Aname mellosa, Harvey, Frances S. B., Framenau, Volker W., Wojcieszek, Janine M., Rix, Michael G. & Harvey, Mark S., 2012

Harvey, Frances S. B., Framenau, Volker W., Wojcieszek, Janine M., Rix, Michael G. & Harvey, Mark S., 2012, Molecular and morphological characterisation of new species in the trapdoor spider genus Aname (Araneae: Mygalomorphae: Nemesiidae) from the Pilbara bioregion of Western Australia, Zootaxa 3383, pp. 15-38 : 22-30

publication ID

https://doi.org/ 10.5281/zenodo.215484

publication LSID

lsid:zoobank.org:pub:3BC5B4FB-8A70-4363-A45E-08F03E2204AA

DOI

https://doi.org/10.5281/zenodo.6168284

persistent identifier

https://treatment.plazi.org/id/067C9365-146E-45FC-81F8-23EF2AF60CB8

taxon LSID

lsid:zoobank.org:act:067C9365-146E-45FC-81F8-23EF2AF60CB8

treatment provided by

Plazi

scientific name

Aname mellosa
status

n. sp.

Aname mellosa n. sp.

Figs 1 –5 View FIGURES 1 – 4 View FIGURES 5 – 8 , 9– 24 View FIGURE 9 View FIGURES 10 – 20 View FIGURES 21 – 24

Type material. AUSTRALIA: Western Australia: holotype male, Jinayri, ca. 60 km NW. of Newman, 22 ° 58 ’05.7”S, 119 ° 17 ’06.1”E, 3 February 2009, pitfall trap, J. Gollan, N. Sullivan, M. Semeniuk, M. Beatson ( WAM T 94870 View Materials ). Paratypes: 1 male, same data as holotype except 22 ° 57 ’ 46.1 ”S, 119 ° 13 ’ 36.3 ”E, 6 February 2009 ( WAM T 96116 View Materials ); 2 males, same data except 22 ° 57 ’ 46.5 ”S, 119 ° 13 ’ 36.5 ”E, 7 February 2009 ( WAM T 96121 View Materials , T 94864 View Materials ); 1 male, same data except 22 ° 57 ’ 47.4 ”S, 119 ° 13 ’ 37.1 ”E, 6 February 2009 ( WAM T 96122 View Materials ); 1 male, same data except 22 ° 57 ’ 47.4 ”S, 119 ° 13 ’ 37.1 ”E, 3 February 2009 (QM S 91581 View Materials ); 1 male, same data except 22 ° 57 ’ 48.7 ”S, 119 ° 13 ’ 38.1 ”E, 5 February 2009 ( WAM T 94881 View Materials ); 1 male, same data except 22 ° 58 ’03.2”S, 119 ° 17 ’06.2”E, 5 February 2009 ( WAM T 96113 View Materials ); 1 male, same data except 22 ° 58 ’04.0”S, 119 ° 17 ’06.4”E, 5 February 2009 ( WAM T 94871 View Materials DNA); 1 male, same data except 22 ° 58 ’05.0”S, 119 ° 17 ’06.2”E, 5 February 2009 ( WAM T 96117 View Materials ); 2 males, same data except 22 ° 58 ’09.5”S, 119 ° 15 ’ 35.1 ”E, 3 February 2009 ( WAM T 94885 View Materials , T 94872 View Materials ); 1 male, same data except 22 ° 58 ’09.5”S, 119 ° 15 ’ 36.2 ”E, 7 February 2009 ( WAM T 96128 View Materials ); 2 males, same data except 22 ° 58 ’09.5”S, 119 ° 15 ’ 36.2 ”E, 1 February 2009 ( WAM T 94884 View Materials DNA, T 94882 View Materials ); 1 male, same data except 22 ° 58 ’09.5”S, 119 ° 15 ’ 36.4 ”E, 5 February 2009 ( WAM T 94857 View Materials ); 1 male, same data except 22 ° 58 ’09.5”S, 119 ° 15 ’ 36.4 ”E, 6 February 2009 ( WAM T 94862 View Materials ); 2 males, same data except 22 ° 58 ’09.8”S, 119 ° 15 ’ 36.5 ”E, 7 February 2009 ( WAM T 96144 View Materials , T 96126 View Materials ); 1 male, same data except 22 ° 58 ’ 10.2 ”S, 119 ° 15 ’ 37.1 ”E, 6 February 2009 ( WAM T 94863 View Materials ); 1 male, same data except 22 ° 58 ’ 26.4 ”S, 119 ° 18 ’00.8”E, 6 February 2009 ( WAM T 94874 View Materials ); 1 female, same data except 22 ° 58 ’ 26.9 ”S, 119 ° 18 ’03.1”E, 1 February 2009 ( WAM T 96145 View Materials ); 2 males, same data except 22 ° 58 ’ 26.9 ”S, 119 ° 18 ’03.1”E, 6 February 2009 ( WAM T 94856 View Materials , T 94860 View Materials ); 1 male, same data except 22 ° 58 ’27.0”S, 119 ° 18 ’04.0”E, 7 February 2009 ( WAM T 94886 View Materials ); 1 male, same data except 22 ° 58 ’ 42.6 ”S, 119 ° 15 ’ 39.8 ”E, 5 February 2009 ( WAM T 96129 View Materials ); 1 male, same data except 22 ° 58 ’ 42.9 ”S, 119 ° 15 ’ 41.2 ”E, 7 February 2009 ( WAM T 94859 View Materials ); 1 male, same data except 22 ° 58 ’ 43.5 ”S, 119 ° 15 ’ 40.5 ”E, 7 February 2009 ( WAM T 96123 View Materials ); 1 male, same data except 22 ° 59 ’03.3”S, 119 ° 16 ’03.3”E, 5 February 2009 ( WAM T 96114 View Materials ); 1 male, same data except 22 ° 59 ’03.7”S, 119 ° 18 ’03.8”E, 2 February 2009 ( WAM T 94879 View Materials DNA); 1 male, same data except 22 ° 59 ’04.5”S, 119 ° 16 ’02.3”E, 7 February 2009 ( WAM T 94866 View Materials ); 1 male, same data except 22 ° 59 ’04.5”S, 119 ° 16 ’02.3”E, 6 February 2009 ( WAM T 94868 View Materials ); 2 males, same data except 22 ° 59 ’08.6”S, 119 ° 15 ’02.8”E, 7 February 2009 ( WAM T 94854 View Materials , T 94861 View Materials ); 1 male, same data except 22 ° 59 ’08.8”S, 119 ° 15 ’03.1”E, 7 February 2009 ( WAM T 94869 View Materials ); 2 males, same data except 22 ° 59 ’09.2”S, 119 ° 15 ’04.4”E, 7 February 2009 ( WAM T 94853 View Materials , T 96118 View Materials ); 1 male, same data except 22 ° 59 ’09.5”S, 119 ° 15 ’ 35.1 ”E, 7 February 2009 ( WAM T 94867 View Materials ); 1 male, same data except 22 ° 59 ’ 14.1 ”S, 119 ° 16 ’09.6”E, 6 February 2009 ( WAM T 94858 View Materials ); 1 male, same data except 22 ° 59 ’ 14.9 ”S, 119 ° 16 ’ 10.3 ”E, 7 February 2009 ( WAM T 96119 View Materials ); 1 male, same data except 22 ° 59 ’ 15.2 ”S, 119 ° 16 ’ 11.3 ”E, 6 February 2009 ( WAM T 96125 View Materials ); 1 male, same data except 22 ° 59 ’ 19.9 ”S, 119 ° 14 ’ 11.3 ”E, 8 February 2009 ( WAM T 96127 View Materials ); 1 male, same data except 22 ° 59 ’ 21.3 ”S, 119 ° 14 ’ 10.7 ”E, 6 February 2009 ( WAM T 94865 View Materials ); 1 male, same data except 22 ° 59 ’ 23.1 ”S, 119 ° 14 ’ 11.1 ”E, 8 February 2009 ( WAM T 94851 View Materials ); 1 male, same data except 22 ° 59 ’ 23.8 ”S, 119 ° 14 ’ 11.5 ”E, 7 February 2009 ( WAM T 96124 View Materials ); 1 male, same data except 22 ° 59 ’ 23.8 ”S, 119 ° 14 ’ 11.5 ”E, 6 February 2009 ( WAM T 96120 View Materials ); 1 male, same data except 22 ° 59 ’ 24.5 ”S, 119 ° 14 ’12.0”E, 5 February 2009 ( WAM T 94883 View Materials ); 1 male, same data except 23 °00’ 23.3 ”S, 119 ° 14 ’ 42.6 ”E, 7 February 2009 ( WAM T 94876 View Materials ); 1 male, same data except 23 °00’ 23.9 ”S, 119 ° 14 ’ 43.6 ”E, 5 February 2009 ( WAM T 96151 View Materials ); 1 male, same data except 23 °00’ 39.8 ”S, 119 ° 14 ’09.0”E, 7 February 2009 ( WAM T 96115 View Materials ); 1 male, same data except 23 °00’ 49.2 ”S, 119 ° 14 ’ 13.9 ”E, 6 February 2009 ( WAM T 94852 View Materials ); 1 male, same data except 23 °01’29.0”S, 119 ° 14 ’ 37.8 ”E, 3 February 2009 ( WAM T 96140 View Materials DNA); 1 male, same data except 23 °01’ 29.4 ”S, 119 ° 14 ’37.0”E, 4 February 2009 ( WAM T 94875 View Materials ); 1 male, same data except 23 °02’26.0”S, 119 ° 16 ’ 45.2 ”E, 2 February 2009 ( WAM T 94880 View Materials DNA); 2 males, same data except 23 °03’08.8”S, 119 ° 16 ’ 14.9 ”E, 3 February 2009 ( WAM T 94873 View Materials , T 94878 View Materials ); 1 male, same data except 23 °03’09.8”S, 119 ° 16 ’ 13.9 ”E, 5 February 2009 ( WAM T 94850 View Materials ); 1 female, same data as holotype except 22 ° 59 ’ 24.5 ”S, 119 ° 14 ’12.0”E, 8 February 2009 ( WAM T 96150 View Materials ).

Other material examined. AUSTRALIA: Western Australia: 1 male, Abydos mine lease, ca. 64 km W. of Marble Bar, 21 °07’ 39.7 ”S, 119 °08’03.2”E, 15 May 2010, dry pitfall trap, P. Bolton ( WAM T 102291); 1 juvenile, same data except 21 °07’ 50.2 ”S, 119 °07’ 32.7 ”E, 14 May 2010 ( WAM T 102290); 1 male, same data except 21 °07’ 53 ”S, 119 °08’07.2”E, 4–10 May 2010 ( WAM T 102287); 1 male, same data except 5–10 May 2010 ( WAM T 102288); 1 male, same data except 21 °07’ 54.9 ”S, 119 °07’ 54 ”E, 28 – 5 May 2010, wet pitfall trap ( WAM T 102284); 1 male, API Hardey, 96.9 km WNW. of Paraburdoo, 22 ° 49 ’04”S, 116 ° 48 ’ 53 ”E, 24 March 2010, dug from burrow, P. Runham ( WAM T 110361); 1 male, Aquila, 12.75 km SE. of Karratha, 20 ° 48 ’ 34 ”S, 116 ° 56 ’ 29 ”E, 8 April 2009, dug from burrow, P. Runham ( WAM T 100079); 1 male, same data except 5 April 2009 ( WAM T 100080); 1 male, Aquila, 20.6 km E. of Karratha, 20 ° 45 ’ 45 ”S, 117 °02’ 31 ”E, 5 April 2009, dug from burrow, P. Runham ( WAM T 100077); 1 male, same data except 7 April 2009 ( WAM T 100078); 1 male, Area C, 101 km NW. of Newman, 23 °00’ 20 ”S, 118 ° 49 ’ 38 ”E, 25 February 2010, dug from burrow, open mulga woodland on clayey loam plain, M. Greenham ( WAM T 101158); 1 female, same data except D. Kamien, T. Sachse ( WAM T 101162); 1 male, Area C, 61.8 km NW. of Newman, 22 ° 59 ’ 55 ”S, 118 ° 52 ’07”E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T 105930); 1 male, same data ( WAM T 105931); 1 male, Area C, 74.1 km NW. of Newman, 22 ° 55 ’04”S, 119 ° 11 ’ 21 ”E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T 105957); 1 female, same data except 22 ° 55 ’05”S, 119 ° 11 ’ 21 ”E, 18 February 2010, dug from burrow, drainage in valley/flats, D. Kamien ( WAM T 101163); 1 female, Area C, 75.7 km NW. of Newman, 22 ° 54 ’ 21 ”S, 119 ° 10 ’ 48 ”E, 19 February 2010, dug from burrow, loamy plain, D. Kamien, T. Sachse ( WAM T 101165); 1 male, Area C, 75.8 km NW. of Newman, 22 ° 54 ’ 21 ”S, 119 ° 10 ’ 48 ”E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T 105943); 1 male, Area C, 78.9 km NW. of Newman, 22 ° 54 ’ 10 ”S, 119 °08’ 38 ”E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T 105958); 1 male, Area C, 81.4 km NW. of Newman, 22 ° 57 ’ 29 ”S, 119 °04’09”E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T 105914); 1 male, Area C, 81.8 km NW. of Newman, 23 °00’ 20 ”S, 119 °01’ 58 ”E, 21 February 2010, dug from burrow, plateau/top of rocky hill, M. Greenham ( WAM T 101156); 1 female, Area C, 82.6 km NW. of Newman, 22 ° 58 ’ 54 ”S, 119 °00’ 49 ”E, 20 February 2010, dug from burrow, Gully, Z. Hamilton ( WAM T 101161); 1 male, same data except 22 ° 58 ’ 55 ”S, 119 °00’ 50 ”E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T 105921); 1 male, Area C, 82.7 km NW. of Newman, 22 ° 53 ’ 57 ”S, 119 °05’ 58 ”E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T 105954); 1 male, Area C, 82.7 km NW. of Newman, 22 ° 57 ’ 49 ”S, 119 °02’ 50 ”E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T 105910); 1 female, Area C, 82.7 km NW. of Newman, 22 ° 57 ’ 53 ”S, 119 °02’ 53 ”E, 20 February 2010, dug from burrow, mulga plain, M. Greenham ( WAM T 101164); 1 male, Area C, 82.7 km NW. of Newman, 23 °00’ 18 ”S, 119 °01’ 26 ”E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T 105960); 1 male, Area C, 83.1 km NW. of Newman, 22 ° 58 ’ 30 ”S, 119 °02’ 16 ”E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T 105956); 1 male, same data ( WAM T 105967); 1 male, Area C, 83.9 km NW. of Newman, 22 ° 59 ’ 35 ”S, 119 °00’ 57 ”E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T 105932); 1 male, Area C, 84.7 km NW. of Newman, 22 ° 54 ’ 21 ”S, 119 °01’ 44 ”E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T 105950); 1 male, same data ( WAM T 105974); 1 male, Area C, 85.2 km NW. of Newman, 23 °00’ 14 ”S, 118 ° 59 ’ 41 ”E, 22 February 2010, dug from burrow, Z. Hamilton ( WAM T 101154); 1 male, Area C, 86.2 km NW. of Newman, 23 °00’05”S, 118 ° 58 ’ 54 ”E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T 105928); 1 male, Area C, 87.2 km NW. of Newman, 22 ° 59 ’ 22 ”S, 118 ° 58 ’ 58 ”E, 22 February 2010, dug from burrow, Loamy clay plain, D. Kamien ( WAM T 101155); 1 male, Area C, 89 km NW. of Newman, 22 ° 57 ’ 23 ”S, 118 ° 59 ’00”E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T 105911); 1 male, same data ( WAM T 105912); 1 male, same data ( WAM T 105975); 1 male, Area C, 89.2 km NW. of Newman, 22 ° 57 ’ 56 ”S, 118 ° 58 ’ 32 ”E, 23 February 2010, dug from burrow, rocky loam plain, D. Kamien, T.Sachse ( WAM T 101157); 1 male, Area C, 89.4 km NW. of Newman, 22 ° 58 ’ 49 ”S, 118 ° 57 ’ 44 ”E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T 105925); 1 male, same data ( WAM T 105944); 1 male, same data ( WAM T 105959); 1 male, Area C, 89.6 km NW. of Newman, 22 ° 54 ’ 31 ”S, 119 °00’ 26 ”E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T 105908); 1 male, same data ( WAM T 105976); 1 female, Area C, 89.6 km NW. of Newman, 23 °00’03”S, 118 ° 56 ’ 49 ”E, 24 February 2001, dug from burrow, Rocky gully, Z. Hamilton ( WAM T 101160); 1 male, Area C, 90.6 km NW. of Newman, 22 ° 53 ’ 56 ”S, 119 °00’07”E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T 105934); 1 male, Area C, 90.6 km NW. of Newman, 22 ° 59 ’ 52 ”S, 119 °01’ 47 ”E, 1 May 2010, wet pit trap, S. Ford, G. O’Connel ( WAM T 105915); 1 male, Area C, 92.7 km NW. of Newman, 23 °01’04”S, 118 ° 54 ’ 24 ”E, 24 February 2010, dug from burrow, open mulga woodland with scattered eucalypts, M. Greenham ( WAM T 101153); 1 female, Area C, 98.2 km NW. of Newman, 23 °00’ 28 ”S, 118 ° 51 ’ 12 ”E, 21 February 2010, dug from burrow, open mulga woodland on clayey loam plain, M. Greenham ( WAM T 101166); 1 male, 36 km NW. of Balfour Downs Homestead, Pilbara Biological Survey site BDRN 11, 22° 30 ’ 59 ”S, 120 ° 39 ’ 34 ”E, 11 September 2005 – 14 August 2006, wet pitfall, CALM Pilbara Survey ( WAM T 109337); 1 male, Bungaroo, 17.1 km S. of Pannawonica, 21 ° 47 ’04”S, 116 ° 15 ’ 31 ”E, 14 March 2010, dug from burrow, J. Alexander ( WAM T 102531); 1 male, Bungaroo, same data except dry pit trap, J. Alexander ( WAM T 102533); 1 juvenile, same data except dug from burrow, D. Keirle ( WAM T 102536); 1 male, Bungaroo, 19.5 km S. of Pannawonica, 21 ° 51 ’00”S, 116 ° 22 ’ 12 ”E, 15 March 2010, dry pit trap, J. Alexander ( WAM T 102530); 1 male, same data except 13 March 2010, D. Keirle ( WAM T 102532); 1 male, same data ( WAM T 102535); 1 male, Bungaroo, 22.1 km S. of Pannawonica, 21 ° 50 ’ 30 ”S, 116 ° 21 ’ 31 ”E, 13 March 2010, Dry pit trap, D. Keirle ( WAM T 102534); 1 male, Bungaroo, 26.9 km S. of Pannawonica, site BUN8, 21° 49 ’08”S, 116 ° 17 ’ 59 ”E, 14–20 March 2005, dry pitfall traps, Z. Hamilton ( WAM T 72405 View Materials ); 1 male, Charles Darwin Reserve, 2.5 km N. of White Wells Homestead, 29 ° 33 ’ 29 ”S, 116 ° 57 ’ 52 ”E, 4–5 March 2008, gas lantern/pitfall, mixed shrubland with emergent eucalypts and native pine, T.F. Houston ( WAM T 88785 View Materials ); 2 males, 9 km NE. of Cowra Line Camp, Pilbara Biological Survey site RHNW 11, 22° 17 ’ 38.8 ”S, 119 °03’ 40.9 ”E, 27 August 2003 – 20 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109348); 1 male, 16 km N. of Cowra Line Camp, Pilbara Biological Survey site RHNW 07, 22 ° 13 ’ 18.2 ”S, 119 °01’ 29.5 ”E, 14 August 2003 – 20 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109353); 1 male, Davidson Creek, ~ 75 km E. of Newman, vert site 1, 23° 31 ’ 29.81 ”S, 120 ° 38 ’ 28.26 ”E, 8 April 2010, dry pitfall, J. Clark ( WAM T 102154); 1 male, Davidson Creek, ~ 75 km E. of Newman, vert site 3, 23° 28 ’ 27.25 ”S, 120 ° 34 ’ 15.86 ”E, 8 April 2010, Dry pitfall, J. Clark ( WAM T 102156); 1 male, Davidson Creek, ~ 75 km E. of Newman, vert site 5, 23° 25 ’ 15.91 ”S, 120 ° 28 ’ 59.56 ”E, 9 April 2010, dry pitfall, J. Clark ( WAM T 102155); 1 male, Fortescue Marsh, 22 ° 17 ’ 32 ”S, 119 °04’ 10 ”E, 2–5 June 2010, dry pitfall, leaf litter, P. Roberts, L. Quinn ( WAM T 107179); 1 male, same data ( WAM T 107180); 1 male, Fortescue Marsh, 22 ° 18 ’ 26 ”S, 119 ° 12 ’ 57 ”E, 2–5 June 2010, dry pitfall, leaf litter, P. Roberts, L. Quinn ( WAM T 107182); 1 male, 67.5 km S. of Fortescue River crossing on Dampier Paraburdoo Railway, Pilbara Biological Survey site PE12, 22°03’ 33.2 ”S, 117 ° 41 ’ 36.5 ”E, 5 September 2005 – 11 August 2006, wet pitfall, CALM Pilbara Survey ( WAM T 106274); 1 male, 7.4 km SW. of Fortescue Roadhouse, 21 ° 20 ’ 58 ”S, 116 °05’ 46 ”E, 24 March 2004, dug from burrow, M. Greenham, P. Runham ( WAM T 88702 View Materials ); 1 male, 13.1 km SW. of Fortescue Roadhouse, 21 ° 23 ’ 15 ”S, 116 °03’05”E, 19 March 2004, dug from burrow, M. Greenham, P. Runham ( WAM T 88698 View Materials ); 1 male, same data except 20 March 2004 ( WAM T 88700 View Materials ); 1 male, same data except 22 March 2004 ( WAM T 88705 View Materials ); 1 male, same data ( WAM T 88708 View Materials ); 1 male, same data ( WAM T 88713 View Materials ); 1 male, 17.9 km SW. of Fortescue Roadhouse, 21 ° 25 ’00”S, 116 °02’ 24 ”E, 19 March 2004, dug from burrow, M. Greenham, P. Runham ( WAM T 88703 View Materials ); 1 male, same data ( WAM T 88706 View Materials ); 1 male, same data except 24 March 2004 ( WAM T 88707 View Materials ); 1 male, 18.4 km NE. of Fortescue Roadhouse, 21 °09’ 14 ”S, 116 ° 13 ’ 59 ”E, 21 March 2004, dug from burrow, M. Greenham, P. Runham ( WAM T 88701 View Materials ); 1 male, same data ( WAM T 88710 View Materials ); 1 male, same data ( WAM T 88712 View Materials ); 1 male, same data except 22 March 2004 ( WAM T 88714 View Materials ); 1 male, 19.3 km SW. of Fortescue Roadhouse, 21 ° 26 ’07”S, 116 °01’ 55 ”E, 24 March 2004, dug from burrow, M. Greenham, P. Runham ( WAM T 88699 View Materials ); 1 male, 42 km N. of Fortescue Roadhouse, 20 ° 54 ’ 53 ”S, 116 ° 14 ’03”E, 20 March 2004, dug from burrow, M. Greenham, P. Runham ( WAM T 88697 View Materials ); 1 male, same data ( WAM T 88709 View Materials ); 1 male, same data ( WAM T 88711 View Materials ); 2 males, Jimblebar minesite, 35 km E. of Newman, 23 ° 22 ’09”S, 120 ° 11 ’ 44 ”E, 9 February 2009, pitfall trap, P. Bolton, C. Weston ( WAM T 95399 View Materials DNA); 1 male, same data except 23 ° 22 ’ 29 ”S, 120 ° 12 ’ 37 ”E, 12 February 2009, active search ( WAM T 95402 View Materials DNA); 1 male, same data except 23 ° 22 ’ 45 ”S, 120 °06’ 59 ”E, 9 February 2009, pitfall trap ( WAM T 95400 View Materials DNA); 1 male, same data except 23 ° 23 ’ 27 ”S, 120 °05’ 43 ”E, 6 February 2009, active search ( WAM T 95401 View Materials DNA); 1 male, Jinayri, ca. 65 km NW. of Newman, 22 ° 51 ’ 10 ”S, 119 ° 16 ’ 34 ”E, 31 March 2009, pitfall trap, J. Gollan, et al. ( WAM T 97013 View Materials DNA); 1 male, same data except 22 ° 54 ’ 37 ”S, 119 ° 12 ’ 30 ”E, 1 April 2009 ( WAM T 97033 View Materials DNA); 7 males, 3.5 km S. of Karratha, Pilbara Biological Survey, site DRC 01, 20 ° 46 ’ 4.7 ”S, 116 ° 50 ’ 31.3 ”E, 1 July 2003 – 2 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109322); 1 male, 6 km S. of Karratha, Pilbara Biological Survey site DRC 03, 20 ° 47 ’ 40.1 ”S, 116 ° 51 ’ 24.1 ”E, 3 July 2003 – 3 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109354); 1 male, 5 km N. of Lake Poongkaliyarra, Pilbara Biological Survey site DRC 07, 20 ° 56 ’04.4”S, 117 °06’ 54.3 ”E, 4 July 2003 – 3 October 2004, wet pitfall, DEC Pilbara survey ( WAM T 109323); 1 male, 9 km NW. of Lake Poongkaliyarra, Pilbara Biological Survey site DRC 05, 20 ° 56 ’ 23.8 ”S, 117 °02’05.3”E, 3 July 2003, 3 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109330); 1 male, ca. 21 km S. of Laverton, site 1, 28° 47 ’ 34 ”S, 122 ° 25 ’ 53 ”E, 8 January 2011, pit trap, S.A. Thompson ( WAM T 110258); 1 male, same data ( WAM T 110263); 1 male, same data ( WAM T 110269); 1 male, same data ( WAM T 110272); 1 male, ca. 21 km S. of Laverton, site 1, 31° 20 ’ 52 ”S, 119 ° 23 ’ 26 ”E, 6 January 2011, pit trap, S.A. Thompson ( WAM T 110264); 1 female, ca. 21 km S. of Laverton, site 11, 28° 49 ’ 35 ”S, 122 ° 23 ’ 20 ”E, 8 January 2011, pit trap, S.A. Thompson ( WAM T 110273); 1 male, ca. 21 km S. of Laverton, site 13, 28° 49 ’ 11 ”S, 122 ° 23 ’ 14 ”E, 6 January 2011, pit trap, S.A. Thompson ( WAM T 110276); 1 male, ca. 21 km S. of Laverton, site 2, 28° 47 ’ 55 ”S, 122 ° 26 ’01”E, 6 January 2011, pit trap, S.A. Thompson ( WAM T 110265); 1 male, same data except 8 January 2011 ( WAM T 110270); 1 male, ca. 21 km S. of Laverton, site 5, 28° 49 ’07”S, 122 ° 26 ’02”E, 6 January 2011, pit trap, S.A. Thompson ( WAM T 110271); 1 male, ca. 21 km S. of Laverton, site 7, 28° 49 ’ 25 ”S, 122 ° 26 ’ 31 ”E, 8 January 2011, pit trap, S.A. Thompson ( WAM T 110262); 1 male, same data except 6 January 2011 ( WAM T 110267); 1 male, ca. 21 km S. of Laverton, site 9, 28° 49 ’ 55 ”S, 122 ° 25 ’ 34 ”E, 8 January 2011, pit trap, S.A. Thompson ( WAM T 110257); 1 male, Lorna Glen Biological Survey, 26 °06’ 46 ”S, 121 ° 30 ’ 14 ”E, 17 March 2004, dry pit, Cowan, M. ( WAM T 88663 View Materials ); 1 male, 3.8 km E. of Lorna Glen Station, 26 ° 13 ’ 35 ”S, 121 ° 31 ’08”E, 16 March 2004, dry pit, M. Cowan ( WAM T 88653 View Materials ); 1 male, same data except 17 March 2004 ( WAM T 88658 View Materials ); 1 male, 11.9 km SW. of Lorna Glen Station, 26 ° 18 ’ 11 ”S, 121 ° 28 ’ 31 ”E, 17 March 2004, dry pit, M. Cowan ( WAM T 88659 View Materials ); 1 male, 25.2 km N. of Lorna Glen Station, 26 °00’05”S, 121 ° 33 ’ 48 ”E, 16 March 2004, dry pit, M. Cowan ( WAM T 88656 View Materials ); 1 male, Mallina Station, SW. of homestead, Pilbara Biological Survey site DRE12, 20° 54 ’ 36 ”S, 117 ° 58 ’ 58 ”E, 14 November 2003 – 12 May 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109343); 5 males, 10 km S. of Mallina Homestead, Pilbara Biological Survey site DRE13, 20° 58 ’ 10.4 ”S, 118 °02’54.0”E, 10 July 2003 – 3 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109327); 1 male, 1.2 km SSE. of Millstream, Pilbara Biological Survey site PW08, 21 ° 36 ’ 15 ”S, 117 °04’ 39 ”E, 14 July 2003 – 12 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109336); 1 juvenile, Mesa A, 43.9 km W. of Pannawonica, site MAT21, 21° 41 ’ 18 ”S, 115 ° 54 ’06”E, 5–11 August 2005, dry pitfall traps, R. Teale ( WAM T 72402 View Materials ); 1 male, Mesa J, 16.6 km SW. of Pannawonica, site MEJ1, 21° 45 ’ 34 ”S, 116 ° 13 ’ 39 ”E, 14–20 March 2005, dry pitfall traps, D. Kamien ( WAM T 72400 View Materials ); 2 males, same data ( WAM T 72401 View Materials ); 3 males, Mesa J, 17 km SW. of Pannawonica, site MEJ2, 21° 45 ’ 37 ”S, 116 ° 13 ’ 16 ”E, 14–20 March 2005, dry pitfall traps, D. Kamien ( WAM T 72399 View Materials ); 3 males, 2.5 km N. of Mile Camp, Pilbara Biological Survey site RHNE 02, 22 ° 45 ’ 57.8 ”S, 119 ° 37 ’ 53.1 ”E, 10 August 2003 – 19 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109321); 1 male, Millstream-Chichester National Park, Pilbara Survey site PW10, 21° 32 ’ 48 ”S, 117 °03’ 33 ”E, 23 November 2003 – 8 May 2004, wet pitfall trap, CALM Pilbara Survey ( WAM T 109320); 1 male, 24 km ENE. of Moorimoordinina Native Well, Pilbara Biological Survey site RHNE 08, 22 ° 27 ’07”S, 119 ° 58 ’ 34 ”E, 9 August 2003 – 20 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109335); 4 males, 10.5 km NW. of Mt Berry, Pilbara Biological Survey site WYE12, 22° 24 ’ 38.6 ”S, 116 ° 23 ’ 30.8 ”E, 10 September 2003 – 9 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109341); 2 males, 19.7 km WNW. of Mt Berry, Pilbara Biological Survey site WYE09, 22 ° 26 ’ 15.3 ”S, 116 ° 16 ’ 27.3 ”E, 8 September 2003 – 10 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109344); 2 males, 8 km NNE. of Mt Edith, Pilbara Biological Survey, site WYE05, 22 ° 34 ’ 16.9 ”S, 116 °08’ 53.8 ”E, 11 September 2003 – 7 October 2004, Ethylene Glycol Pit, CALM Pilbara Survey ( WAM T 109345); 1 male, 14.5 km SE. of Mt Florance Homestead, Pilbara Biological Survey site PE06, 21 ° 51 ’ 22.2 ”S, 117 ° 58 ’ 47.3 ”E, 3 September 2003 – 11 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109362); 6 males, 24 km WSW. of Mt Marsh, Pilbara Biological Survey site RHNW 02, 22 ° 32 ’09”S, 118 ° 59 ’ 51 ”E, 12 August 2003 – 19 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109329); 1 male, 26 km WSW. of Mt Marsh, Pilbara Biological Survey site RHNW 01, 22 ° 32 ’06”S, 118 ° 58 ’ 39 ”E, 12 August 2003 – 19 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109350); 1 male, 7 km SSE. of Mt Minnie, Pilbara Biological Survey site WYW03, 22 ° 10 ’ 10.8 ”S, 115 ° 33 ’ 39 ”E, 27 September 2003 – 30 September 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109325); 1 male, Mount Webber, 20 ° 31 ’ 21 ”S, 119 ° 18 ’ 41 ”E, 31 March 2010 – 7 May 2010, wet pitfall trap, gorge, P. Bolton ( WAM T 103131); 1 male, Mount Webber, 21 ° 31 ’ 57 ”S, 119 ° 17 ’ 44 ”E, 30 March 2010 – 7 May 2010, wet pitfall trap, east facing ridge, P. Bolton ( WAM T 103128); 1 male, Mount Webber, 21 ° 32 ’00”S, 119 ° 17 ’ 33 ”E, 29 March 2010 – 7 May 2010, wet pitfall trap, gorge, P. Bolton ( WAM T 103129); 2 males, Mt Stuart Station, NE. of homestead, Pilbara Survey site WYE13, 22° 25 ’ 30 ”S, 116 ° 25 ’ 57 ”E, 1 May 2004 – 10 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109349); 1 male, North Star, Pilbara, 21 ° 16 ’ 23.81 ”S, 119 ° 3 ’ 24.48 ”E, 11–18 February 2011, leaf litter, Thompson ( WAM T 112591); 1 male, 12.5 km N. of Nullagine, Pilbara Biological Survey site NW03, 21 ° 46 ’ 13.1 ”S, 120 °05’ 30.7 ”E, 5 August 2003 – 18 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109331); 1 male, 30 km NE. of Nullagine, 21 ° 48 ’ 16.08 ”S, 120 ° 23 ’18.00”E, 29 April 2010, C. Jackson ( WAM T 102065); 2 males, 56.3 km N. of Nullagine, Pilbara Biological Survey site NW04, 21 ° 40 ’ 42.7 ”S, 120 °05’ 18.2 ”E, 19 May 2004 – 18 May 2005, ethylene glycol pits, CALM Pilbara Survey ( WAM T 104454); 1 male, Orebody 35, ca. 8 km W. of Newman, Site 10 –P6, 23° 23 ’ 59.65 ”S, 119 ° 39 ’07.47”E, 12 March 2010 – 18 May 2010, pitfall trap, open floodplain, J. Gollan ( WAM T 104794); 1 male, Orebody 35, ca. 8 km W. of Newman, Site 3 –HC1, 23° 24 ’17.00”S, 119 ° 36 ’ 18.90 ”E, 15 March 2010, hand collected, drainage gully, south facing ridge, deeply vegetated, J. Gollan, M. Bulbert, M. Semeniuk, J. Bevan ( WAM T 104784); 1 male, Orebody 35, ca. 8 km W. of Newman, Site 6 –P8, 23° 24 ’07.00”S, 119 ° 34 ’ 27.88 ”E, 12 March– 18 May 2010, pitfall trap, slope with spinifex, J. Gollan ( WAM T 104810); 1 male, Orebody 35, ca. 8 km W. of Newman, Site 9 –P10, 23° 23 ’ 22.32 ”S, 119 ° 38 ’ 36.06 ”E, 12 March– 18 May 2010, pitfall trap, open floodplain, J. Gollan ( WAM T 104807); 1 male, 2.3 km ESE. of Python Pool, Pilbara Biological Survey site PW01, 21 ° 20 ’ 22.8 ”S, 117 ° 15 ’ 36.3 ”E, 16 July 2003 – 11 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109356); 2 males, 5 km WSW. of Python Pool, Pilbara Biological Survey site PW03, 21 ° 20 ’ 28.7 ”S, 117 ° 11 ’ 18.9 ”E, 8 May 2004 – 12 May 2005, wet pitfall, CALM Pilbara Survey ( WAM T 109339); 5 males, 1 juvenile, 11.5 km SW. of Rhodes Ridge, Pilbara Biological Survey site RHNC 07, 23 °08’ 45.7 ”S, 119 ° 15 ’ 56.9 ”E, 25 May 2004 – 11 May 2005, wet pitfall, CALM Pilbara Survey ( WAM T 109324); 1 male, Roy Hill, 22 ° 25 ’ 35.77 ”S, 119 ° 56 ’ 32.69 ”E, 11–17 March 2010, wet pitfall trap, N. Thompson, L. Roque-Albelo ( WAM T 102618); 1 female, South Parmelia, 66 km NW. of Newman, 23 °02’08”S, 119 ° 11 ’ 17 ”E, 12 April 2011, dug from burrow, R. Teale, M. Greenham ( WAM T 113574); 1 female, same data except 23 °02’04”S, 119 ° 11 ’ 32 ”E ( WAM T 113576); 1 female, South Parmelia, 52 km NW. of Newman, 23 °05’ 11 ”S, 119 ° 19 ’07”E, 16 April 2011, dug from burrow, R. Teale, M. Greenham ( WAM T 113582); 1 female, Southern Flank, 71 km NW. of Newman 22 ° 59 ’ 40 ”S, 119 °09’ 55 ”E, 17 April 2011, dug from burrow, R. Teale, M. Greenham ( WAM T 113612); 1 male, Tom Price, 22 ° 42 ’S, 117 ° 47 ’E, 3 January 1994, M. Hyams ( WAM T 30075 View Materials , formerly 94 / 486); 1 female, Tom Price Airport, 14 km NW. of Tom Price, 22 ° 34 ’ 48 ”S, 117 ° 43 ’ 51 ”E, 1–30 November 2007, D. Kamien, J. Adcroft ( WAM T 102911); 2 males, Warramboo, 50.5 km W. of Pannawonica, site WA3, 21° 40 ’00”S, 115 ° 50 ’06”E, 14–20 March 2005, dry pitfall traps, D. Kamien ( WAM T 72404 View Materials ); 1 male, Warramboo, 52.1 km W. of Pannawonica, site WA2, 21° 39 ’ 49 ”S, 115 ° 49 ’ 10 ”E, 14–20 March 2005, dry pitfall traps, D. Kamien ( WAM T 72403 View Materials ); 1 male, 14 km SE. of Wheelarra Hill, Pilbara Biological Survey site BDRS 10, 23° 28 ’ 16 ”S, 120 ° 12 ’ 54 ”E, 6 September 2005 – 10 August 2006, wet pitfall, CALM Pilbara Survey ( WAM T 109352); 1 male, 34 km E. of Wheelarra Hill, Pilbara Biological Survey site BDRS 03, 23 ° 21 ’ 31 ”S, 120 ° 27 ’ 33 ”E, 4 September 2005 – 11 August 2006, wet pitfall, CALM Pilbara Survey ( WAM T 109347); 1 male, 11 km SSE. of Whim Creek Hotel, Pilbara Biological Survey site DRE10, 20° 55 ’ 11.4 ”S, 117 ° 51 ’ 40.6 ”E, 11 July 2003 – 3 October 2004, ethylene glycol pits, CALM Pilbara Survey ( WAM T 109333); 2 males, 1 female, 2 juveniles, 5 km E. of Whim Creek Hotel, Pilbara Biological Survey site DRE08, 20 ° 50 ’ 54.6 ”S, 117 ° 51 ’ 16.2 ”E, 11 July 2003 – 3 October 2004, ethylene glycol pits, CALM Pilbara Survey ( WAM T 109319); 1 male, ca. 7 km NE. of Wodgina mine site, 21 °09’ 10.04 ”S, 118 ° 43 ’ 10.03 ”E, 1 April– 4 May 2010, wet pitfall trap, minor drainage line, P. Bolton ( WAM T 107002); 1 male, ca. 70 km SE. of Wodgina mine, 21 ° 35 ’ 22 ”S, 119 ° 11 ’ 10 ”E, 31 March– 6 April 2010, wet pitfall trap, gully, P. Bolton ( WAM T 105227); 1 male, same data ( WAM T 105228); 1 male, 8.5 km WSW. of Yanyare River Mouth, Pilbara Biological Survey site DRW02, 20 ° 50 ’ 40.6 ”S, 116 ° 22 ’02.9”E, 23 September 2003 – 2 October 2004, wet pitfall, CALM Pilbara Survey ( WAM T 109326).

Etymology. The specific epithet refers to the yellow-brown colouration of parts of the carapace ( mellosa, Latin , of honey, honey-coloured) ( Brown 1956).

Diagnosis. Males of A. mellosa differ from all other named species of the genus by the colouration of the carapace which is overall light yellowish-brown with the cephalic region honey-coloured to very dark reddish-brown ( Fig. 10 View FIGURES 10 – 20 ). Females also have a somewhat darker cephalic region but the colour difference between the cephalic and thoracic regions is much less pronounced ( Fig. 21 View FIGURES 21 – 24 ). Tibia I of males has a large spur and megaspine located medially, and metatarsus I is incrassate with the ‘heel’ of the thickening about half-way; this conformation most closely resembles A. pallida from eastern Queensland from which it differs by the less pronounced metatarsal process and the smaller tibial spur and megaspine. Female genitalia resemble those of A. longitheca Raven, 1984 from central Queensland in possessing long lateral tubes ( Figs 23, 24 View FIGURES 21 – 24 ), but they differ in the shape of the median spermathecae which are coiled in A. longitheca and S-shaped in A. mellosa . See Remarks for further clarification.

Description. Adult male (based on holotype WAM T 94870 View Materials ): medium-sized nemesiid spider (total length 18.4).

Colour: carapace ( Fig. 10 View FIGURES 10 – 20 ) light yellow-brown in thoracic region turning darker towards anterior parts, cephalic region dark reddish-brown, eye region black ( Fig. 13 View FIGURES 10 – 20 ); sternum light yellow-brown, sigilla and labium largely brown ( Fig. 12 View FIGURES 10 – 20 ); abdomen dorsally dark brownish-grey, mottled with small light-brown spots ( Fig. 11 View FIGURES 10 – 20 ), ventrally light yellowish; chelicerae very dark reddish-brown; pedipalp segments reddish-brown, tibia somewhat lighter ( Figs 17, 18 View FIGURES 10 – 20 ); legs light yellowish-brown ( Figs 15, 16 View FIGURES 10 – 20 ).

Carapace: 7.0 long, 5.6 wide; eye group ( Fig. 13 View FIGURES 10 – 20 ) 1.4 wide, 0.7 long; fovea procurved ( Fig. 10 View FIGURES 10 – 20 ).

Sternum and labium: 4.4 long, 2.8 wide; 2 pairs of elongated sigilla in posterior half ( Fig. 12 View FIGURES 10 – 20 ); labium wider than long, slightly indented anteriorly ( Fig. 14 View FIGURES 10 – 20 ).

Maxillae: with> 50 pin-like cuspules, extending to heel ( Fig. 14 View FIGURES 10 – 20 ).

Chelicerae: without rastellum ( Fig. 12 View FIGURES 10 – 20 ), promargin with 9 large teeth, retromargin with 4 smaller teeth proximally.

Abdomen: 7.4 long, 5.0 wide ( Fig. 11 View FIGURES 10 – 20 ). Four spinnerets, terminal segment of posterior lateral spinnerets digitiform.

Pedipalp: length of femur 4.0, patella 2.2, tibia 3.1, tarsus 3.1. Femur with 1 disto-prolateral spine; patella with 3 prolateral spines; tibia with 2 baso-ventral spines and 7–8 prolateral spines in apical half, tibia with slight distoventral depression without setae (accommodates bulb and embolus) ( Figs 17, 18 View FIGURES 10 – 20 ); tarsus terminally blunt; bulb globular, embolus ca. 0.8 long, slightly curved and tapering ( Figs 17–20 View FIGURES 10 – 20 ).

Legs: femur I with row of 7 dorsal spines in basal half and 2 disto-prolateral spines, patella with 2 prolateral spines, metatarsi and tarsi without spines. Tibia I with large spur and megaspine, metatarsi incrassate ( Fig. 16 View FIGURES 10 – 20 ). Tarsi and apical half of metatarsi of legs I and II ventrally with entire dense scopulae, tarsi of legs III and IV with divided weak scopulae. Tarsi, metatarsi and tibiae with numerous dorsal trichobothria of variable length. Paired tarsal claws of legs with two rows of 7–10 teeth each; third tarsal claw reduced and very small.

Leg measurements: length of legs I>IV>II>III. leg I: femur 6.0, patella 3.5, tibia 4.6, metatarsus 4.2, tarsus 2.5, total = 20.8. Dorsal proximal width of patella I = 1.4, tibial index = 0.12, leg I formula = 2.97. Leg II: femur 6.0, patella 3.4, tibia 4.5, metatarsus 4.9, tarsus 3.1, total = 21.9. Dorsal proximal width of patella II = 1.2, tibial index = 0.15; leg II formula = 3.13. Leg III: femur 5.2, patella 3.0, tibia 3.2, metatarsus 6.6, tarsus 2.9, total = 20.9. Dorsal proximal width of patella III = 1.3, tibial index = 0.19; leg III formula = 2.99. Leg IV: femur 7.0, patella 3.5, tibia 5.9, metatarsus 6.3, tarsus 2.9, total = 25.6. Dorsal proximal width of patella IV = 1.3, tibial index = 0.14; leg III formula = 3.66.

Variation: total length 5.1–6.2, carapace 6.1–7.9 long, 16.9–20 wide (n = 12).

Adult female (based on WAM T 96150 View Materials ): medium-sized nemesiid spider (total length 19.5).

Colour: light yellowish-brown with indistinctly darker cephalic region ( Fig. 21 View FIGURES 21 – 24 ), eye region brown; sternum, sigilla and labium uniformly light yellowish-brown; abdomen dorsally dark greyish-brown with purple tinge and mottled with small lighter small spots, ventrally light yellowish-brown; chelicerae orange-brown; pedipalps and legs uniformly light yellow-brown.

Carapace: 7.5 long, 6.4 wide; eye group 1.4 wide, 0.8 long; fovea slightly procurved.

Sternum and labium: 5.0 long, 3.2 wide ( Fig. 22 View FIGURES 21 – 24 ); 2 pairs of indistinct elongated sigilla in posterior half, 1 pair of round sigilla at base of leg II; labium wider than long, slightly indented anteriorly.

Maxillae: with> 100 pin-like cuspules, extending to heel and covering in low density almost entire retrolateral half of maxillae.

Chelicerae: without rastellum, promargin with 9 large teeth, retromargin with 4 smaller teeth proximally.

Abdomen: 8.9 long, 5.9 wide. Four spinnerets, terminal segment of posterior lateral spinnerets digitiform.

Genitalia: median spermathecal stalks S-shaped, spermathecae slightly wider than spermathecal stalks; separate straight and elongated tube accompanies spermatheca ( Figs 23, 24 View FIGURES 21 – 24 ).

Pedipalp: length of femur 3.6, patella 2.3, tibia 2.5, tarsus 2.7. Femur without spine, patella with 2 prolateral spines; tibia with 11–13 irregularly placed spines in mainly apical half (not dorsal); tarsus with scopulae ventrally and 2 baso-ventral spines and curved terminal claw.

Legs: femur I with 1 dorsal spines proximally and 1 disto-prolateral spine, patella with 2 (left leg 1) prolateral spine, tibia with a row of three ventral spines (5 on right leg of which 2 are paired) and 2 disto-ventral spines, metatarsi 1 basal and 1 apical pair of ventral spines (left leg) (3 additional medio-ventral spines on left leg). Tarsi and apical half of metatarsi of leg I and II ventrally with entire dense scopulae, tarsi of legs III and IV with divided very weak scopulae. Tarsi, metatarsi and tibiae with numerous dorsal trichobothria of variable length. Paired tarsal claws of legs with two rows of 10–13 (legs I–II) or 5–7 (legs III–IV) teeth each; third tarsal claw reduced and very small.

Leg measurements: length of legs IV>I>II>III. leg I: femur 6.0, patella 3.1, tibia 4.0, metatarsus 3.5, tarsus 2.3, total = 18.9. Dorsal proximal width of patella I = 1.0, tibial index = 0.14, leg I formula = 2.12. Leg II: femur 5.0, patella 3.0, tibia 3.3, metatarsus 3.1, tarsus 2.1, total = 16.5. Dorsal proximal width of patella II = 0.9, tibial index = 0.14; leg II formula = 1.85. Leg III: femur 4.0, patella 2.5, tibia 2.0, metatarsus 2.9, tarsus 1.7, total = 13.1. Dorsal proximal width of patella III = 1.1, tibial index = 0.24; leg III formula = 1.47. Leg IV: femur 5.5, patella 3.2, tibia 4.0, metatarsus 4.4, tarsus 2.2, total = 19.3. Dorsal proximal width of patella IV = 1.1, tibial index = 0.15; leg III formula = 2.17.

Variation: total length 19.3 –25.0, carapace 5.4–6.6 long, 6.6–8.7 wide (n = 8).

Distribution. Aname mellosa has been found throughout the central Pilbara, from Jimblebar minesite near the township of Newman in the east to Mount Minnie in the west, and south into the Midwest region of Western Australia ( Fig. 5 View FIGURES 5 – 8 ).

Remarks. Care must be taken when identifying A. mellosa as a number of undescribed species from northern Western Australia have a very similar somatic appearance, but have a different genital morphology. The embolus of male A. mellosa is long and slightly curved, but it is often much shorter and/or much more curved in the undescribed species. An unambiguous identification of females or juveniles in this group is currently not possible, however, we have listed females and juveniles when they have been collected with or in close proximity to males. As stated in the Diagnosis, females of A. mellosa have an unusual genital morphology with long, straight lateral tubes in addition to the median pair of spermathecae. This morphology is similar to that of A. longitheca ( Raven 1985 a), but both species differ in the shape of the median spermathecae.

All records of mature males were collected between January and early June suggesting this species to be reproductively active towards the end of the wet season. Many mature males of A. mellosa were removed from their burrows (R. Teale, pers. comm.) where it is likely that they had matured and were waiting for suitable conditions to emerge and search for adult females. The silk-lined open burrow of A. mellosa is dug at an angle into the substrate and lacks a lid ( Figs 2–4 View FIGURES 1 – 4 ) (R. Teale, pers. comm.).

WAM

Western Australian Museum

DNA

Department of Natural Resources, Environment, The Arts and Sport

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Nemesiidae

Genus

Aname