S R12E, 38.025056 N 110.570986 W, 1494 m, 18 Jun 1983, R.T. Schuh and M.D. Schwartz, Ericameria nauseosa   , 18 (00108394), 1♀ (00108395) (AMNH). Iron Co.   : Bear Valley, 37.926626 N 112.607346 W, 2295 m, 09 Jun 1966, G.F. Knowlton, 18 (00134611) (USNM). Cedar City, 37.675216 N 113.050686 W, 1798 m, 22 Jun 1933, O.H. Swezey, 58 (00242463– 00242467), 10♀ (00242468–00242477) (CAS). Kanarraville, 37.538896 N 113.183336 W, 09 Jun 1960, G.F. Knowlton, 1♀ (00164887) (USNM). Juab Co.: Deep Creek Mountains, 39.843096 N 113.822886 W, 1753 m, Jun 1928, Vasco M. Tanner, 2♀ (00134616, 00164882) (USNM). Kane Co.: Whitehouse, Utah Exp. Sta., 37.078326 N 111.889896 W, 1316 m, 20 Jun 1933, G.F. Knowlton, 18 (00112036) (CNC), 18 (00134612) (USNM). Millard Co.: Skull Rock Pass, 53 mi WSW of Delta   , 39.033296 N 113.339696 W, 1596 m, 12 Jun 1992, W.F. Chamberlain, 18 (00119048) (TAMU). Sevier Co.: 11 mi W of Sevier on I- 70, 38.575016 N 112.461326 W, 1999 m, 15 Jul 1980, R.T. Schuh, G.M. Stonedahl, Chrysothamnus viscidiflorus   , 18 (00170271), 19♀ (00169986, 00170272–00170289) (AMNH); 15 Jul 1980, G.M. Stonedahl, Chrysothamnus viscidiflorus   , 78 (00170253–00170258, 00107900), 12♀ (00170259–00170270) (AMNH). Clear Creek Narrows Summit on Rt 4, 38.581566 N 112.256786 W, 2243 m, 15 Jun 1980, R.T. Schuh and G.M. Stonedahl, 58 (00107907, 00170239–00170242), 10♀ (00170243–00170252) (AMNH). Monroe, 38.636 N 112.126 W, 25 Jul 1906, unknown, 1♀ (00134618) (USNM). Salina, 38.948286 N 111.857326 W, 1577 m, 28 Jun 1954, G.L. Nielsen, 18 (00070117) (USNM). Tooele Co.: 4 mi NE of Camel Back Mountain [Camels Back Ridge], 40.185866 N 112.916546 W, 1326 m, 07 Jun 1954, P.D. Ashlock, Ericameria nauseosa   , 28 (00134373– 00134374), 1♀ (00134375) (USNM). Clover Creek, 40.4086 N 112.404116 W, 1516 m, 23 Jul 1954, G.F. Edmunds, 1♀ (00119434) (UCB). Fisher’s Pass, 40.337966 N 112.567396 W, 1957 m, 06 Jul 1932, G.F. Knowlton and M.J. Janes, Chrysothamnus sp.   , 1♀ (00134613) (USNM). Uintah Co.: Fort Duchesne, 40.288066 N 109.853896 W, 15 Jun 1932, Lowell Cutler, 1♀ (00164881) (USNM). Utah Co.: American Fork, 40.41576 N 111.78366 W, 1475 m, 22 Jun [18]91, O. Heidemann, 1♀ (00242478) (CAS), 38 (00134356, 00164883–00164884), 2♀ (00134355, 00164885) (USNM); 22 Jun [18]91, P.R. Uhler Collection, 1♀ (00122112) (USNM); 24 Jun 1891, P.R. Uhler Collection, 1♀ (00122113) (USNM); 24 Jun, unknown, 18 (00070118) (USNM); 28 Jun 1965, H.H. Knight, 18 (00111892), 1♀ (00112108) (CNC). Washington Co.: 3 mi NW of Pine Valley, 37.41716 N 113.54546 W, 1890 m, 15 Jun 1967, unknown, Ericameria nauseosa   , 18 (00134610) (USNM). Saint George, 37.104176 N 113.583336 W, 853 m, 30 Jun 1965, H.H. Knight, 28 (00134371–00134372) (USNM).

Slaterocoris croceipes (Uhler)  

Figures 12 View Fig , 20A, 20E View Fig , 21–23 View Fig View Fig View Fig ; plates 1 View Plate 1 , 4F; map 3, cover

Stiphrosoma croceipes Uhler, 1893: 373   (orig. desc.).

Strongylocoris croceipes: Van Duzee, 1921: 136   (disc., in part).

Slaterocoris croceipes: Kelton, 1968: 1128   , fig. 13 (disc.); Knight, 1970: 253, fig. 14, (disc.); Henry and Wheeler, 1988: 449 (catalog); Schuh, 1995: 197 (catalog).

DIAGNOSIS: Distinguished from other apache   -group species by the variable bright yellowish to orange-brown distal portions of the femora, the yellowish orange tibiae (fig. 12); the sparse, recumbent, moderately long, golden brown vestiture (pl. 4F); and the male genitalia with the apex of the basal lobe in the right paramere usually serrate (fig. 21A, D–I) and the endosomal spicule with distal portion of the ventral lobe somewhat expanded and moderately serrate (fig. 22A, B, D, E, G). Slaterocoris croceipes   differs from S. sheridani   in antennal segment II longer than the foretibia, mostly yellowish to orange-brown femora, and untwisted dorsal lobe of the endosomal spicule. The more sparse dorsal vestiture, mostly pale antennal segment I, pale yellowish orange femur, shorter hemelytron, and the twisted dorsal lobe of the endosomal spicule (fig. 60D, E) of S. sparsus   will distinguish it from S. croceipes   ; both species can have the femora mostly pale, although the femora in S. sparsus   are usually more extensively pale yellowish orange than in S. croceipes   , whose palest specimens have a more orange or reddish-brown tint. When antennal segment I is pale in the female, S. croceipes   , S. sheridani   , and S. sparsus   can be difficult to separate. Antennal segment II is on average longer and the costal margin is somewhat curved, and the pale portions of the femora are pale orange-brown in S. croceipes   . In the latter two species antennal segment II is generally shorter, the costal margin is arcuate, and the pale portions of the femora have a more yellow tint.

REDESCRIPTION: Male: Moderately large, elongate; length 3.15–5.28, width 1.58–1.90 (pl. 1). COLORATION: Antennal segments I and II and base of femur black, variable distal portion of femur pale orange-brown, tibia yellow (fig. 12). VESTITURE: Consisting of sparse, recumbent to suberect, moderate length, golden brown setae (pl. 4F). STRUCTURE: Costal margin of hemelytron curved to subparallel; frons smooth (fig. 20A); antennal segment II longer than foretibia; mesothoracic spiracle and metathoracic scent-efferent system as in figure 20E. GENITALIA: Tergal process: Short, with one sharp point. Endosomal spicule: Distal portion of ventral lobe variable, somewhat narrow (fig. 22F) or slightly expanded (fig. 22B), usually with moderately dense marginal serration, length short to moderately long; dorsal lobe parallel sided, predominantly wide distally (fig. 22E), sometimes narrow (fig. 22F), terminating with apical serration of variable number and size; lateral edge usually without conspicuous medial and basal spine (fig. 22A–H). Right paramere: Number of apical spines variable, basal lobe usually with apex serrate (fig. 21A–I).

Female: Moderately large, obovate; length 3.45–4.20, width 1.60–2.15 (pl. 1). COLOR- ATION: Sometimes antennal segment II dusky to pale. STRUCTURE: Costal margin curved. GENITALIA: As in species group except: Ventral labiate plate: Strongly produced, base triangular, moderately wide, apex entire or cleft (fig. 23A, B). Interramal lobe: Dorsomedial margin with short process (fig. 23D, E).

HOSTS: Many specimens, from a large number of collection events, indicate that Ericameria nauseosa   ( Asteraceae   ) is a host plant. Other probable host plants are E. parryi var. vulcanica   and Chrysothamnus viscidiflorus   . A number of specimens were also collected from species of Artemisia   ( A. filifolia   and A. tridentata   ) and Chrysothamnus   . The latter generic identification could pertain to either Chrysothamnus   of Erica- meria considering the current state of botanical taxonomy. The asteraceous genus, Aster   , and three non-asteraceous plants are considered to be sitting records.

DISTRIBUTION: From the mountain ranges of southern California that border the western portion of the Mojave Desert, north along the eastern foothills of the Sierra   Nevada, isolated ranges of the Great Basin in Nevada and Utah, lower elevations of northwestern Colorado, and scattered populations in the Great Plains of Colorado, South Dakota, and Texas (map 3).

COLLECTION SUMMARY: 719 specimens from 112 collection events with the earliest record from April in California and the majority of specimens from June to early August.

DISCUSSION: The original description of croceipes ( Uhler, 1893)   stated that it was ‘‘based upon two males specimens from Los Angeles, Cal., which were sent to me by Mr. D.W. Coquillett.’’ Kelton received these two males, deposited in the USNM, and designated one the lectotype ( Kelton, 1968), stating that it bore the No. 69539. At the beginning of the current project both males were found at the CNC without labels indicating the lectotype designation or the USNM type number. To correct the ambiguous lectotype designation, I have chosen the male in the best condition, and applied two labels to it in order to stabilize the concept of croceipes: AMNH   _PBI 00122110 and LECTOTYPE 8 Stiphrosoma croceipes Uhler, 1893   , desig. by M.D. Schwartz, 2010 [red label]. I also labeled the other syntype [right hemelytron missing] to recognize it as the paralectotype (AMNH_PBI 00070120).

In the original description, ‘‘several specimens were collected at American Fork, [UT], June 22 [91]’’ and included in croceipes   , by Uhler (1893). During the course of this current project seven specimens (38, 4♀) fitting the original description have been located—six in the USNM and one female in the CAS. The CAS specimen has an additional printed label, ‘‘ Heideman [sic] Collector’   ’; I assume Heidemann collected all seven specimens. Van Duzee (1921) probably included the latter specimen in his treatment of the genus (see Other Specimens Examined). The seven Utah specimens are not considered to be syntypes and along with the lectotypes and paralectotype have been returned to the museum of provenance   .

Easternmost specimens are somewhat smaller than western specimens. The size difference is easily apprehended by comparing the length of antennal segment II in both sexes; those from Colorado, South Dakota, and Texas are shorter (0.80–0.938, 0.78– 0.81 ♀) than those from California, Nevada, and Utah (1.00–1.318, 1.06–1.23 ♀). The hemelytron length in males is conspicuously longer in California specimens (total body length 4.13–5.28) than in those from the remainder of the range (3.15–4.10).

As noted for S. apache   , variation also occurs in the number and conformation of the apical spines and apical serration on the basal lobe of the right paramere in S. croceipes   (cf. fig. 21A–I). Variable structure was observed during this study in specimens from California, Nevada, Texas, and Utah. The dorsal and ventral lobes of the endosomal spicule (fig. 22A–H) are also variable across the distribution of the species. The variation of the width of the dorsal lobe was deemed great enough to code S. croceipes   as polymorphic for this character (37-0, 1; table 2).

Specimens of S. croceipes   from Oelrich, South Dakota, and near Lefors, Texas, in the eastern reaches of the distribution, have femoral coloration as observed in the specimens from the western portion of the range in California, Nevada, and Utah. The proportions of basal black, medial orangebrown, and apical yellow coloration regions on the femora are variable across the distribution and within populations. The extent of black varies from a basal onefourth to one-half, with the reciprocal extent of orange-brown varying from a medial three-quarters to one-half. The orange-brown color also grades from a darkest tone to apical yellow in a variable fashion.

LECTOTYPE: USA: California: Los Angeles Co.: Los Angeles [34.052226 N 118.242786 W], May , Coquillett Collection , 18 ( AMNH _ PBI 00122110 View Materials ) ( USNM). GoogleMaps  

OTHER SPECIMENS EXAMINED: USA: Arizona: Coconino Co.: Grand Canyon, south Rim, 36.054446 N 112.138616 W, 03 Jun 1930, R.L. Usinger, Artemisia sp.   , 18 (00119454), 2♀ (00119455–00119456) (UCB). Gila Co.   : nr Roosevelt Lake, 33.673236 N 111.149136 W, 686 m, 27 Jun 1947, H. and M. Townes, 1♀ (00134619) (USNM). California: Inyo Co.   : 2 mi E of Big Pine, 37.1656 N 118.252326 W,

03 Jun 1975, J.D. Pinto, 1♀ (00119871) (UCR). 5 mi E of Big Pine, 37.17096 N 118.21146 W, 1341 m, 11 Jun 1969, W.F. Barr, Aster sp.   , 1♀ (00122347) (UID). 9 mi NE of Big Pine, 37.2576 N 118.173016 W, 1920 m, 09 Jun 1966, W. Gagne, 18 (00119435), 1♀ (00276763) (UCB). 10 mi NE of Big Pine, 37.26116 N 118.15086 W, 2103 m, 09 Jun 1966, C.W.O’Brien, 98 (00119436, 00276754–00276761), 1♀ (00276762) (UCB); 09 Jun 1966, W. Gagne, Chrysothamnus sp.   , 18 (00119437), Chrysothamnus sp.   , 48 (00276773–00276776), 3♀ (00276777–00276779) (UCB). 30 mi N of Saline Valley Lake, 36.677136 N 117.768166 W, 335 m, 04 May 1974, J.D. Pinto, 28 (00119865, 00120016), 1♀ (00120017) (UCR). China Lake Naval Weapons Center, Coso Mountains, Silver Peak Springs on west side, 35.736 N 117.6346 W, 1996 m, 27 Jul 1999, M. Gates, 1♀ (00119866) (UCR). Deep [Spring] Valley, Antelope Spring[s], 37.330246 N 118.090436 W, 1698 m, 02 Jun 1987, W.F. Barr, Salix sp.   ( Salicaceae   ), 1♀ (00122425) (UID). Independence, 36.802786 N 118.199176 W, 17 Jun 1937, Norman W. Frasier, 1♀ (00119445) (UCB). Olancha, Owens Valley, 36.28066 N 118.01696 W, 1122 m, 20 May 1937, J.W. Johnson   , 18 (00119439) (UCB). Panamint Mountains, 36.40036 N 117.27976 W, 1512 m, 24 May 1937, Norman W. Frasier, 1♀ (00119444) (UCB). Saline Valley Road, 19.3 mi N of Hwy 190, 36.59316 N 117.62946 W, 919 m, 04 Jun 1974, J.D. Pinto, 38 (00119864, 00120013– 00120014), 1♀ (00120015) (UCR). Kern Co.: 11 mi NW of Tehachapi on Rt 58, 35.244666 N 118.585746 W, 25 Apr 1983, G.M. Stonedahl, Ericameria nauseosa   , 78 (00108331, 00170227–00170232), 3♀ (00170233–00170235), 2 immatures (00170236–00170237) (AMNH). 20 km W of Wofford Heights on Rt 155, 35.706946 N 118.67646 W, 1500 m, 26 Jul 1999, M.D. Schwartz, Chrysothamnus viscidiflorus   , 18 (00110996), 2♀ (00112105– 00112106) (CNC). E of Caliente on Caliente Creek Rd, Twin Oaks, 35.307026 N 118.42336 W, 836 m, 22 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Ericameria nauseosa   , det. A. Sanders UCR140644 [PBI_CAL04-H126], 108 (00088461–00088470), 14♀ (00088471– 00088484) (AM), 18 (00108333), 1♀ (00322727), 3 immatures (00322728–00322730) (AMNH). Kernville, 35.754726 N 118.424446 W, 06 Jun 1952, G.L. Smith, 1♀ (00119443) (UCB); 09 Jun 1952, unknown, 1♀ (00276781) (UCB); 09 Jun 1959, G.L. Smith, 1♀ (00276782) (UCB). Mount Pinos, 34.82346 N 119.13316 W, 2438 m, 01 Jul 1965, unknown, 1♀ (00119446) (UCB). Red Rock Canyon, 35.3256 N 117.949726 W, 18 May 1937, E.P. Van Duzee, Asteraceae   , 18 (00242481) (CAS). S of Bodfish on Caliente- Bodfish Road, 35.560896 N 118.50536 W, 1094 m, 22 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Fremontodendron californicum (Torr.) Corville   ( Sterculiaceae   ), det. Field ID, 1♀ (00322745) (AMNH). S of Tehachapi on Tehachapi Willow Springs Rd, 35.055026 N 118.36966 W, 1329 m, 21 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Ericameria nauseosa   , det. A. Saunders UCR 140653 [PBI_CAL04-H92], 88 (00091708, 00088439– 00088445), 7♀ (00088446–00088452) (AM). S of Tehachapi on Tehachapi Willow Springs Rd, 35.068896 N 118.39596 W, 1393 m, 21 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Ericameria nauseosa   , det. Field ID, 98 (00322736–00322744), 5♀ (00322746–00322750) (AMNH). W of Mojave on Oak Creek Road, 35.038486 N 118.28076 W, 1055 m, 21 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Ericameria nauseosa   , det. A. Sanders UCR140653, [PBI CAL04-H92], 68 (00137262, 00322731–00322735), 10♀ (00322751–00322760) (AMNH). Los Angeles Co.: 1 mi S of Gorman, 34.781656 N 118.851676 W, 13 Jun 1964, A.G. Raske, 38 (00119410, 00276769–00276770), 2♀ (00276771–00276772) (UCB). Big Rock Creek Wash, Mohave Desert, 34.66646 N 117.87266 W, 770 m, 12 May 1944, Timberlake, Chrysothamnus sp.   , 18 (00119862), 2♀ (00120018–00120019) (UCR). Largo Vista Rd 3.1 mi S of Rt 18, SE of Llano, 34.452516 N 117.76516 W, 1275 m, 17 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Ericameria nauseosa   , det. A. Saunders UCR 140595 [PBI_CAL04-H2], 178 (00088414–00088429, 00091710), 9♀ (00088430–00088438) (AM), 228 (00107881, 00108136, 00108853, 00137236–00137238, 00322993–00323008), 17♀ (00108135, 00108970, 00322987–00322992, 00323009– 00323017) (AMNH). Los Angeles, 34.052226 N 118.242786 W, Coquillett Collection, 1♀ (00122111) (USNM); May, Coquillett Collection, 18 (00070119) (USNM), 18 paralectotype [abdomen missing] (00070120) (USNM). Rt 18 at W 263 St, 34.498336 N 117.6696 W, 1033 m, 17 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Ericameria nauseosa UCR   140595 [PBI CAL04-H2], 98 (00137263, 00322687–00322694), 23♀ (00322695– 00322717) (AMNH). SE of jct Pearblossom Hwy (Rt 18) and 263rd Street 3300N, 34.50006 N 117.668666 W, 1035 m, 19 May 2000, M.D. Schwartz, Ericameria nauseosa   , 118 (00108334, 00170213–00170222), 4♀ (00170223–00170226) (AMNH). Mono Co.: Mono Lake, 1 mi S of Lee Vining, 37.944466 N 119.102146 W, 2064 m, 20 Aug 1987, D.A. Rider, 1♀ (00242954) (DAR). Oasis, 37.487716 N 117.915656 W, 1538 m, 27 May 1954, M. Cazier, 28 (00108332, 00170238), 1♀ (00108895) (AMNH). Riverside Co.: 2.5 mi NW of Desert Springs, 33.98866 N 116.52776 W, 408 m, 25 May 1955, Timberlake, Ericameria nauseosa   , 78 (00119860 [cat UCRC ENT 32032 View Materials ], 00120004 [cat UCRC ENT 32025 View Materials ]–00120009 [cat UCRC ENT 32035 View Materials ]), 3♀ (00120010 [cat UCRC ENT 32026 View Materials ]–00120012 [cat UCRC ENT 32031 View Materials ]) (UCR). Round Valley, 33.805026 N 116.662796 W, 2754 m, 27 Jul 1947, R.H. Beamer, 2♀ (00117852–00117853) (KU). Upper Palm Canyon, 14 mi SE of Keen Camp Summit, 33.63776 N 116.64526 W, 1341 m, 08 Aug 1964, E.I. Schlinger, 18 (00119868) (UCR). Vandevanter Flat, San Jacinto Mountains, 33.783896 N 116.957786 W, 04 May 1940, unknown, Asteraceae   , 18 (00242480), 1♀ (00242479) (CAS); 04 Jun 1940, unknown, Chrysothamnus sp.   , 18 (00119438), Chrysothamnus sp.   , 3♀ (00276764–00276766) (UCB); 04 Jun 1940, H.T. Reynolds, Chrysothamnus sp.   , 1♀ (00276767) (UCB); 04 Jun 1940, C.D. Michener, 1♀ (00276768) (UCB); top of Berdoo Canyon, 33.83386 N 116.07336 W, 1128 m, 14 Jun 1963, E.I. Schlinger, 1♀ (00119872) (UCR). San Bernardino Co.: 1 mi N of X of FS Rd 2N93 and Hwy 38, 34.21846 N 116.81626 W, 2350 m, 24 Jun 1997, J. Heraty, 18 (00119870) (UCR). 5.4 mi W of Lucerne Valley, 34.443896 N 117.06166 W, 935 m, 13 May 1978, J.D. Pinto and R.T. Schuh, Ericameria nauseosa   , 668 (00108329, 00170290–00170313, 00322857–00322897), 37♀ (00169981, 00322898–00322933) (AMNH); 13 May 1978, John D. Pinto, Chrysothamnus sp.   , 158 (00119983–00119997), 7♀ (001198756, 00119998–00120003) (UCR). Apple Valley, 34.531396 N 117.282786 W, 15 May 1955, W.R.M. Mason, 18 (00112099) (CNC). Barton Flats, 34.172516 N 116.863086 W, 1997 m, 22 Jul 1981, G. Gordh, 18 (00119863) (UCR). Cajon Pass Summit on I-15, 34.298896 N 117.455566 W, 02 Jul 1949, Flock, 18 (00119869) (UCR). Camp Baldy, 34.295286 N 116.913926 W, 11 Jul 1950, R. DeNoble, 28 (00119440, 00276780) (UCB). Ironwood Campground, 34.304166 N 117.012226 W, 2130 m, 22 Aug 1999, M. Gates, 18 (00119867) (UCR). NW of Pioneertown, T1N R4E S12, 34.18446 N 116.52696 W, 1280 m, 03 May 1994, M. Cooperband, 18 (00119861) (UCR). Phelan, Rt 138 at Phelan Road, 34.425316 N 117.61746 W, 1310 m, 16 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Ericameria nauseosa   , det. A. Saunders UCR 140595 [PBI_CAL04-H2], 38 (00088453–00088455), 5♀ (00088456– 00088460) (AM), 238 (00107875, 00322718– 00322721, 00322934–00322951), 41♀ (00169985, 00322722–00322726, 00322952– 00322986) (AMNH). Tulare Co.: 21 mi W of Rt 395 toward Kennedy Meadows, 36.009446 N 118.503086 W, 1875 m, 01 Jul 1980, R.T. Schuh, Ericameria parryi var. vulcanica   , 88 (00108335, 00108843– 00108849), 3♀ (00108850–00108852) (AMNH). Colorado: Delta Co.   : Delta   , 38.742226 N 108.068336 W, 27 Jun 1938, R. Bauer, 2♀ (00134594–00134595) (USNM); 27 Jun 1938, U. Lanham, 1♀ (00134596) (USNM). Mesa Co.   : Collbran, 39.240166 N 107.959736 W, 1829 m, 08 Aug 1922, E.R. Kalmbach, 1♀ (00164886) (USNM). Glade Park, 38.994746 N 108.741176 W, 2105 m, 09 Aug 1931, F.C. Hottes, 18 (00070130), 1♀ (00134615) (USNM). Moffat Co.: Massadona, 40.25286 N 108.64986 W, 1768 m, 01 Jul 1931, R.H. Beamer, 1♀ (00117855) (KU). Maybell, 40.517756 N 108.080286 W, 1805 m, 30 Jun 1931, R.H. Beamer, 1♀ (00117854) (KU). Morgan Co.: 5 mi N of Wiggins, 40.303036 N 104.072226 W, 12 Jun 1979, D.A. Polhemus, 28 (00116819, 00116953) (JTP). Weld Co.: 5 mi NW of Roggen, 40.218616 N 104.43866 W, 1461 m, 16 Jul 1983, D.A. and J.T. Polhemus, Artemisia filifolia Torr.   , 7♀ (00116958–00116964) (JTP). 5 mi W of Roggen, 40.16756 N 104.466266 W, 1585 m, 16 Jul 1983, R.T. Schuh, D.A. and J.T. Polhemus, Artemisia filifolia Torr.   , 18 (00108396), 1♀ (00108396) (AMNH). 10 mi E of Roggen along I-25, 40.20186 N 104.25496 W, 1426 m, 11 Jul 1979, D.A. Polhemus, 18 (00116727), 1♀ (00116965) (JTP). Fort Lupton, 40.084726 N 104.81256 W, 10 Jun, unknown, 18 (00134368) (USNM); 10 Jun 1910, unknown, 1♀ (00134614) (USNM). Keenesburg, 40.108336 N 104.519446 W, 11 Jun 1961, B.H. Poole, 3♀ (00112102–00112104) (CNC). Unknown co.: Colo 2304, unknown, 1♀ (00134357) (USNM). Nevada: Carson City Co.: 4 mi S of Carson City, 39.11626 N 119.78156 W, 1463 m, 30 May 1930, W.F. Chamberlain, Artemisia sp.   , 28 (00119044–00119045), 1♀ (00119046) (TAMU). Clark Co.: Charleston Peak, 36.272226 N 115.694176 W, 2286 m, 22 Jul 1982, J.T. Polhemus, 18 (00116728), 3♀ (00116954–00116956) (JTP). Kyle Canyon, 36.262746 N 115.606686 W, 2090 m, 26 Jun 1966, W.F. Barr, 18 (00122345), 1♀ (00122346) (UID). Lee Canyon, Charleston Peak, 36.354516 N 115.64056 W, 2286 m, 20 Jul 1982, J.T. Polhemus, 1♀ (00116957) (JTP). Esmeralda Co.   : 2 mi W of Lida, 37.45136 N 117.514916 W, 1935 m, 08 Jun 1966, W. Gagne, Light Trap, 18 (00119441), 38 (00276790–00276792) (UCB). 3 mi W of Lida, 37.458336 N 117.551856 W, 2073 m, 07 Jul 1966, C.W. O’Brien, 18 (00276783), 2♀ (00276784–00276785) (UCB); 07 Jul 1966, L. and C.W. O’Brien, 18 (00119411) (UCB). 6 mi N of Goldfield, 37.795356 N 117.234726 W, 1615 m, 08 Jun 1966, W. Gagne, 18 (00119039) (TAMU), 18 (00276793) (UCB). Lincoln Co.: Beaver Dam State Park, 37.518136 N 114.076366 W, 1555 m, 18 Jun 1986, J.B. Knight and K.R. Helms, 1♀ (00108896) (AMNH). Pioche, 37.929696 N 114.452216 W, 1852 m, 09 Jul 1965, W.F. Barr, Penstemon sp.   ( Scrophulariaceae   ), 28 (00122344, 00122439), 1♀ (00122440) (UID). Lyon Co.: 7 mi N of Sweetwater Summit on Rt 338, Toiyabe National Forest, 38.613626 N 119.240546 W, 1865 m, 11 Jul 1980, R.T. Schuh and G.M. Stonedahl, Ericameria nauseosa   , 28 (00108336, 00170314), 9♀ (00170315–00170323) (AMNH). 8 mi N of Sweetwater summit on highway 22, Toiyabe National Forest, 38.630466 N 119.260976 W, 1811 m, 11 Jul 1980, G.M. Stonedahl, Ericameria nauseosa   , 4♀ (00108337, 00170324–00170326) (AMNH). Nye Co.: 2 mi E of Tonopah, 38.067226 N 117.192446 W, 1890 m, 07 Jun 1966, W. Gagne, 28 (00119442, 00276786), 3♀ (00276787– 00276789) (UCB). 5.5 mi S of Belmont on Hwy 82, 38.514016 N 116.932316 W, 1981 m, 13 Jul 1980, G.M. Stonedahl, Artemisia tridentata   , 1♀ (00108897) (AMNH). Mercury, 18 M, 36.660566 N 115.993616 W, 20 Jun 1965, H.H. Knight and J. Merino, Ericameria nauseosa   , 1♀ (00119067) (TAMU). Mercury, 401 M, 36.660566 N 115.993616 W, 18 Jun 1965, H.H. Knight and J. Merino, Ericameria nauseosa   , 18 (00112100) (CNC), 28 (00119063–00119064), 2♀ (00119065– 00119066) (TAMU). White Pine Co.   : Lehman Caves, Great Basin National Park, 39.005556 N 114.219446 W, 24 Jun 1966, W. Gagne, J. Haddock, 18 (00119429) (UCB). South Dakota: Fall River Co.: Angostura Dam, Hot Springs, 43.344636 N 103.439396 W, 967 m, 06 Jul 1968, A.T. Howden, 1♀ (00115908) (CNC). Oelrichs, 43.178896 N 103.230286 W, 1022 m, 28 Jun 1973, L.A. Kelton, Artemisia sp.   , 28 (00110993, 00112135), Artemisia sp.   , 178 (00115880– 00115896), 11♀ (00115897–00115907) (CNC). Texas: Gray Co.: 3 mi E of Lefors, 35.44146 N 100.79696 W, 878 m, 29 May 2002, J.C. Schaffner, 58 (00118222, 00119058– 00119060, 00245354), 2♀ (00119061, 00245355) (TAMU). Hemphill Co.: 7.5 mi N of Canadian River on Hwy 83, 35.983616 N 100.465276 W, 18 May 2002, E.G. Riley, 1♀ (00119062) (TAMU). Utah: Beaver Co.: Milford, 39.396916 N 113.017896 W, 1513 m, 07 Jun 1969, W.F. Chamberlain, 18 (00119047) (TAMU). Garfield Co.: Capitol Reef National Park, Grand Wash, Cobab Canyon Trail, 38.156 N 111.166676 W, 1827 m, 21 Jun 1983, R.T. Schuh, M.D. Schwartz, Ericameria nauseosa   , 28 (00108390–00108391), 2♀ (00108392– 00108393) (AMNH). Jct. Rts 95 & 276, S of Hanksville, T32S R12E, 38.025056 N 110.570986 W, 1494 m, 18 Jun 1983, R.T. Schuh and M.D. Schwartz, Ericameria nauseosa   , 18 (00108394), 1♀ (00108395) (AMNH). Iron Co.   : Bear Valley, 37.926626 N 112.607346 W, 2295 m, 09 Jun 1966, G.F. Knowlton, 18 (00134611) (USNM). Cedar City, 37.675216 N 113.050686 W, 1798 m, 22 Jun 1933, O.H. Swezey, 58 (00242463– 00242467), 10♀ (00242468–00242477) (CAS). Kanarraville, 37.538896 N 113.183336 W, 09 Jun 1960, G.F. Knowlton, 1♀ (00164887) (USNM). Juab Co.: Deep Creek Mountains, 39.843096 N 113.822886 W, 1753 m, Jun 1928, Vasco M. Tanner, 2♀ (00134616, 00164882) (USNM). Kane Co.: Whitehouse, Utah Exp. Sta., 37.078326 N 111.889896 W, 1316 m, 20 Jun 1933, G.F. Knowlton, 18 (00112036) (CNC), 18 (00134612) (USNM). Millard Co.: Skull Rock Pass, 53 mi WSW of Delta   , 39.033296 N 113.339696 W, 1596 m, 12 Jun 1992, W.F. Chamberlain, 18 (00119048) (TAMU). Sevier Co.: 11 mi W of Sevier on I- 70, 38.575016 N 112.461326 W, 1999 m, 15 Jul 1980, R.T. Schuh, G.M. Stonedahl, Chrysothamnus viscidiflorus   , 18 (00170271), 19♀ (00169986, 00170272–00170289) (AMNH); 15 Jul 1980, G.M. Stonedahl, Chrysothamnus viscidiflorus   , 78 (00170253–00170258, 00107900), 12♀ (00170259–00170270) (AMNH). Clear Creek Narrows Summit on Rt 4, 38.581566 N 112.256786 W, 2243 m, 15 Jun 1980, R.T. Schuh and G.M. Stonedahl, 58 (00107907, 00170239–00170242), 10♀ (00170243–00170252) (AMNH). Monroe, 38.636 N 112.126 W, 25 Jul 1906, unknown, 1♀ (00134618) (USNM). Salina, 38.948286 N 111.857326 W, 1577 m, 28 Jun 1954, G.L. Nielsen, 18 (00070117) (USNM). Tooele Co.: 4 mi NE of Camel Back Mountain [Camels Back Ridge], 40.185866 N 112.916546 W, 1326 m, 07 Jun 1954, P.D. Ashlock, Ericameria nauseosa   , 28 (00134373– 00134374), 1♀ (00134375) (USNM). Clover Creek, 40.4086 N 112.404116 W, 1516 m, 23 Jul 1954, G.F. Edmunds, 1♀ (00119434) (UCB). Fisher’s Pass, 40.337966 N 112.567396 W, 1957 m, 06 Jul 1932, G.F. Knowlton and M.J. Janes, Chrysothamnus sp.   , 1♀ (00134613) (USNM). Uintah Co.: Fort Duchesne, 40.288066 N 109.853896 W, 15 Jun 1932, Lowell Cutler, 1♀ (00164881) (USNM). Utah Co.: American Fork, 40.41576 N 111.78366 W, 1475 m, 22 Jun [18]91, O. Heidemann, 1♀ (00242478) (CAS), 38 (00134356, 00164883–00164884), 2♀ (00134355, 00164885) (USNM); 22 Jun [18]91, P.R. Uhler Collection, 1♀ (00122112) (USNM); 24 Jun 1891, P.R. Uhler Collection, 1♀ (00122113) (USNM); 24 Jun, unknown, 18 (00070118) (USNM); 28 Jun 1965, H.H. Knight, 18 (00111892), 1♀ (00112108) (CNC). Washington Co.: 3 mi NW of Pine Valley, 37.41716 N 113.54546 W, 1890 m, 15 Jun 1967, unknown, Ericameria nauseosa   , 18 (00134610) (USNM). Saint George, 37.104176 N 113.583336 W, 853 m, 30 Jun 1965, H.H. Knight, 28 (00134371–00134372) (USNM).

AMNH

American Museum of Natural History

USNM

Smithsonian Institution, National Museum of Natural History

CAS

California Academy of Sciences

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hemiptera

Family

Miridae

Genus

Slaterocoris

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hemiptera

Family

Miridae

Genus

Slaterocoris

Loc

Slaterocoris atratus (Uhler) Slaterocoris croceipes (Uhler)

Schwartz, Michael D. 2011
2011
Loc

Slaterocoris atratus: Kelton, 1968: 1135

Schuh, R. T. 1995: 197
Henry, T. J. & A. G. Wheeler, Jr. 1988: 448
Knight, H. H. 1970: 246
Kelton, L. A. 1968: 1135
1968
Loc

Slaterocoris croceipes: Kelton, 1968: 1128

Schuh, R. T. 1995: 197
Henry, T. J. & A. G. Wheeler, Jr. 1988: 449
Knight, H. H. 1970: 253
Kelton, L. A. 1968: 1128
1968
Loc

Strongylocoris atratus: Van Duzee, 1921: 135

Knight, H. H. 1938: 7
Van Duzee, E. P. 1921: 135
1921
Loc

Strongylocoris croceipes: Van Duzee, 1921: 136

Van Duzee, E. P. 1921: 136
1921
Loc

Stiphrosoma atrata, Uhler, 1894: 268

Uhler, P. R. 1894: 268
1894
Loc

Stiphrosoma croceipes

Uhler, P. R. 1893: 373
1893