Clavatula ’ orientoromana Báldi, 1960

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 95-96

‘ Clavatula ’ orientoromana Báldi, 1960

Figs 29C, D 1 –D 2 , 4D

* Clavatula orientoromana n. sp. — Báldi 1960: 81, pl. 3, figs 6a–b.

Clavatula (Surcula) nodosa borsodensis n. ssp. — Csepreghy-Meznerics 1969: 96, pl. 6, fig. 5.

Clavatula (Surcula) nodosa borsodensis Csepr. Mezn. — Csepreghy-Meznerics 1972: 32, pl. 16, fig. 17. Clavatula orientoromana Báldi, 1960 — Pálfy et al. 2008: 105.

Clavatula (Surcula) nodosa borsodensis Csepreghy-Meznerics, 1969 — Pálfy et al. 2008: 105.

Type material. Holotype: M.60.145, Szokolya ( Hungary), SL: 34 mm, MD: 11.3 mm, illustrated in Báldi (1960, pl. 3, figs 6a–b), stored in the Hungarian Natural History Museum , Budapest ( Hungary), fig. 28C, 4D . Paratypes: M60.146, M60.147, Szokolya ( Hungary), stored in the Hungarian Natural History Museum , Budapest ( Hungary), figs 29D 1 –D 2 .

Additional studied material. M.70.619, Rendek-völgyi quarry, Borsodbóta ( Hungary), holotype of Clavatula (Surcula) nodosa borsodensis Csepreghy-Meznerics, 1969 , stored in the Hungarian Natural History Museum , Budapest ( Hungary) .

Revised description. Shell medium-sized, solid with high, conical spire. Protoconch paucispiral, poorly preserved, high conical. Teleoconch of at least eight whorls. Early teleoconch whorls with two prominent adsutural spiral rows of rounded beads separated by narrow concavity. Abapically mid-portion widening, suprasutural beads slightly larger, forming periphery; weak spiral cords covering entire whorl. On fifth whorl subsutural row of beads weakens, cord swells to form subsutural collar. Suprasutural beads strengthen and become tubercles. Spiral cords becoming more prominent covering collar and suprasutural tubercles. Suture narrowly incised, running below beads. Last whorl 55% of total height. Subsutural collar narrow, low, poorly delimited. Subsutural ramp concave, delimited by row of rounded tubercles, convex below, strongly constricted at base. Fasciole poorly delimited. Spiral sculpture of fine threads on collar and ramp, below shoulder with narrow cords of roughly alternate strength extending to fasciole. Peribasal cord slightly more prominent.Aperture and outer lip incomplete.Anal sinus moderately deep and wide, symmetrically U-shaped, with apex on adapical half of ramp. Siphonal canal long, narrow, recurved at tip. Columella straight to weakly twisted, smooth.

Discussion. Despite the fragmentary preservation, this species is well-defined by its two prominent adsutural cords of relatively large beads on early whorls. ‘ Clavatula ’ orientoromana Báldi, 1960 is superficially reminiscent of ‘ Clavatula ’ romana ( Defrance, 1826) , from the Pliocene of Italy, but differs in its much higher spire and the comparatively smooth subsutural collar on later teleoconch whorls. Moreover, ‘ C. ’ orientoromana differs in its neanic whorl sculpture being composed of two adsutural beaded cords separated by a smooth mid-whorl, as opposed to a tripartite sculpture in ‘ C. ’ romana , which is herein considered part of the mystica -clade. ‘ Clavatula ’ orientoromana differs from Clavatula in its more slender fusiform shell and from the bimarginata -clade in not having axial riblets at mid-whorl.

The early Miocene ‘ Clavatula ’ mariae ( Hoernes & Auinger, 1891) has a tripartite sculpture on early teleoconch whorls, less conical whorls, and a less constricted base. ‘ Clavatula ’ boreoromana Kautsky, 1925, from the early Miocene of the North Sea, agrees with ‘ C. ’ orientoromana in its general shape with a high spire, but has slightly opisthocline beads and coarser spiral cords (see Kautsky 1925: 154, pl. 10, figs 28–29). The Egerian (Chattian/ Aquitanian) ‘ Clavatula ’ danuvii Vicián, Kovács & Stein, 2019, from Hungary, lacks prominent beads (see Vicián et al. 2019, 256. pl. 5, figs 19–20).

Kovács & Vicián (2021) recognized Clavatula (Surcula) nodosa borsodensis Csepreghy-Meznerics, 1969 as subjective junior synonym of Clavatula orientoromana , which is followed herein.

Paleoenvironment. The fauna from Szokolya ( Hungary) corresponds to that of the basinal clays of the Baden Formation in the Vienna Basin, which formed in middle to outer neritic environments ( Kranner et al. 2021).

Distribution in Central Paratethys. Badenian (middle Miocene): Bükk Mountains: Borsodbóta ( Hungary) ( Csepreghy-Meznerics 1969, 1972); Pannonian Basin: Szokolya ( Hungary) ( Báldi 1960).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width.

FIGURE 4A. ‘Clavatula’ justinae (Hoernes & Auinger, 1891), NHMW ex1949/0005/0110b, Buituri (Romania). B. ‘Clavatula’ mariae (Hoernes & Auinger, 1891), lectotype, NHMW 1849/0013/0009, Mold (Austria). C. ‘Clavatula’ olgae (Hoernes & Auinger, 1891), lectotype, NHMW 1862/0001/0552, Buituri (Romania). D. ‘Clavatula’ orientoromana Báldi, 1960, holotype, M.60.145, Szokolya (Hungary). E. ‘Clavatula’ juliae (Hoernes & Auinger 1891), paralectotype, NHMW 1949/0005/0043c, Lăpugiu de Sus (Romania). F. ‘Clavatula’ apolloniae (Hoernes & Auinger, 1891), NHMW 1868/0001/0474a, Lăpugiu de Sus (Romania). G. Neoperrona harzhauseri (Kovács & Vicián, 2021), NHMW 2019/0151/019a, Letkés (Hungary). H. Neoperrona zoltanorum nov. sp., holotype, NHMW 2019/0151/0108, Letkés (Hungary). I. Scaevatula sidoniae (Hoernes & Auinger, 1891), NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). J. Pusionella palatina Strausz, 1954, NHMW 2021/0159/0003, Várpalota (Hungary). K. Pusionella hofmanni nov. sp., NHMW 1867/0019/0136, holotype, Coşteiu de Sus (Romania). L. Perrona barbarae (Hoernes & Auinger, 1891) NHMW 1949/0005/0103a, Rückersdorf (Austria). M. Perrona descendens (Hilber, 1879), NHMW 1846/0049/0017, St. Florian (Austria). N. Perrona emmae (Hoernes & Auinger 1891), NHMW 2021/0127/0005, Enzesfeld (Austria), NHMW 1970/1396/1449, Baden (Austria). O. Perrona ernae (Šuklje, 1929), NHMW 2021/0156/0001, Weinsteig (Austria). P. Perrona floriana (Hilber, 1879), holotype, NHMW 1858/0020/0012, Guglitz at St. Florian (Austria). Q. Perrona ilonae nov. nom., M.61.4367, Szentkút, Mátraverebély (Hungary). R. Perrona lydiae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0112, Gainfarn (Austria). S. Perrona oliviae (Hoernes & Auinger 1891), paralectotype, NHMW 1873/0026/0027, Lăpugiu de Sus (Romania). T. Perrona rosaliae (Hoernes & Auinger 1891), lectotype, NHMW 1999z0098/0002, Vienna/Pötzleinsdorf (Austria). U. Perrona sabinae (Hoernes & Auinger 1891), paralectotype, NHMW 2021/0154/0001, Lăpugiu de Sus (Romania). V. Perrona styriaca (Hilber, 1879), lectotype, NHMW 1846/0037/0309, Pöls (Austria). W. Perrona ursulae (Hoernes & Auinger, 1891), paralectotype, NHMW 1846/0001/1294, Buituri (Romania). X. Perrona vindobonensis (Quenstedt, 1884), paralectotype, NHMW 1997z0178/1467, Gainfarn (Austria), Y. Perrona wanzenboecki nov. nom., NHMW 2021/0158/0001, neotype, St. Veit an der Triesting (Austria). Z. Tomellana dulaii nov. sp., NHMW 2021/0148/0002, holotype, Nemeşeşti (Romania). Aa. Tomellana aueri nov. sp., NHMW 2013/0078/0436, Baden (Austria). Ab. Striopusionella szekelyhidiae (Kovács & Vicián, 2021), holotype, PAL 2021.19.1, Letkés (Hungary). Ac. Scaevatula sidoniae (Hoernes & Auinger, 1891). NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). Scale bar is 5 mm for all specimens except for Perrona barbarae and protoconch of Scaevatula sidoniae.

FIGURE 29 A. ‘Clavatula’ inedita Bellardi, 1877, Letkés (Hungary), private collection of Zoltán Kovács and Zoltán Vicián. B. ‘Clavatula’ rusticula (Mayer-Eymar, 1886), lectotype, S 1615, Grund (Austria). C. ‘Clavatula’ orientoromana Báldi, 1960, holotype, M.60.145, Szokolya (Hungary). D1–D2. ‘Clavatula’ orientoromana Báldi, 1960, paratype, M60.146, Szokolya (Hungary). E1–E2. ‘Clavatula’ justinae (Hoernes & Auinger, 1891), paralectotypes, NHMW 1849/0005/0110b, Buituri (Romania).F1–F3. ‘Clavatula’ justinae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0110a, Buituri (Romania). G. ‘Clavatula’ justinae (Hoernes & Auinger, 1891), lectotype, NHMW 1849/0005/0110c, Buituri (Romania).