Perrona floriana ( Hilber, 1879 )

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 106

Perrona floriana ( Hilber, 1879)

Figs 31E 1 –E 3 , 4P , 5 , 7

* Pleurotoma (Clavatula) Floriana Hilb. — Hilber 1879: 18, pl. 3, figs 4a–d.

Pleurotoma (Clavatula) Floriana Hilb. — Hoernes & Auinger 1891: 357, pl. 48, figs 16a–b.

? Clavatula floriana (Hilber) 1879 — Bohn-Havas 1973: 1063.

non Perrona floriana (Hilb.) — Sieber 1947: 111 [= Perrona loetschi nov. sp.].

non Clavatula (Perrona) floriana Hilber — Švagrovský 1958: 20, pl. 6, fig. 3 [= Perrona vindobonensis (Quenstedt, 1884) ]. non Clavatula (Perrona) floriana ( Hilber, 1879) — Mikuž 1998: 77, pl. 5, figs 3, 6, 7 [= Perrona ernae ( Šuklje, 1929) ].

non Perrona floriana ( Hilber, 1879) — Mikuž 2009: 33, pl.11, fig. 147 [= Perrona ernae ( Šuklje, 1929) ].

Type material. Holotype: NHMW 1858/0020/0012, Guglitz at St. Florian ( Austria), SL: 24.5 mm, MD: 10.4 mm, illustrated in Hilber (1879, pl. 3, figs 4a–d) and in Hoernes & Auinger (1891, pl. 48, figs 16a–b); figs 31E 1 –E 3, 4P. Revised description. Shell medium-sized, moderately solid, broad fusiform, with weakly gradate spire; apical angle 34°. Protoconch not preserved. Teleoconch of at least nine whorls. Early teleoconch whorls with two adsutural smooth, narrow spiral cords separated by deep concavity with strongly opisthocline axial riblets. On fifth teleoconch whorl profile subcylindrical, subsutural cord weakens. Subsutural collar broad, flat, not swollen. Whorl profile flat-sided with periphery at abapical suture. Suprasutural cord disappears, surface below collar area covered in fine flattened cords separated by narrow, shallow grooves. Suture narrowly incised, superficial. Last whorl ~65% of total height, broad. Collar and subsutural ramp not sharply delimited. Low-set shoulder inflated and broadly rounded, moderately constricted below, base not delimited. Siphonal fasciole weakly developed. Sculpture of fine spiral cords very weak over subsutural collar, slightly stronger over ramp, subobsolete at mid-whorl, strengthening again with cords of alternate strength over base and fasciole. Aperture moderately wide, pyriform. Outer lip thin, smooth within. Anal sinus wide, moderately deep, asymmetrically U-shaped, with apex just below ramp. Siphonal canal moderately long, wide, slightly deflected, shallowly notched at tip. Columella moderately excavated in upper third, straight below, twisted at fasciole, smooth. Columellar and parietal callus slightly thickened, sharply delimited, forming moderately broad callus rim.

Discussion. This species is documented so far by a single specimen, which makes it difficult to evaluate the status of Perrona floriana ( Hilber, 1879) . However, it is so unlike any of its congeners that it probably represents a good species. It is unique amongst the Paratethyan Perrona species in having the subsutural collar not swollen and completely devoid of tubercles on the last two whorls, on the last whorl the subsutural ramp flat and hardly delimited and the shoulder broadly rounded, and again hardly delimited, and the base is not delimited at all. Perrona floriana lacks a shoulder like P. ilonae nov. nom., has a more conical spire whorls and a much broader last whorl. Perrona floriana differs from Perrona vindobonensis (Quenstedt, 1884) by the rounded shoulder and the absence of any subsutural swelling. The same feature allows a separation from P. lydiae ( Hoernes & Auinger, 1891) and P. descendens ( Hilber, 1879) . The species was listed by Bohn-Havas (1973: 1063) but this identification will need confirmation. All records from the literature, except for the original descriptions by Hilber (1879) and Hoernes & Auinger (1891) represent other species.

Paleoenvironment. The assemblage from St. Florian suggests inner neritic, lagoonal environments (own data).

Distribution in Central Paratethys. Badenian (middle Miocene): Styrian Basin: St. Florian ( Austria) ( Hilber 1879).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 4A. ‘Clavatula’ justinae (Hoernes & Auinger, 1891), NHMW ex1949/0005/0110b, Buituri (Romania). B. ‘Clavatula’ mariae (Hoernes & Auinger, 1891), lectotype, NHMW 1849/0013/0009, Mold (Austria). C. ‘Clavatula’ olgae (Hoernes & Auinger, 1891), lectotype, NHMW 1862/0001/0552, Buituri (Romania). D. ‘Clavatula’ orientoromana Báldi, 1960, holotype, M.60.145, Szokolya (Hungary). E. ‘Clavatula’ juliae (Hoernes & Auinger 1891), paralectotype, NHMW 1949/0005/0043c, Lăpugiu de Sus (Romania). F. ‘Clavatula’ apolloniae (Hoernes & Auinger, 1891), NHMW 1868/0001/0474a, Lăpugiu de Sus (Romania). G. Neoperrona harzhauseri (Kovács & Vicián, 2021), NHMW 2019/0151/019a, Letkés (Hungary). H. Neoperrona zoltanorum nov. sp., holotype, NHMW 2019/0151/0108, Letkés (Hungary). I. Scaevatula sidoniae (Hoernes & Auinger, 1891), NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). J. Pusionella palatina Strausz, 1954, NHMW 2021/0159/0003, Várpalota (Hungary). K. Pusionella hofmanni nov. sp., NHMW 1867/0019/0136, holotype, Coşteiu de Sus (Romania). L. Perrona barbarae (Hoernes & Auinger, 1891) NHMW 1949/0005/0103a, Rückersdorf (Austria). M. Perrona descendens (Hilber, 1879), NHMW 1846/0049/0017, St. Florian (Austria). N. Perrona emmae (Hoernes & Auinger 1891), NHMW 2021/0127/0005, Enzesfeld (Austria), NHMW 1970/1396/1449, Baden (Austria). O. Perrona ernae (Šuklje, 1929), NHMW 2021/0156/0001, Weinsteig (Austria). P. Perrona floriana (Hilber, 1879), holotype, NHMW 1858/0020/0012, Guglitz at St. Florian (Austria). Q. Perrona ilonae nov. nom., M.61.4367, Szentkút, Mátraverebély (Hungary). R. Perrona lydiae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0112, Gainfarn (Austria). S. Perrona oliviae (Hoernes & Auinger 1891), paralectotype, NHMW 1873/0026/0027, Lăpugiu de Sus (Romania). T. Perrona rosaliae (Hoernes & Auinger 1891), lectotype, NHMW 1999z0098/0002, Vienna/Pötzleinsdorf (Austria). U. Perrona sabinae (Hoernes & Auinger 1891), paralectotype, NHMW 2021/0154/0001, Lăpugiu de Sus (Romania). V. Perrona styriaca (Hilber, 1879), lectotype, NHMW 1846/0037/0309, Pöls (Austria). W. Perrona ursulae (Hoernes & Auinger, 1891), paralectotype, NHMW 1846/0001/1294, Buituri (Romania). X. Perrona vindobonensis (Quenstedt, 1884), paralectotype, NHMW 1997z0178/1467, Gainfarn (Austria), Y. Perrona wanzenboecki nov. nom., NHMW 2021/0158/0001, neotype, St. Veit an der Triesting (Austria). Z. Tomellana dulaii nov. sp., NHMW 2021/0148/0002, holotype, Nemeşeşti (Romania). Aa. Tomellana aueri nov. sp., NHMW 2013/0078/0436, Baden (Austria). Ab. Striopusionella szekelyhidiae (Kovács & Vicián, 2021), holotype, PAL 2021.19.1, Letkés (Hungary). Ac. Scaevatula sidoniae (Hoernes & Auinger, 1891). NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). Scale bar is 5 mm for all specimens except for Perrona barbarae and protoconch of Scaevatula sidoniae.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 7. Apertures of Paratethyan Clavatulidae (O. = Olegia, ‘C.’ = ‘Clavatula’, N. = Neoperrona, P. = Perrona, Pu. = Pusionella, Sc. = Scaevatula, S. = Striopusionella, T. = Tomellana).

FIGURE 31 A1–A3. Perrona descendens (Hilber, 1879), lectotype, NHMW 1862/0033/0034, Pöls (Austria). B1–B3. Perrona descendens (Hilber, 1879), paralectotype, NHMW 1949/0005/0101, St. Florian (Austria). C1–C3. Perrona descendens (Hilber, 1879), paralectotype, NHMW 1861/0001/0260, Pöls (Austria). D1–D3. Perrona descendens (Hilber, 1879), paralectotype, NHMW 1861/0001/0260a, Pöls (Austria). E1–E3. Perrona floriana (Hilber, 1879), holotype, NHMW 1858/0020/0012, Guglitz at St. Florian (Austria). F1–F2. Perrona ilonae nov. nom., M.61.4367, Szentkút, Mátraverebély (Hungary).